<document id="55206267803768FE7A27897BA410B0C8" ID-CLB-Dataset="312568" ID-DOI="10.5852/ejt.2025.1016.3063" ID-GBIF-Dataset="aa1f865e-3066-437c-a9fb-5d4dc5ab00b8" ID-ISSN="2118-9773" ID-Zenodo-Dep="17245005" ID-ZooBank="971686D8-3C1F-4F8F-9AA1-FBB8BC97E175" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="julia" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1759333875172" checkinUser="plazi" docAuthor="Shafaie, Sepideh, Stano PEKÁR &amp; Osman SEYYAR" docDate="2025" docId="F910AB10A14BFF9D163FFCB75465FC39" docLanguage="en" docName="EJT.2025.1016.1-139.pdf" docOrigin="European Journal of Taxonomy 1016" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3063/13687" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Zodarion levyi Shafaie &amp; Pekar 2025, sp. nov." docType="treatment" docVersion="9" lastPageNumber="90" masterDocId="0529D368A11DFFC41411FFD15518FFDF" masterDocTitle="New data on zodariid spiders (Arachnida, Araneae) in the Middle East and Eastern Mediterranean" masterLastPageNumber="139" masterPageNumber="1" pageNumber="87" updateTime="1760042680557" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="F3A44F5E082FF15A6D6566FFC2B8CBDA">New data on zodariid spiders (Arachnida, Araneae) in the Middle East and Eastern Mediterranean</mods:title>
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<mods:namePart id="E7193F9932120A5C002C87FC39564718">Shafaie, Sepideh</mods:namePart>
<mods:affiliation id="C45A643B99F3664B22A8E5D553E3F541">Department of Botany and Zoology, Faculty of Sciences, Masaryk University, Kotlářská 2, 61137 Brno, Czech Republic.</mods:affiliation>
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<mods:namePart id="EAEBC5F8011069BCF7A46FB348EFD63D">Stano PEKÁR</mods:namePart>
<mods:affiliation id="2D59A3E4E21266D0B1D57D923D3E3465">Department of Botany and Zoology, Faculty of Sciences, Masaryk University, Kotlářská 2, 61137 Brno, Czech Republic.</mods:affiliation>
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<mods:namePart id="A5499309327FCF156D8AB68BC7F100C5">Osman SEYYAR</mods:namePart>
<mods:affiliation id="CD3A665AEF548DFE2A24944213388672">Department of Biology, Faculty of Art &amp; Sciences, Niğde Ömer Halisdemir University, TR- 51240 Niğde, Turkey</mods:affiliation>
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<heading id="2A4EAD6AA14BFF92163FFCB7511EFC5E" box="[558,1030,870,897]" centered="true" fontSize="11" level="2" pageId="86" pageNumber="87" reason="2">
<taxonomicName id="B6B96185A14BFF92163FFCB756BDFC5E" authority="Shafaie &amp; Pekar" authorityName="Shafaie &amp; Pekar" authorityYear="2025" box="[558,933,870,897]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="levyi" status="sp. nov." subFamily="Zodariinae">
<emphasis id="43CDC614A14BFF92163FFCB757C4FC5E" bold="true" box="[558,732,870,897]" italics="true" pageId="86" pageNumber="87">Zodarion levyi</emphasis>
Shafaie &amp; Pekár
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<taxonomicNameLabel id="58FE7B6FA14BFF9217BDFCB6511EFC5E" box="[940,1030,871,897]" pageId="86" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act: 
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<figureCitation id="E9820683A14BFF9216AAFC7C5659FC18" box="[699,833,941,967]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="87.[189,232,1927,1953]" captionStartId-1="88.[189,233,1928,1954]" captionStartId-2="89.[189,232,1889,1915]" captionTargetBox-0="[189,1398,468,1905]" captionTargetBox-1="[189,1385,573,1890]" captionTargetBox-2="[193,1394,1047,1827]" captionTargetId-0="figure-104@87.[189,1398,468,1905]" captionTargetId-1="figure-181@88.[189,1398,573,1903]" captionTargetId-2="figure-239@89.[189,1398,1043,1851]" captionTargetPageId-0="87" captionTargetPageId-1="88" captionTargetPageId-2="89" captionText-0="Fig. 67. Habitus of Zodarion levyi Shafaie &amp; Pekár sp. nov. A–B. Holotype, ♂ (HUJINV-Ar 21303). C–D. Paratype, ♀ (HUJINV-Ar 21304). A. Dorsal view. B. Ventral view. C. Dorsal view. D. Ventral view. Scale bars: 1 mm." captionText-1="Fig. 68. Male palp of Zodarion levyi Shafaie &amp; Pekár sp. nov., holotype (HUJINV-Ar 21303). A. Prolateral view. B. Ventral view. C. Retrolateral view. D. Prolateral view. E. Ventral view. F. Retrolateral view. G. Embolar tip, ventral view. Scale bars: 0.2 mm." captionText-2="Fig. 69. Female copulatory organ of Zodarion levyi Shafaie &amp; Pekár sp. nov., paratype (HUJINV-Ar 21304). A. Intact epigyne, ventral view. B. Epigyne, ventral view. C. Vulva, ventral view. D. Vulva, ventral view. Scale bars: 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.17245254" figureDoi-1="http://doi.org/10.5281/zenodo.17245256" figureDoi-2="http://doi.org/10.5281/zenodo.17245258" httpUri-0="https://zenodo.org/record/17245254/files/figure.png" httpUri-1="https://zenodo.org/record/17245256/files/figure.png" httpUri-2="https://zenodo.org/record/17245258/files/figure.png" pageId="86" pageNumber="87">Figs 67–69</figureCitation>
, 
<figureCitation id="E9820683A14BFF92175CFC7C5661FC18" box="[845,889,941,967]" captionStart="Fig" captionStartId="130.[189,233,1786,1812]" captionTargetBox="[189,1398,987,1746]" captionTargetId="figure-370@130.[189,1398,987,1746]" captionTargetPageId="130" captionText="Fig. 106. Distribution records across the Middle East and Eastern Mediterranean. 1 = Indozodion lindbergi (Roewer, 1960); 2 = Lachesana gavishae Shafaie &amp; Pekár sp. nov.; 3 = L. lubinae Shafaie &amp; Pekár sp. nov.; 4 = Parazodarion raddei (Simon, 1889); 5 = Pax akilae Shafaie &amp; Pekár sp. nov.; 6 = Zodarion crewsae Coşar, Danışman &amp; Kunt, 2022; 7 = Z. jakubi Shafaie &amp; Pekár sp. nov.; 8 = Z. levyi Shafaie &amp; Pekár sp. nov.; 9 = Z. milani Shafaie &amp; Pekár sp. nov.; 10 = Z. namrun Dimitrov, 2024; 11 = Z. nicki Strand, 1914; 12 = Z. nitidum Audouin, 1826; 13 = Z. tunetiacum Strand, 1906; 14 = Z. zonsteini Shafaie &amp; Pekár sp. nov." figureDoi="http://doi.org/10.5281/zenodo.17245348" httpUri="https://zenodo.org/record/17245348/files/figure.png" pageId="86" pageNumber="87">106</figureCitation>
</paragraph>
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<subSubSection id="39A3498DA14BFF9214ACFC3F51C4FAD5" pageId="86" pageNumber="87" type="diagnosis">
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<emphasis id="43CDC614A14BFF9214ACFC3F542EFBD7" bold="true" box="[189,310,1006,1032]" pageId="86" pageNumber="87">Diagnosis</emphasis>
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This new species is most similar to 
<taxonomicName id="B6B96185A14BFF921660FBCF57C7FBE8" authorityName="Komnenov &amp; Chatzaki" authorityYear="2016" box="[625,735,1053,1079]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="beroni">
<emphasis id="43CDC614A14BFF921660FBCF57C7FBE8" box="[625,735,1053,1079]" italics="true" pageId="86" pageNumber="87">Z. beroni</emphasis>
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by the small body size and asymmetrically bi-concave embolar tip (
<figureCitation id="E9820683A14BFF92154AFB9154D0FB85" box="[347,456,1088,1114]" captionStart="Fig" captionStartId="88.[189,233,1928,1954]" captionTargetBox="[189,1385,573,1890]" captionTargetId="figure-181@88.[189,1398,573,1903]" captionTargetPageId="88" captionText="Fig. 68. Male palp of Zodarion levyi Shafaie &amp; Pekár sp. nov., holotype (HUJINV-Ar 21303). A. Prolateral view. B. Ventral view. C. Retrolateral view. D. Prolateral view. E. Ventral view. F. Retrolateral view. G. Embolar tip, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.17245256" httpUri="https://zenodo.org/record/17245256/files/figure.png" pageId="86" pageNumber="87">Fig. 68G</figureCitation>
vs 
<figureCitation id="E9820683A14BFF9215E5FB915744FB84" box="[500,604,1088,1115]" captionStart="Fig" captionStartId="52.[189,232,1893,1919]" captionTargetBox="[189,1397,847,1846]" captionTargetId="figure-115@52.[189,1398,847,1850]" captionTargetPageId="52" captionText="Fig. 37. Copulatory organs of Zodarion beroni Komnenov &amp; Chatzaki, 2016 (CMU). A–C, F. Male palp, with rotated bulb inside. D–E. Female copulatory organ. A. Prolateral view. B. Ventral view. C. Retrolateral view. D. Intact epigyne, ventral view. E. Vulva, ventral view. F. Detail of embolus, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.17245121" httpUri="https://zenodo.org/record/17245121/files/figure.png" pageId="86" pageNumber="87">Fig. 37F</figureCitation>
) but can be distinguished by the following characters: (1) a more extended prolateral process of median apophysis (vs elongated, cf. 
<figureCitation id="E9820683A14BFF9217CDFBB5517EFBA1" box="[988,1126,1124,1150]" captionStart="Fig" captionStartId="88.[189,233,1928,1954]" captionTargetBox="[189,1385,573,1890]" captionTargetId="figure-181@88.[189,1398,573,1903]" captionTargetPageId="88" captionText="Fig. 68. Male palp of Zodarion levyi Shafaie &amp; Pekár sp. nov., holotype (HUJINV-Ar 21303). A. Prolateral view. B. Ventral view. C. Retrolateral view. D. Prolateral view. E. Ventral view. F. Retrolateral view. G. Embolar tip, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.17245256" httpUri="https://zenodo.org/record/17245256/files/figure.png" pageId="86" pageNumber="87">Fig. 68B, E</figureCitation>
and 
<figureCitation id="E9820683A14BFF9210B0FBB55014FBA1" box="[1185,1292,1124,1150]" captionStart="Fig" captionStartId="52.[189,232,1893,1919]" captionTargetBox="[189,1397,847,1846]" captionTargetId="figure-115@52.[189,1398,847,1850]" captionTargetPageId="52" captionText="Fig. 37. Copulatory organs of Zodarion beroni Komnenov &amp; Chatzaki, 2016 (CMU). A–C, F. Male palp, with rotated bulb inside. D–E. Female copulatory organ. A. Prolateral view. B. Ventral view. C. Retrolateral view. D. Intact epigyne, ventral view. E. Vulva, ventral view. F. Detail of embolus, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.17245121" httpUri="https://zenodo.org/record/17245121/files/figure.png" pageId="86" pageNumber="87">Fig. 37B</figureCitation>
); and (2) slightly convergent embolar arms (vs divergent arms, cf. 
<figureCitation id="E9820683A14BFF92177DFB5656CEFB7E" box="[876,982,1159,1185]" captionStart="Fig" captionStartId="88.[189,233,1928,1954]" captionTargetBox="[189,1385,573,1890]" captionTargetId="figure-181@88.[189,1398,573,1903]" captionTargetPageId="88" captionText="Fig. 68. Male palp of Zodarion levyi Shafaie &amp; Pekár sp. nov., holotype (HUJINV-Ar 21303). A. Prolateral view. B. Ventral view. C. Retrolateral view. D. Prolateral view. E. Ventral view. F. Retrolateral view. G. Embolar tip, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.17245256" httpUri="https://zenodo.org/record/17245256/files/figure.png" pageId="86" pageNumber="87">Fig. 68G</figureCitation>
and 
<figureCitation id="E9820683A14BFF921003FB565165FB7E" box="[1042,1149,1159,1185]" captionStart="Fig" captionStartId="52.[189,232,1893,1919]" captionTargetBox="[189,1397,847,1846]" captionTargetId="figure-115@52.[189,1398,847,1850]" captionTargetPageId="52" captionText="Fig. 37. Copulatory organs of Zodarion beroni Komnenov &amp; Chatzaki, 2016 (CMU). A–C, F. Male palp, with rotated bulb inside. D–E. Female copulatory organ. A. Prolateral view. B. Ventral view. C. Retrolateral view. D. Intact epigyne, ventral view. E. Vulva, ventral view. F. Detail of embolus, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.17245121" httpUri="https://zenodo.org/record/17245121/files/figure.png" pageId="86" pageNumber="87">Fig. 37F</figureCitation>
). Females of 
<taxonomicName id="B6B96185A14BFF921131FB595060FB7E" authorityName="Shafaie &amp; Pekar" authorityYear="2025" box="[1312,1400,1159,1185]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="levyi" status="sp. nov.">
<emphasis id="43CDC614A14BFF921131FB595060FB7E" box="[1312,1400,1159,1185]" italics="true" pageId="86" pageNumber="87">Z. levyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58FE7B6FA14BFF9214ACFB7A5401FB1A" box="[189,281,1195,1221]" pageId="86" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
can be recognized from the other members of the 
<taxonomicName id="B6B96185A14BFF921764FB7B56A8FB1B" authorityName="Nosek" authorityYear="1905" box="[885,944,1194,1220]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="thoni">
<emphasis id="43CDC614A14BFF921764FB7B56A8FB1B" box="[885,944,1194,1220]" italics="true" pageId="86" pageNumber="87">thoni</emphasis>
</taxonomicName>
-group by an extended median septum (
<figureCitation id="E9820683A14BFF9214D6FB1C544FFB38" box="[199,343,1229,1255]" captionStart="Fig" captionStartId="89.[189,232,1889,1915]" captionTargetBox="[193,1394,1047,1827]" captionTargetId="figure-239@89.[189,1398,1043,1851]" captionTargetPageId="89" captionText="Fig. 69. Female copulatory organ of Zodarion levyi Shafaie &amp; Pekár sp. nov., paratype (HUJINV-Ar 21304). A. Intact epigyne, ventral view. B. Epigyne, ventral view. C. Vulva, ventral view. D. Vulva, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.17245258" httpUri="https://zenodo.org/record/17245258/files/figure.png" pageId="86" pageNumber="87">Fig. 69A–B</figureCitation>
), whereas it is less extended in 
<taxonomicName id="B6B96185A14BFF9216D7FB1F5629FB38" authorityName="Komnenov &amp; Chatzaki" authorityYear="2016" box="[710,817,1229,1255]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="beroni">
<emphasis id="43CDC614A14BFF9216D7FB1F5629FB38" box="[710,817,1229,1255]" italics="true" pageId="86" pageNumber="87">Z. beroni</emphasis>
</taxonomicName>
(
<figureCitation id="E9820683A14BFF921750FB1C56B6FB38" box="[833,942,1229,1255]" captionStart="Fig" captionStartId="52.[189,232,1893,1919]" captionTargetBox="[189,1397,847,1846]" captionTargetId="figure-115@52.[189,1398,847,1850]" captionTargetPageId="52" captionText="Fig. 37. Copulatory organs of Zodarion beroni Komnenov &amp; Chatzaki, 2016 (CMU). A–C, F. Male palp, with rotated bulb inside. D–E. Female copulatory organ. A. Prolateral view. B. Ventral view. C. Retrolateral view. D. Intact epigyne, ventral view. E. Vulva, ventral view. F. Detail of embolus, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.17245121" httpUri="https://zenodo.org/record/17245121/files/figure.png" pageId="86" pageNumber="87">Fig. 37D</figureCitation>
), 
<taxonomicName id="B6B96185A14BFF9217D2FB1F5172FB38" authorityName="Kulczynski" authorityYear="1908" box="[963,1130,1229,1255]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="granulatum">
<emphasis id="43CDC614A14BFF9217D2FB1F5172FB38" box="[963,1130,1229,1255]" italics="true" pageId="86" pageNumber="87">Z. granulatum</emphasis>
</taxonomicName>
(
<figureCitation id="E9820683A14BFF92106BFB1C51FFFB37" box="[1146,1255,1229,1256]" captionStart="Fig" captionStartId="70.[187,231,1262,1288]" captionTargetBox="[189,1392,265,1216]" captionTargetId="figure-179@70.[189,1398,265,1217]" captionTargetPageId="70" captionText="Fig. 53. Copulatory organs of Zodarion granulatum Kulczyński, 1908 (CMU). A–C, F. Male palp. D–E. Female copulatory organ. A. Prolateral view. B. Ventral view. C. Retrolateral view. D. Intact epigyne, ventral view. E. Vulva, ventral view. F. Detail of embolus, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.17245204" httpUri="https://zenodo.org/record/17245204/files/figure.png" pageId="86" pageNumber="87">Fig. 53D</figureCitation>
), 
<taxonomicName id="B6B96185A14BFF9210EDFB1F506EFB38" authorityName="Dimitrov" authorityYear="2024" box="[1276,1398,1230,1255]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="namrun">
<emphasis id="43CDC614A14BFF9210EDFB1F506EFB38" box="[1276,1398,1230,1255]" italics="true" pageId="86" pageNumber="87">Z. namrun</emphasis>
</taxonomicName>
(
<figureCitation id="E9820683A14BFF9214D7FB21544DFAD4" box="[198,341,1264,1291]" captionStart="Fig" captionStartId="104.[189,233,1929,1955]" captionTargetBox="[196,1398,1090,1894]" captionTargetId="figure-271@104.[189,1398,1085,1909]" captionTargetPageId="104" captionText="Fig. 82. Female copulatory organ of Zodarion namrun Dimitrov, 2024 (NMP P6A 7505). A. Intact epigyne, ventral view. B. Epigyne, ventral view. C. Vulva, ventral view. D. Vulva, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.17245292" httpUri="https://zenodo.org/record/17245292/files/figure.png" pageId="86" pageNumber="87">Fig. 82A–B</figureCitation>
), 
<taxonomicName id="B6B96185A14BFF92157BFB205717FAD5" authorityName="Kulczynski" authorityYear="1908" box="[362,527,1264,1290]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="reticulatum">
<emphasis id="43CDC614A14BFF92157BFB205717FAD5" box="[362,527,1264,1290]" italics="true" pageId="86" pageNumber="87">Z. reticulatum</emphasis>
</taxonomicName>
(
<figureCitation id="E9820683A14BFF92160EFB215795FAD4" box="[543,653,1264,1291]" captionStart="Fig" captionStartId="113.[189,233,1523,1549]" captionTargetBox="[189,1398,265,1491]" captionTargetId="figure-100@113.[189,1398,265,1491]" captionTargetPageId="113" captionText="Fig. 90. Copulatory organs of Zodarion reticulatum Kulczyński, 1908 (CMU). A–C. Male palp. D–E. Female copulatory organ. A. Prolateral view. B. Ventral view. C. Retrolateral view. D. Intact epigyne, ventral view. E. Vulva, ventral view. F. Detail of embolus, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.17245308" httpUri="https://zenodo.org/record/17245308/files/figure.png" pageId="86" pageNumber="87">Fig. 90D</figureCitation>
), 
<taxonomicName id="B6B96185A14BFF9216B0FB20562BFAD5" authorityName="Caporiacco" authorityYear="1948" box="[673,819,1264,1290]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="rhodiense">
<emphasis id="43CDC614A14BFF9216B0FB20562BFAD5" box="[673,819,1264,1290]" italics="true" pageId="86" pageNumber="87">Z. rhodiense</emphasis>
</taxonomicName>
(
<figureCitation id="E9820683A14BFF921752FB2156A9FAD4" box="[835,945,1264,1291]" captionStart="Fig" captionStartId="115.[189,233,1893,1919]" captionTargetBox="[189,1398,819,1865]" captionTargetId="figure-168@115.[189,1398,819,1865]" captionTargetPageId="115" captionText="Fig. 92. Copulatory organs of Zodarion rhodiense Caporiacco, 1948 (CMU). A–C, F. Male palp. D–E. Female copulatory organ. A. Prolateral view. B. Ventral view. C. Retrolateral view. D. Intact epigyne, ventral view. E. Vulva, ventral view. F. Detail of embolus, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.17245315" httpUri="https://zenodo.org/record/17245315/files/figure.png" pageId="86" pageNumber="87">Fig. 92D</figureCitation>
), and 
<taxonomicName id="B6B96185A14BFF9217E7FB205149FAD5" authorityName="Nosek" authorityYear="1905" box="[1014,1105,1264,1290]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="thoni">
<emphasis id="43CDC614A14BFF9217E7FB205149FAD5" box="[1014,1105,1264,1290]" italics="true" pageId="86" pageNumber="87">Z. thoni</emphasis>
</taxonomicName>
(
<figureCitation id="E9820683A14BFF921073FB2151C8FAD5" box="[1122,1232,1264,1290]" captionStart="Fig" captionStartId="119.[189,232,1929,1955]" captionTargetBox="[189,1398,801,1894]" captionTargetId="figure-278@119.[189,1398,801,1897]" captionTargetPageId="119" captionText="Fig. 96. Copulatory organs of Zodarion thoni Nosek, 1905 (CMU). A–C, F. Male palp. D–E. Female copulatory organ. A. Prolateral view. B. Ventral view. C. Retrolateral view. D. Intact epigyne, ventral view. E. Vulva, ventral view. F. Detail of embolus, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.17245323" httpUri="https://zenodo.org/record/17245323/files/figure.png" pageId="86" pageNumber="87">Fig. 96D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="39A3498DA14BFF9214ACFAE3512CFAA5" box="[189,1076,1330,1403]" pageId="86" pageNumber="87" type="etymology">
<paragraph id="71061A06A14BFF9214ACFAE3545CFA93" blockId="86.[189,324,1330,1356]" box="[189,324,1330,1356]" pageId="86" pageNumber="87">
<heading id="2A4EAD6AA14BFF9214ACFAE3545CFA93" bold="true" box="[189,324,1330,1356]" fontSize="11" level="3" pageId="86" pageNumber="87" reason="3">
<emphasis id="43CDC614A14BFF9214ACFAE3545CFA93" bold="true" box="[189,324,1330,1356]" pageId="86" pageNumber="87">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="71061A06A14BFF9214ACFAB1512CFAA5" blockId="86.[189,1076,1376,1403]" box="[189,1076,1376,1403]" pageId="86" pageNumber="87">This species is named after the late Gershom Levy, an Israeli arachnologist.</paragraph>
</subSubSection>
<subSubSection id="39A3498DA14BFF9214ACFA7356E5F852" pageId="86" pageNumber="87" type="materials_examined">
<paragraph id="71061A06A14BFF9214ACFA735474FA63" blockId="86.[189,364,1442,1468]" box="[189,364,1442,1468]" pageId="86" pageNumber="87">
<heading id="2A4EAD6AA14BFF9214ACFA735474FA63" bold="true" box="[189,364,1442,1468]" fontSize="11" level="3" pageId="86" pageNumber="87" reason="3">
<emphasis id="43CDC614A14BFF9214ACFA735474FA63" bold="true" box="[189,364,1442,1468]" pageId="86" pageNumber="87">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="71061A06A14BFF9214FBFA005444FA34" blockId="86.[189,1248,1489,1550]" box="[234,348,1489,1515]" pageId="86" pageNumber="87">
<emphasis id="43CDC614A14BFF9214FBFA005444FA34" bold="true" box="[234,348,1489,1515]" pageId="86" pageNumber="87">
<typeStatus id="AE02A4A4A14BFF9214FBFA005444FA34" box="[234,348,1489,1515]" pageId="86" pageNumber="87" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="71061A06A14BFF9214ACFA2551F8F9D1" blockId="86.[189,1248,1489,1550]" box="[189,1248,1523,1550]" pageId="86" pageNumber="87">
<materialsCitation id="C1D1105BA14BFF9214ACFA2551F8F9D1" ID-GBIF-Occurrence="5839008359" box="[189,1248,1523,1550]" collectingDate="1996-05-01" collectorName="J. Levy" country="Israel" location="Odem Forest" pageId="86" pageNumber="87" specimenCode="HUJINV-Ar 21303" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="09AE5A96A14BFF9214ACFA25543CF9D1" box="[189,292,1524,1550]" name="Israel" pageId="86" pageNumber="87">ISRAEL</collectingCountry>
• 
<specimenCount id="67BFD18FA14BFF92152AFA25544CF9D1" box="[315,340,1524,1550]" count="1" pageId="86" pageNumber="87" type="male">♂</specimenCount>
; 
<taxonomicName id="B6B96185A14BFF921570FA2557AEF9D1" authority="Forest, Golan Heights" authorityName="Forest, Golan Heights" box="[353,694,1524,1550]" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="odem">
<location id="74664CDDA14BFF921570FA2554E4F9D1" LSID="urn:lsid:plazi:treatment:F910AB10A14BFF9D163FFCB75465FC39:74664CDDA14BFF921570FA2554E4F9D1" box="[353,508,1524,1550]" country="Israel" name="Odem Forest" pageId="86" pageNumber="87">Odem Forest</location>
, 
<location id="74664CDDA14BFF921619FA2557AEF9D1" LSID="urn:lsid:plazi:treatment:F910AB10A14BFF9D163FFCB75465FC39:74664CDDA14BFF921619FA2557AEF9D1" box="[520,694,1524,1550]" country="Israel" name="Golan Heights" pageId="86" pageNumber="87">Golan Heights</location>
</taxonomicName>
; 
<date id="05073CC6A14BFF9216D3FA25564BF9D2" box="[706,851,1523,1550]" pageId="86" pageNumber="87" value="1996-05-01">
<collectingDate id="1543C52EA14BFF9216D3FA25564BF9D2" box="[706,851,1523,1550]" pageId="86" pageNumber="87" value="1996-05-01">1 May 1996</collectingDate>
</date>
; 
<collectorName id="DC4C7FD0A14BFF92174EFA2556ADF9D1" box="[863,949,1524,1550]" pageId="86" pageNumber="87">J. Levy</collectorName>
leg.; 
<specimenCode id="211FB27DA14BFF9217E6FA2551C4F9D1" box="[1015,1244,1524,1550]" pageId="86" pageNumber="87">HUJINV-Ar 21303</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="71061A06A14BFF9214FBF9E45444F990" blockId="86.[189,1046,1589,1651]" box="[234,348,1589,1615]" pageId="86" pageNumber="87">
<emphasis id="43CDC614A14BFF9214FBF9E45444F990" bold="true" box="[234,348,1589,1615]" pageId="86" pageNumber="87">
<typeStatus id="AE02A4A4A14BFF9214FBF9E45444F990" box="[234,348,1589,1615]" pageId="86" pageNumber="87" type="paratype">Paratype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="71061A06A14BFF9214ACF989510EF9AD" blockId="86.[189,1046,1589,1651]" box="[189,1046,1624,1651]" pageId="86" pageNumber="87">
<materialsCitation id="C1D1105BA14BFF9214ACF9895109F9AD" ID-GBIF-Occurrence="5839008520" box="[189,1041,1624,1651]" collectingDate="1996-05-01" collectorName="J. Levy" country="Israel" location="ISRAEL" pageId="86" pageNumber="87" specimenCode="HUJINV-Ar 21304" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<collectingCountry id="09AE5A96A14BFF9214ACF989543CF9AD" box="[189,292,1624,1650]" name="Israel" pageId="86" pageNumber="87">ISRAEL</collectingCountry>
• 
<specimenCount id="67BFD18FA14BFF92152AF989547FF9AC" box="[315,359,1624,1651]" count="1" pageId="86" pageNumber="87" type="female">1 ♀</specimenCount>
; same collection data as for holotype; 
<specimenCode id="211FB27DA14BFF92173CF9895109F9AD" box="[813,1041,1624,1650]" pageId="86" pageNumber="87">HUJINV-Ar 21304</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="71061A06A14BFF9214ACF94854E1F96C" blockId="86.[189,505,1689,1715]" box="[189,505,1689,1715]" pageId="86" pageNumber="87">
<heading id="2A4EAD6AA14BFF9214ACF94854E1F96C" bold="true" box="[189,505,1689,1715]" fontSize="11" level="3" pageId="86" pageNumber="87" reason="3">
<emphasis id="43CDC614A14BFF9214ACF94854E1F96C" bold="true" box="[189,505,1689,1715]" pageId="86" pageNumber="87">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="71061A06A14BFF9214ACF9195410F893" blockId="86.[189,1399,1736,1868]" pageId="86" pageNumber="87">
<materialsCitation id="C1D1105BA14BFF9214ACF919567EF93D" ID-GBIF-Occurrence="5839008317" box="[189,870,1736,1763]" collectingDate="1996-05-01" collectionCode="CMU" collectorName="J. Levy" country="Israel" location="ISRAEL" pageId="86" pageNumber="87" specimenCount="1" specimenCount-female="1">
<collectingCountry id="09AE5A96A14BFF9214ACF919543BF93D" box="[189,291,1736,1762]" name="Israel" pageId="86" pageNumber="87">ISRAEL</collectingCountry>
• 
<specimenCount id="67BFD18FA14BFF92152BF919547DF93C" box="[314,357,1736,1763]" count="1" pageId="86" pageNumber="87" type="female">1 ♀</specimenCount>
; same collection data as for holotype; 
<collectionCode id="17A882C3A14BFF921735F919567EF93D" box="[804,870,1736,1762]" pageId="86" pageNumber="87">CMU</collectionCode>
</materialsCitation>
• 
<materialsCitation id="C1D1105BA14BFF92176FF9195617F8D9" ID-GBIF-Occurrence="5839008488" collectingDate="2023-03-29" collectionCode="CMU" collectorName="S. Pekar &amp; V. Opatova" country="Israel" latitude="33.206" location="Odem Forest" longLatPrecision="72" longitude="35.736" municipality="Odem Forest" pageId="86" pageNumber="87" specimenCount="3" specimenCount-female="3">
<specimenCount id="67BFD18FA14BFF92176FF91956D8F93C" box="[894,960,1736,1763]" count="3" pageId="86" pageNumber="87" type="female">3 ♀♀</specimenCount>
; 
<taxonomicName id="B6B96185A14BFF9217DEF9195172F93C" authority="Forest" authorityName="Forest" box="[975,1130,1736,1763]" class="Arachnida" family="Zodariidae" genus="Acanthinozodium" kingdom="Animalia" order="Araneae" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="odem">
<collectingMunicipality id="9162807CA14BFF9217DEF9195172F93C" box="[975,1130,1736,1763]" pageId="86" pageNumber="87">Odem Forest</collectingMunicipality>
</taxonomicName>
; 
<geoCoordinate id="148D7CC1A14BFF921065F91951F7F93C" box="[1140,1263,1736,1763]" degrees="33.206" direction="north" orientation="latitude" pageId="86" pageNumber="87" precision="55" value="33.206">33.206° N</geoCoordinate>
, 
<geoCoordinate id="148D7CC1A14BFF9210EBF919506AF93C" box="[1274,1394,1736,1763]" degrees="35.736" direction="east" orientation="longitude" pageId="86" pageNumber="87" precision="55" value="35.736">35.736° E</geoCoordinate>
; 
<date id="05073CC6A14BFF9214ACF93D5446F8D9" box="[189,350,1772,1798]" pageId="86" pageNumber="87" value="2023-03-29">
<collectingDate id="1543C52EA14BFF9214ACF93D5446F8D9" box="[189,350,1772,1798]" pageId="86" pageNumber="87" value="2023-03-29">29 Mar. 2023</collectingDate>
</date>
; 
<collectorName id="DC4C7FD0A14BFF92157AF93D54D5F8DA" box="[363,461,1771,1798]" pageId="86" pageNumber="87">S. Pekár</collectorName>
and 
<collectorName id="DC4C7FD0A14BFF921617F93D5791F8D9" box="[518,649,1772,1798]" pageId="86" pageNumber="87">V. Opatová</collectorName>
leg.; 
<collectionCode id="17A882C3A14BFF9216DDF93D5617F8D9" box="[716,783,1772,1798]" pageId="86" pageNumber="87">CMU</collectionCode>
</materialsCitation>
• 
<materialsCitation id="C1D1105BA14BFF921738F93D54E2F8F6" ID-GBIF-Occurrence="5839008337" collectingDate="2023-05-04" collectionCode="CMU" collectorName="M. Rezac" country="Israel" latitude="33.206" longLatPrecision="72" longitude="35.736" pageId="86" pageNumber="87" specimenCount="1" specimenCount-female="1">
<specimenCount id="67BFD18FA14BFF921738F93D564EF8D9" box="[809,854,1772,1798]" count="1" pageId="86" pageNumber="87" type="female">1 ♀</specimenCount>
; same collection data as for preceding; 
<date id="05073CC6A14BFF92113BF93D55E3F8F6" pageId="86" pageNumber="87" value="2023-05-04">
<collectingDate id="1543C52EA14BFF92113BF93D55E3F8F6" pageId="86" pageNumber="87" value="2023-05-04">4 May 2023</collectingDate>
</date>
; 
<collectorName id="DC4C7FD0A14BFF921517F8DE546EF8F4" box="[262,374,1807,1835]" pageId="86" pageNumber="87">M. Řezáč</collectorName>
leg.; 
<collectionCode id="17A882C3A14BFF9215A6F8DE54E2F8F6" box="[439,506,1807,1833]" pageId="86" pageNumber="87">CMU</collectionCode>
</materialsCitation>
• 
<materialsCitation id="C1D1105BA14BFF921603F8DE541CF893" ID-GBIF-Occurrence="5839008321" collectingDate="2023-05-05" collectionCode="CMU" collectorName="M. Rezac" country="Israel" latitude="33.288" location="Har Hermon" longLatPrecision="72" longitude="35.764" pageId="86" pageNumber="87" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="67BFD18FA14BFF921603F8DE5727F8F6" box="[530,575,1807,1833]" count="1" pageId="86" pageNumber="87" type="male">1 ♂</specimenCount>
, 
<specimenCount id="67BFD18FA14BFF92165AF8DE576FF8F6" box="[587,631,1807,1833]" count="1" pageId="86" pageNumber="87" type="female">1 ♀</specimenCount>
; 
<location id="74664CDDA14BFF921697F8DE5604F8F6" LSID="urn:lsid:plazi:treatment:F910AB10A14BFF9D163FFCB75465FC39:74664CDDA14BFF921697F8DE5604F8F6" box="[646,796,1807,1833]" country="Israel" latitude="33.288" longLatPrecision="72" longitude="35.764" name="Har Hermon" pageId="86" pageNumber="87">Har Hermon</location>
; 
<geoCoordinate id="148D7CC1A14BFF921736F8DE56BAF8F6" box="[807,930,1807,1833]" degrees="33.288" direction="north" orientation="latitude" pageId="86" pageNumber="87" precision="55" value="33.288">33.288° N</geoCoordinate>
, 
<geoCoordinate id="148D7CC1A14BFF9217BFF8DF513DF8F6" box="[942,1061,1806,1833]" degrees="35.764" direction="east" orientation="longitude" pageId="86" pageNumber="87" precision="55" value="35.764">35.764° E</geoCoordinate>
; 
<date id="05073CC6A14BFF921020F8DF51D9F8F6" box="[1073,1217,1806,1833]" pageId="86" pageNumber="87" value="2023-05-05">
<collectingDate id="1543C52EA14BFF921020F8DF51D9F8F6" box="[1073,1217,1806,1833]" pageId="86" pageNumber="87" value="2023-05-05">5 May 2023</collectingDate>
</date>
; 
<collectorName id="DC4C7FD0A14BFF9210DCF8DE5024F8F4" box="[1229,1340,1807,1835]" pageId="86" pageNumber="87">M. Řezáč</collectorName>
leg.; 
<collectionCode id="17A882C3A14BFF9214ACF8E3541CF893" box="[189,260,1842,1868]" pageId="86" pageNumber="87">CMU</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="71061A06A14BFF9214ACF8A256E5F852" blockId="86.[189,1021,1907,1936]" box="[189,1021,1907,1936]" pageId="86" pageNumber="87">
<materialsCitation id="C1D1105BA14BFF9214ACF8A256E1F852" ID-GBIF-Occurrence="5839008510" box="[189,1017,1907,1936]" collectingDate="2001-03-28" collectionCode="CMU" collectorName="O. Rezac" country="Syria" location="Al-Zarah" pageId="86" pageNumber="87" specimenCount="1" specimenCount-male="1" stateProvince="Homs">
<collectingCountry id="09AE5A96A14BFF9214ACF8A2540DF852" box="[189,277,1907,1933]" name="Syria" pageId="86" pageNumber="87">SYRIA</collectingCountry>
• 
<specimenCount id="67BFD18FA14BFF92153DF8A25443F851" box="[300,347,1907,1934]" count="1" pageId="86" pageNumber="87" type="male">1 ♂</specimenCount>
; W 
<collectingRegion id="B37DD4E4A14BFF92159BF8A554CBF851" box="[394,467,1908,1934]" country="Syria" name="Hims" pageId="86" pageNumber="87">Homs</collectingRegion>
, 
<location id="74664CDDA14BFF9215CCF8A25754F852" LSID="urn:lsid:plazi:treatment:F910AB10A14BFF9D163FFCB75465FC39:74664CDDA14BFF9215CCF8A25754F852" box="[477,588,1907,1933]" country="Syria" name="Al-Zarah" pageId="86" pageNumber="87" stateProvince="Homs">Al-Zarah</location>
; 
<date id="05073CC6A14BFF921649F8A257E0F852" box="[600,760,1907,1934]" pageId="86" pageNumber="87" value="2001-03-28">
<collectingDate id="1543C52EA14BFF921649F8A257E0F852" box="[600,760,1907,1934]" pageId="86" pageNumber="87" value="2001-03-28">28 Mar. 2001</collectingDate>
</date>
; 
<collectorName id="DC4C7FD0A14BFF921714F8A25668F84F" box="[773,880,1907,1936]" pageId="86" pageNumber="87">O. Řezáč</collectorName>
leg.; 
<collectionCode id="17A882C3A14BFF9217A3F8A256E1F852" box="[946,1017,1907,1933]" pageId="86" pageNumber="87">CMU</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="39A3498DA14AFF9D14ACFED95157FCA9" lastPageId="89" lastPageNumber="90" pageId="87" pageNumber="88" type="description">
<paragraph id="71061A06A14AFF9314ACFED95456FEFD" blockId="87.[189,334,264,290]" box="[189,334,264,290]" pageId="87" pageNumber="88">
<heading id="2A4EAD6AA14AFF9314ACFED95456FEFD" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="87" pageNumber="88" reason="3">
<emphasis id="43CDC614A14AFF9314ACFED95456FEFD" bold="true" box="[189,334,264,290]" pageId="87" pageNumber="88">Description</emphasis>
</heading>
</paragraph>
<paragraph id="71061A06A14AFF9314FBFEE6578BFE8E" blockId="87.[189,1399,311,443]" box="[234,659,311,337]" pageId="87" pageNumber="88">
<emphasis id="43CDC614A14AFF9314FBFEE65432FE8E" bold="true" box="[234,298,311,337]" pageId="87" pageNumber="88">Male</emphasis>
(
<typeStatus id="AE02A4A4A14AFF931528FEE654BAFE8E" box="[313,418,311,337]" pageId="87" pageNumber="88" type="holotype">holotype</typeStatus>
HUJINV-Ar 21303)
</paragraph>
<paragraph id="71061A06A14AFF9314ACFE8A518BFE64" blockId="87.[189,1399,311,443]" pageId="87" pageNumber="88">
<smallCapsWord id="77E08CDAA14AFF9314ACFE8A5403FEAC" baselines="366,366" box="[189,283,347,373]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Habitus" pageId="87" pageNumber="88">HABITUS</smallCapsWord>
(
<figureCitation id="E9820683A14AFF93153AFE8B54A0FEAB" box="[299,440,346,372]" captionStart="Fig" captionStartId="87.[189,232,1927,1953]" captionTargetBox="[189,1398,468,1905]" captionTargetId="figure-104@87.[189,1398,468,1905]" captionTargetPageId="87" captionText="Fig. 67. Habitus of Zodarion levyi Shafaie &amp; Pekár sp. nov. A–B. Holotype, ♂ (HUJINV-Ar 21303). C–D. Paratype, ♀ (HUJINV-Ar 21304). A. Dorsal view. B. Ventral view. C. Dorsal view. D. Ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.17245254" httpUri="https://zenodo.org/record/17245254/files/figure.png" pageId="87" pageNumber="88">Fig. 67A–B</figureCitation>
). Carapace and chelicerae light brown; mouthparts basally light brown and apically pale; sternum yellow. All legs yellow. Palpal segments yellow. Abdomen dorsally black, with a shiny, black scutum, venter of abdomen and spinnerets slightly dirty yellow; PVS present.
</paragraph>
<caption id="25C64A8EA14AFF9314ACF85654CEF837" ID-DOI="http://doi.org/10.5281/zenodo.17245254" ID-Zenodo-Dep="17245254" httpUri="https://zenodo.org/record/17245254/files/figure.png" pageId="87" pageNumber="88" startId="87.[189,232,1927,1953]" targetBox="[189,1398,468,1905]" targetPageId="87">
<paragraph id="71061A06A14AFF9314ACF85654CEF837" blockId="87.[189,1399,1927,2024]" pageId="87" pageNumber="88">
<emphasis id="43CDC614A14AFF9314ACF8565403F87E" bold="true" box="[189,283,1927,1953]" pageId="87" pageNumber="88">Fig. 67.</emphasis>
Habitus of 
<taxonomicName id="B6B96185A14AFF9315BAF856563DF87E" authority="Shafaie &amp; Pekar" authorityName="Shafaie &amp; Pekar" authorityYear="2025" box="[427,805,1927,1953]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="levyi" status="sp. nov.">
<emphasis id="43CDC614A14AFF9315BAF8565742F87E" box="[427,602,1927,1953]" italics="true" pageId="87" pageNumber="88">Zodarion levyi</emphasis>
Shafaie &amp; Pekár
</taxonomicName>
<taxonomicNameLabel id="58FE7B6FA14AFF93173FF8595693F87D" box="[814,907,1928,1954]" pageId="87" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="43CDC614A14AFF931784F85656D3F87E" bold="true" box="[917,971,1927,1953]" pageId="87" pageNumber="88">A–B</emphasis>
. Holotype, ♂ (HUJINV-Ar 21303). 
<emphasis id="43CDC614A14AFF9314ACF87B55EEF81B" bold="true" box="[189,246,1962,1988]" pageId="87" pageNumber="88">C–D</emphasis>
. Paratype, ♀ (HUJINV-Ar 21304). 
<emphasis id="43CDC614A14AFF9316B7F87B57A3F81B" bold="true" box="[678,699,1962,1988]" pageId="87" pageNumber="88">A</emphasis>
. Dorsal view. 
<emphasis id="43CDC614A14AFF93177FF87A5698F81A" bold="true" box="[878,896,1963,1989]" pageId="87" pageNumber="88">B</emphasis>
. Ventral view. 
<emphasis id="43CDC614A14AFF931026F87B5153F81B" bold="true" box="[1079,1099,1962,1988]" pageId="87" pageNumber="88">C</emphasis>
. Dorsal view. 
<emphasis id="43CDC614A14AFF9310ECF87A5009F81A" bold="true" box="[1277,1297,1963,1989]" pageId="87" pageNumber="88">D</emphasis>
. Ventral view. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="71061A06A145FF9C14ACFED85115FE6F" blockId="88.[189,1400,265,433]" pageId="88" pageNumber="89">
<smallCapsWord id="77E08CDAA145FF9C14ACFED85468FEFE" baselines="284,284" box="[189,368,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="88" pageNumber="89">MEASUREMENTS</smallCapsWord>
. Total length 2.44; carapace 1.25 long, 0.89 wide. Clypeus 0.14 high. Eye sizes and interdistances: AME 0.13, ALE=PME 0.09, PLE 0.08, AME–AME 0.05, AME–ALE 0.03, ALE–ALE 0.29, AME–PME 0.05, PME–PME 0.33, PME–PLE 0.05, PLE–PLE 0.26, ALE–PLE 0.1. Legs: I 3.55 (0.99, 0.39, 0.72, 0.89, 0.56), II 2.97 (0.82, 0.36, 0.52, 0.75, 0.52), III 2.89 (0.79, 0.33, 0.56, 0.75, 0.46), IV 4.31 (1.15, 0.36, 0.92, 1.22, 0.66). Palp: 0.89 (0.29, 0.13, 0.05, 0.42).
</paragraph>
<paragraph id="71061A06A145FF9D14ACFE0957B5FEFC" blockId="88.[188,1398,471,533]" lastBlockId="89.[189,1398,265,362]" lastPageId="89" lastPageNumber="90" pageId="88" pageNumber="89">
<smallCapsWord id="77E08CDAA145FF9C14ACFE0955EBFE2F" baselines="491,491" box="[189,243,472,498]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Palp" pageId="88" pageNumber="89">PALP</smallCapsWord>
(
<figureCitation id="E9820683A145FF9C1512FE065489FE2E" box="[259,401,471,497]" captionStart="Fig" captionStartId="88.[189,233,1928,1954]" captionTargetBox="[189,1385,573,1890]" captionTargetId="figure-181@88.[189,1398,573,1903]" captionTargetPageId="88" captionText="Fig. 68. Male palp of Zodarion levyi Shafaie &amp; Pekár sp. nov., holotype (HUJINV-Ar 21303). A. Prolateral view. B. Ventral view. C. Retrolateral view. D. Prolateral view. E. Ventral view. F. Retrolateral view. G. Embolar tip, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.17245256" httpUri="https://zenodo.org/record/17245256/files/figure.png" pageId="88" pageNumber="89">Fig. 68A–G</figureCitation>
). RTA broad, with a claw-shaped extension. Median apophysis broad, rectangular, with an oval leaf-shaped PMA. Embolus short, with a bifurcated tip including two convergent arms. 
<emphasis id="43CDC614A144FF9D14FBFED8545DFEFC" bold="true" box="[234,325,265,291]" pageId="89" pageNumber="90">Female</emphasis>
(
<typeStatus id="AE02A4A4A144FF9D1544FED854A4FEFC" box="[341,444,265,291]" pageId="89" pageNumber="90" type="paratype">paratype</typeStatus>
HUJINV-Ar 21304)
</paragraph>
<caption id="25C64A8EA145FF9C14ACF85956E4F836" ID-DOI="http://doi.org/10.5281/zenodo.17245256" ID-Zenodo-Dep="17245256" httpUri="https://zenodo.org/record/17245256/files/figure.png" pageId="88" pageNumber="89" startId="88.[189,233,1928,1954]" targetBox="[189,1385,573,1890]" targetPageId="88">
<paragraph id="71061A06A145FF9C14ACF85956E4F836" blockId="88.[189,1398,1928,2025]" pageId="88" pageNumber="89">
<emphasis id="43CDC614A145FF9C14ACF8595439F87D" bold="true" box="[189,289,1928,1954]" pageId="88" pageNumber="89">Fig. 68.</emphasis>
Male palp of 
<taxonomicName id="B6B96185A145FF9C15CEF859566BF87D" authority="Shafaie &amp; Pekar" authorityName="Shafaie &amp; Pekar" authorityYear="2025" box="[479,883,1928,1954]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="levyi" status="sp. nov.">
<emphasis id="43CDC614A145FF9C15CEF859578EF87D" box="[479,662,1928,1954]" italics="true" pageId="88" pageNumber="89">Zodarion levyi</emphasis>
Shafaie &amp; Pekár
</taxonomicName>
<taxonomicNameLabel id="58FE7B6FA145FF9C176EF85856FDF87C" box="[895,997,1929,1955]" pageId="88" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (HUJINV-Ar 21303). 
<emphasis id="43CDC614A145FF9C14ACF87A55CAF81A" bold="true" box="[189,210,1963,1989]" pageId="88" pageNumber="89">A</emphasis>
. Prolateral view. 
<emphasis id="43CDC614A145FF9C15A8F87D54D5F819" bold="true" box="[441,461,1964,1990]" pageId="88" pageNumber="89">B</emphasis>
. Ventral view. 
<emphasis id="43CDC614A145FF9C1684F87A57B2F81A" bold="true" box="[661,682,1963,1989]" pageId="88" pageNumber="89">C</emphasis>
. Retrolateral view. 
<emphasis id="43CDC614A145FF9C17BAF87D56D8F819" bold="true" box="[939,960,1964,1990]" pageId="88" pageNumber="89">D</emphasis>
. Prolateral view. 
<emphasis id="43CDC614A145FF9C10B9F87D51A4F819" bold="true" box="[1192,1212,1964,1990]" pageId="88" pageNumber="89">E</emphasis>
. Ventral view. 
<emphasis id="43CDC614A145FF9C14ACF81E55D7F836" bold="true" box="[189,207,1999,2025]" pageId="88" pageNumber="89">F</emphasis>
. Retrolateral view. 
<emphasis id="43CDC614A145FF9C15A4F81F54D4F837" bold="true" box="[437,460,1998,2024]" pageId="88" pageNumber="89">G</emphasis>
. Embolar tip, ventral view. Scale bars: 0.2 mm.
</paragraph>
</caption>
<paragraph id="71061A06A144FF9D14ACFEFC5193FEB5" blockId="89.[189,1398,265,362]" pageId="89" pageNumber="90">
<smallCapsWord id="77E08CDAA144FF9D14ACFEFC5404FE9A" baselines="320,320" box="[189,284,301,327]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Habitus" pageId="89" pageNumber="90">HABITUS</smallCapsWord>
(
<figureCitation id="E9820683A144FF9D153CFEFD54A4FE99" box="[301,444,300,326]" captionStart="Fig" captionStartId="87.[189,232,1927,1953]" captionTargetBox="[189,1398,468,1905]" captionTargetId="figure-104@87.[189,1398,468,1905]" captionTargetPageId="87" captionText="Fig. 67. Habitus of Zodarion levyi Shafaie &amp; Pekár sp. nov. A–B. Holotype, ♂ (HUJINV-Ar 21303). C–D. Paratype, ♀ (HUJINV-Ar 21304). A. Dorsal view. B. Ventral view. C. Dorsal view. D. Ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.17245254" httpUri="https://zenodo.org/record/17245254/files/figure.png" pageId="89" pageNumber="90">Fig. 67C–D</figureCitation>
). Carapace, chelicerae and palp yellow; mouthparts, sternum, spinnerets, legs and abdomen ventrally pale. Abdomen dorsum brownish without scutum; PVS present.
</paragraph>
<paragraph id="71061A06A144FF9D14ACFE405115FDE8" blockId="89.[189,1400,401,568]" pageId="89" pageNumber="90">
<smallCapsWord id="77E08CDAA144FF9D14ACFE405469FE76" baselines="420,420" box="[189,369,401,427]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="89" pageNumber="90">MEASUREMENTS</smallCapsWord>
. Total length 2.9; carapace 1.25 long, 0.89 wide. Clypeus 0.14 high. Eye sizes and interdistances: AME 0.12, ALE= PME 0.1, PLE 0.09, AME–AME 0.09, AME–ALE 0.03, ALE–ALE 0.36, AME–PME 0.09, PME–PME 0.39, PME–PLE 0.09, PLE–PLE 0.23, ALE–PLE 0.15. Legs: I 3.25 (0.89, 0.36, 0.62, 0.79, 0.59), II 2.85 (0.75, 0.29, 0.59, 0.66, 0.56), III 2.98 (0.79, 0.39, 0.52, 0.79, 0.49), IV 4.28 (1.32, 0.46, 0.66, 1.25, 0.59). Palp: 0.98 (0.31, 0.14, 0.07, 0.46).
</paragraph>
<paragraph id="71061A06A144FF9D14ACFDB15673FD3C" blockId="89.[189,1399,607,739]" pageId="89" pageNumber="90">
<smallCapsWord id="77E08CDAA144FF9D14ACFDB15405FDA7" baselines="627,627" box="[189,285,608,634]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Epigyne" pageId="89" pageNumber="90">EPIGYNE</smallCapsWord>
(
<figureCitation id="E9820683A144FF9D1523FD8E54D4FDA6" box="[306,460,607,633]" captionStart="Fig" captionStartId="89.[189,232,1889,1915]" captionTargetBox="[193,1394,1047,1827]" captionTargetId="figure-239@89.[189,1398,1043,1851]" captionTargetPageId="89" captionText="Fig. 69. Female copulatory organ of Zodarion levyi Shafaie &amp; Pekár sp. nov., paratype (HUJINV-Ar 21304). A. Intact epigyne, ventral view. B. Epigyne, ventral view. C. Vulva, ventral view. D. Vulva, ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.17245258" httpUri="https://zenodo.org/record/17245258/files/figure.png" pageId="89" pageNumber="90">Fig. 69A–D</figureCitation>
). Epigyne comprising a clearly distinctive, transverse brace-shaped median septum, with deep copulatory openings, anteriorly covered by a hooded, emarginated arch and laterally surrounded by coiled spermathecae running divergently, visible by transparency. Vulva comprises divergent spermathecal ducts terminally expanded inward.
</paragraph>
<paragraph id="71061A06A144FF9D14ACFCDB5471FCFB" blockId="89.[189,361,778,804]" box="[189,361,778,804]" pageId="89" pageNumber="90">
<heading id="2A4EAD6AA144FF9D14ACFCDB5471FCFB" bold="true" box="[189,361,778,804]" fontSize="11" level="3" pageId="89" pageNumber="90" reason="3">
<emphasis id="43CDC614A144FF9D14ACFCDB5471FCFB" bold="true" box="[189,361,778,804]" pageId="89" pageNumber="90">Size variation</emphasis>
</heading>
</paragraph>
<paragraph id="71061A06A144FF9D14ACFCE85157FCA9" blockId="89.[189,1398,824,887]" pageId="89" pageNumber="90">
Total body length 2.44–3.06, carapace length/width 1.25–1.53/ 
<quantity id="B641B7E3A144FF9D17B8FCE95158FC8C" box="[937,1088,824,851]" metricMagnitude="-2" metricUnit="m" metricValue="2.4765" metricValueMax="2.6924" metricValueMin="2.2606" pageId="89" pageNumber="90" unit="in" value="0.975" valueMax="1.06" valueMin="0.89">0.89–1.06 in</quantity>
males (N =2). Total body length 2.8–2.9, carapace length/width 1.25–1.33/ 
<quantity id="B641B7E3A144FF9D16EBFC8D5689FCA9" box="[762,913,860,887]" metricMagnitude="-2" metricUnit="m" metricValue="2.2225" metricValueMax="2.2606" metricValueMin="2.1844" pageId="89" pageNumber="90" unit="in" value="0.875" valueMax="0.89" valueMin="0.86">0.86–0.89 in</quantity>
females (N=2).
</paragraph>
</subSubSection>
<subSubSection id="39A3498DA144FF9D14ACFC4C5465FC39" box="[189,381,925,998]" pageId="89" pageNumber="90" type="distribution">
<paragraph id="71061A06A144FF9D14ACFC4C544EFC68" blockId="89.[189,342,925,951]" box="[189,342,925,951]" pageId="89" pageNumber="90">
<heading id="2A4EAD6AA144FF9D14ACFC4C544EFC68" bold="true" box="[189,342,925,951]" fontSize="11" level="3" pageId="89" pageNumber="90" reason="3">
<emphasis id="43CDC614A144FF9D14ACFC4C544EFC68" bold="true" box="[189,342,925,951]" pageId="89" pageNumber="90">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="71061A06A144FF9D14ACFC1D5465FC39" blockId="89.[189,381,972,998]" box="[189,381,972,998]" pageId="89" pageNumber="90">
<collectingCountry id="09AE5A96A144FF9D14ACFC1D55E6FC39" box="[189,254,972,998]" name="Israel" pageId="89" pageNumber="90">Israel</collectingCountry>
and 
<collectingCountry id="09AE5A96A144FF9D1526FC1D5461FC39" box="[311,377,972,998]" name="Syria" pageId="89" pageNumber="90">Syria</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>