The comparative study of Alkanna and Huynhia (Boraginaceae, Lithospermeae), with description of a new species in Iran Author Khalvati, Sina 0000-0002-1782-6672 Department of Biology, Herbarium Division, Bu-Ali Sina University, P. O. Box 65175 / 4111, Hamedan, Iran. & sinakhalvati @ gmail. com; https: // orcid. org / 0000 - 0002 - 1782 - 6672 sinakhalvati@gmail.com Author Ranjbar, Massoud 0000-0003-0939-3878 Department of Biology, Herbarium Division, Bu-Ali Sina University, P. O. Box 65175 / 4111, Hamedan, Iran. & ranjbar @ basu. ac. ir; https: // orcid. org / 0000 - 0003 - 0939 - 3878 ranjbar@basu.ac.ir text Phytotaxa 2021 2021-11-09 524 4 216 234 journal article 3654 10.11646/phytotaxa.524.4.1 c7ecb63e-4d46-4af3-b512-cb76ba9e0caa 1179-3163 5656352 1. Alkanna frigida Boissier (1846: 32) . Type: — IRAN . Kohgiluyeh and Boyer-Ahmad : Dena Mt. [In argillosis humidis alpis Kuh-Daëna post nivem deliquescentem], 8 July 1842 , Kotschy 573 ( holotype G ! [G-00176782]; isotype BM ! [ BM013839850 ], C ! [ C10008670 ], E ! [ E00284793 ], FI ! [ FI 009503], GOET ! [ GOET005793 ], H ! [ H1613716 ], K ! [K-001085737], MO ! [MO-503924], P ! [ P00606828 ], US! [ US 01050205], W ! [W-0051322], WAG ! [ WAG0058072 ]). Fig. 9 . = Alkanna frigida variety pallida Bornmüller (1907: 784) . Type: — IRAN . Tehran : Tochal Mt. [ M . Elburs , in jugo Lädd alpium Totschal], 3000 m , 4 June 1902 , Bornmüller 7703 ( holotype JE ! [JE-00012012]) . FIGURE 9. Alkanna frigida Boiss. ( Kotschy 573 , G) (reproduced with permission). Description: —Perennial, root rather thick, stem 3–10 (–15) cm long, multi-branched, branched from base, densely patent-pilose. Basal leaves oblong to ovate, 30–100 × 5–12 mm , acute to sub-acute, tapering into the petiole, with entire, flat, or sometimes slightly undulate margins, white densely tuberculate setose. Cauline leaves oblong to linear, 10–20 × 3–5 mm , acute or subacute, sessile, attenuate at base, with an evident midrib, veinless, densely white setose. Inflorescence a compact terminal head; pedicels short, leafy, peduncle 1–2 cm long; bracts oblong to ovate, 5–10 × 2–5 mm , densely to sparsely tuberculate setiform and pilose. Pedicels erect, 1–2 mm long at flowering, recurved 2–3 mm long in fruit and pilose. Calyx 3–5 mm long in flowering time, densely hirsute, and eglandular hairy. Corolla sulphur yellow, occasionally with purple spots, 10–12 (–15) mm long, glabrous outside, limb 3–5 mm in diam., tube 2–3 mm wide, and 6–10 mm long. Nutlets 5–6 × 3–4 mm , densely tuberculate; beak horizontal to strongly curved. Additional specimens examined: — IRAN . Alborz : Kandavan , 36º 5’ N , 51º 11’ E , s.d., Wendelbo & Shirdelpour 11650 ( TARI ) ; Chaharmahal and Bakhtiari : Lordegan , between Monj & Char-Tagh , Tang-e Zendan , 31º 30’ N , 50º 49’ E , s.d., Mozaffarian 59924 ( TARI ) ; Isfahan : Semirom , Khafr , Dena Mt. , 30º 58’ N , 51º 30’ E , s.d., Riazi 5982 ( TARI ) ; Ghazvin : Alamut , 36º 31’ N , 50º 28’ E , s.d., Mirfakhrayi 1382 ( TARI ) ; Semnan : 35 km north of Damghan , Touyeh , 36º 2’ N , 53º 51’ E , s.d., Wendelbo & Assadi 29488 ( TARI ) ; Tehran : Damavand , Valiran village , 35º 40’ N , 52º 5’ E , s.d., Mozaffarian 32345 ( TARI ) .