Taxonomic revision of the genus Nychiodes Lederer, 1853 (Geometridae: Ennominae: Boarmiini) with description of three new species-an integrative approach Author Wanke, Dominic Author Hausmann, Axel Author Krogmann, Lars Author Petrányi, Gergely Author Rajaei, Hossein text Zootaxa 2020 2020-07-14 4812 1 1 61 journal article 10.11646/zootaxa.4812.1.1 1175-5326 3944102 74FE6D6A-B41D-4FC9-BC26-B07DDEB9A0F6 The waltheri species-group The species of this group can be diagnosed by the following characters (after Müller et al. 2019 ): In male genitalia, the costa of the valva is sclerotized up to the subapical part, humped medially (in most species); apex of ampulla superior and ampulla inferior spinose; sacculus process strongly pronounced, slender and pointed ( Müller et al. 2019 ); tip of aedeagus with digitiform extension (straight or curved). In female genitalia, ductus bursae shorter than half length of corpus bursae; lamella postvaginalis conically broadened, strongly sclerotized. The following species are included in this group: N. waltheri Wagner, 1919 , N. palaestinensis Wagner, 1919 , N. muelleri Hausmann, 1991 and N. aphrodite Hausmann & Wimmer, 1994 . Remarks. Two species, N. dalmatina and N. waltheri were included in the recent revision of European species ( Müller et al. 2019 ). However, as N. waltheri is the iconic species of this group and mainly distributed outside Europe, we included this species also to the present study.