<document id="5026076820297C451AC4F287CA5F3230" ID-CLB-Dataset="39184" ID-DOI="10.11646/zootaxa.4105.5.2" ID-GBIF-Dataset="60ca4dbd-002b-4ca6-999d-9174f9b02324" ID-ISSN="1175-5326" ID-Zenodo-Dep="255425" ID-ZooBank="52401DF9-2901-4AF6-BA17-696D39280B3C" IM.bibliography_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1461736713249" checkinUser="plazi" docAuthor="Sanborn, Allen F." docDate="2016" docId="E338D67535102169FF05F97C72A76F79" docLanguage="en" docName="zootaxa.4105.5.2.pdf.imf" docOrigin="Zootaxa 4105 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Guyalna panamensis, n. comb" docType="treatment" docVersion="23" lastPageNumber="445" masterDocId="1F01AE0D351E2166FF92FF9876486E39" masterDocTitle="The cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Cicadinae: Fidicinini): generic description, twelve new combinations, and a key to species" masterLastPageNumber="454" masterPageNumber="430" pageNumber="444" updateTime="1732490992240" updateUser="ExternalLinkService">
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<mods:title id="32EB2F293079EE90CC00B2407E61CF18">The cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Cicadinae: Fidicinini): generic description, twelve new combinations, and a key to species</mods:title>
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<mods:namePart id="4B3C30EA2133D46871C5BDE66E172034">Sanborn, Allen F.</mods:namePart>
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<treatment id="E338D67535102169FF05F97C72A76F79" ID-DOI="http://doi.org/10.5281/zenodo.4951946" ID-GBIF-Taxon="119651802" ID-Zenodo-Dep="4951946" LSID="urn:lsid:plazi:treatment:E338D67535102169FF05F97C72A76F79" httpUri="http://treatment.plazi.org/id/E338D67535102169FF05F97C72A76F79" lastPageId="15" lastPageNumber="445" pageId="14" pageNumber="444" scope_family="Cicadidae" scope_order="Hemiptera" scope_subFamily="Cicadinae" scope_tribe="Fidicinini">
<subSubSection id="238B34E835102168FF05F97C74F968C7" box="[151,689,1764,1790]" pageId="14" pageNumber="444" type="nomenclature">
<paragraph id="6B2E676335102168FF05F97C74F968C7" blockId="14.[151,689,1764,1824]" box="[151,689,1764,1790]" pageId="14" pageNumber="444">
<heading id="3066D00F35102168FF05F97C74F968C7" bold="true" box="[151,689,1764,1790]" fontSize="11" level="1" pageId="14" pageNumber="444" reason="1">
<emphasis id="59E5BB7135102168FF05F97C74F968C7" bold="true" box="[151,689,1764,1790]" pageId="14" pageNumber="444">
<taxonomicName id="AC911CE035102168FF05F97C740868C7" ID-CoL="6KV9V" authority="(Davis, 1939)" baseAuthorityName="Davis" baseAuthorityYear="1939" box="[151,576,1764,1790]" class="Insecta" family="Cicadidae" genus="Guyalna" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="444" phylum="Arthropoda" rank="species" species="panamensis" status="comb. nov.">
<emphasis id="59E5BB7135102168FF05F97C77DD68C7" bold="true" box="[151,405,1764,1790]" italics="true" pageId="14" pageNumber="444">Guyalna panamensis</emphasis>
(
<bibRefCitation id="0F001A9235102168FE37F97C747F68C7" author="Davis, W. T." box="[421,567,1764,1790]" firstAuthor="Davis" journalOrPublisher="Journal of the New York Entomological Society" pageId="14" pageNumber="444" pagination="287 - 302" part="47" refId="ref12817" refString="Davis, W. T. (1939) New cicadas from North Carolina and the West Indies. Journal of the New York Entomological Society, 47, 287 - 302." title="New cicadas from North Carolina and the West Indies" type="journal article" year="1939">Davis, 1939</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="42D6060A35102168FDD5F97C74E568C7" box="[583,685,1764,1790]" pageId="14" pageNumber="444">n. comb</taxonomicNameLabel>
.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="238B34E835102168FF05F89F76B16919" box="[151,249,1799,1824]" pageId="14" pageNumber="444" type="description">
<paragraph id="6B2E676335102168FF05F89F76B16919" blockId="14.[151,689,1764,1824]" box="[151,249,1799,1824]" pageId="14" pageNumber="444">
(
<figureCitation id="F3AA7BE635102168FF0DF89F76B96919" box="[159,241,1799,1824]" captionStart="FIGURE 19" captionStartId="15.[151,250,926,948]" captionTargetBox="[151,1435,375,904]" captionTargetId="figure@15.[151,1436,375,904]" captionTargetPageId="15" captionText="FIGURE 19. Guyalna panamensis (Davis, 1939) n. comb. A, male, Panama (AFSC, compared to holotype specimen). Scale bar: 2 cm; B, timbal cover. Scale bar: 2 mm." httpUri="https://zenodo.org/record/255444/files/figure.png" pageId="14" pageNumber="444">Fig. 19</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="238B34E835102168FF05F8D7747F69BD" pageId="14" pageNumber="444" type="reference_group">
<paragraph id="6B2E676335102168FF05F8D77457695C" blockId="14.[151,567,1871,1924]" box="[151,543,1871,1893]" pageId="14" pageNumber="444">
<treatmentCitationGroup id="4B81404D35102168FF05F8D77457695C" box="[151,543,1871,1893]" pageId="14" pageNumber="444">
<taxonomicName id="AC911CE035102168FF05F8D77453695C" ID-CoL="6HZ2Z" authority="Davis, 1939" authorityName="Davis" authorityPageNumber="288" authorityYear="1939" box="[151,539,1871,1893]" class="Insecta" family="Cicadidae" genus="Fidicina" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="444" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="59E5BB7135102168FF05F8D77723695C" box="[151,363,1871,1893]" italics="true" pageId="14" pageNumber="444">Fidicina panamensis</emphasis>
<treatmentCitation id="EA30417235102168FEE3F8D77453695C" author="Davis" box="[369,539,1871,1893]" page="288" pageId="14" pageNumber="444" year="1939">
<bibRefCitation id="0F001A9235102168FEE3F8D77453695C" author="Davis, W. T." box="[369,539,1871,1893]" firstAuthor="Davis" journalOrPublisher="Journal of the New York Entomological Society" page="288" pageId="14" pageNumber="444" pagination="287 - 302" part="47" refId="ref12817" refString="Davis, W. T. (1939) New cicadas from North Carolina and the West Indies. Journal of the New York Entomological Society, 47, 287 - 302." title="New cicadas from North Carolina and the West Indies" type="journal article" year="1939">Davis 1939: 288</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="6B2E676335102168FF05F8F7747F69BD" blockId="14.[151,567,1871,1924]" box="[151,567,1902,1924]" pageId="14" pageNumber="444">
<treatmentCitationGroup id="4B81404D35102168FF05F8F7747F69BD" box="[151,567,1902,1924]" pageId="14" pageNumber="444">
<taxonomicName id="AC911CE035102168FF05F8F7747A69BD" ID-CoL="8HX5L" authority="Wolda 1977: 244" authorityName="Wolda" authorityPageNumber="244" authorityYear="1977" baseAuthorityName="Davis" baseAuthorityYear="1939" box="[151,562,1902,1924]" class="Insecta" family="Cicadidae" genus="Dorisiana" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="444" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="59E5BB7135102168FF05F8F7773369BD" box="[151,379,1903,1924]" italics="true" pageId="14" pageNumber="444">Dorisiana panamensis</emphasis>
<treatmentCitation id="EA30417235102168FE10F8F6747A69BD" author="Wolda" box="[386,562,1902,1924]" page="244" pageId="14" pageNumber="444" year="1977">
<bibRefCitation id="0F001A9235102168FE10F8F6747A69BD" author="Wolda, H." box="[386,562,1902,1924]" firstAuthor="Wolda" journalOrPublisher="Geo-Eco-Trop" page="244" pageId="14" pageNumber="444" pagination="229 - 257" part="3" refId="ref14908" refString="Wolda, H. (1977) Fluctuations in abundance of some Homoptera in a neotropical forest. Geo-Eco-Trop, 3, 229 - 257." title="Fluctuations in abundance of some Homoptera in a neotropical forest" type="journal article" year="1977">Wolda 1977: 244</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="238B34E835102168FF05F82874E369F1" box="[151,683,1968,1993]" pageId="14" pageNumber="444" type="materials_examined">
<paragraph id="6B2E676335102168FF05F82874E369F1" blockId="14.[151,1436,1968,2029]" box="[151,683,1968,1993]" pageId="14" pageNumber="444">
<materialsCitation id="DBF96D3E35102168FF05F82874E369F1" ID-GBIF-Occurrence="4115800313" box="[151,683,1968,1993]" country="Panama" location="Tabernilla" pageId="14" pageNumber="444" specimenCount="1" stateProvince="Canal Zone" typeStatus="type">
<emphasis id="59E5BB7135102168FF05F828777D69F0" bold="true" box="[151,309,1968,1993]" pageId="14" pageNumber="444">
<typeStatus id="B42AD9C135102168FF05F828769A69F0" box="[151,210,1968,1993]" pageId="14" pageNumber="444">Type</typeStatus>
locality.
</emphasis>
<location id="6E4E31B835102168FEAEF82877F869F0" LSID="urn:lsid:plazi:treatment:E338D67535102169FF05F97C72A76F79:6E4E31B835102168FEAEF82877F869F0" box="[316,432,1968,1993]" country="Panama" name="Tabernilla" pageId="14" pageNumber="444" stateProvince="Canal Zone">Tabernilla</location>
, 
<collectingRegion id="A955A98135102168FE29F828747769F1" box="[443,575,1968,1993]" pageId="14" pageNumber="444">Canal Zone</collectingRegion>
, 
<collectingCountry id="138627F335102168FDD9F82974EF69F1" box="[587,679,1969,1992]" name="Panama" pageId="14" pageNumber="444">Panama</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="238B34E835102169FF55F84C772C6F25" lastPageId="15" lastPageNumber="445" pageId="14" pageNumber="444" type="discussion">
<paragraph id="6B2E676335102169FF55F84C772C6F25" blockId="14.[151,1436,1968,2029]" lastBlockId="15.[151,1436,151,320]" lastPageId="15" lastPageNumber="445" pageId="14" pageNumber="444">
<emphasis id="59E5BB7135102168FF55F84C777369D4" bold="true" box="[199,315,2004,2029]" pageId="14" pageNumber="444">Remarks.</emphasis>
The species was last transferred to 
<taxonomicName id="AC911CE035102168FD6AF84E752269D4" box="[760,874,2006,2029]" class="Insecta" family="Cicadidae" genus="Dorisiana" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="59E5BB7135102168FD6AF84E752269D4" box="[760,874,2006,2029]" italics="true" pageId="14" pageNumber="444">Dorisiana</emphasis>
</taxonomicName>
without comment by 
<bibRefCitation id="0F001A9235102168FBE8F84C735669D4" author="Wolda, H." box="[1146,1310,2004,2029]" firstAuthor="Wolda" journalOrPublisher="Geo-Eco-Trop" pageId="14" pageNumber="444" pagination="229 - 257" part="3" refId="ref14908" refString="Wolda, H. (1977) Fluctuations in abundance of some Homoptera in a neotropical forest. Geo-Eco-Trop, 3, 229 - 257." title="Fluctuations in abundance of some Homoptera in a neotropical forest" type="journal article" year="1977">Wolda (1977)</bibRefCitation>
. However, 
<taxonomicName id="AC911CE035112169FF05FF0F76B06E89" box="[151,248,151,176]" class="Insecta" family="Cicadidae" genus="Guyalna" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="445" phylum="Arthropoda" rank="genus">
<emphasis id="59E5BB7135112169FF05FF0F76B06E89" box="[151,248,151,176]" italics="true" pageId="15" pageNumber="445">Guyalna</emphasis>
</taxonomicName>
did not exist at the time and the timbal cover fits the expanded description of 
<taxonomicName id="AC911CE035112169FBF3FF0F72896E89" box="[1121,1217,151,176]" class="Insecta" family="Cicadidae" genus="Guyalna" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="445" phylum="Arthropoda" rank="genus">
<emphasis id="59E5BB7135112169FBF3FF0F72896E89" box="[1121,1217,151,176]" italics="true" pageId="15" pageNumber="445">Guyalna</emphasis>
</taxonomicName>
provided here with the rounded apex and approximate parallel sides as it extends anteriorly (
<figureCitation id="F3AA7BE635112169FC5AFF2472536EED" box="[968,1051,188,213]" captionStart="FIGURE 19" captionStartId="15.[151,250,926,948]" captionTargetBox="[151,1435,375,904]" captionTargetId="figure@15.[151,1436,375,904]" captionTargetPageId="15" captionText="FIGURE 19. Guyalna panamensis (Davis, 1939) n. comb. A, male, Panama (AFSC, compared to holotype specimen). Scale bar: 2 cm; B, timbal cover. Scale bar: 2 mm." httpUri="https://zenodo.org/record/255444/files/figure.png" pageId="15" pageNumber="445">Fig. 19</figureCitation>
). As a result, the species is hereby transferred to 
<taxonomicName id="AC911CE035112169FEA6FF4777DC6EC1" box="[308,404,223,248]" class="Insecta" family="Cicadidae" genus="Guyalna" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="445" phylum="Arthropoda" rank="genus">
<emphasis id="59E5BB7135112169FEA6FF4777DC6EC1" box="[308,404,223,248]" italics="true" pageId="15" pageNumber="445">Guyalna</emphasis>
</taxonomicName>
to become 
<taxonomicName id="AC911CE035112169FD85FF4775EC6EC1" authority="Davis, 1939" authorityName="Davis" authorityYear="1939" box="[535,932,223,248]" class="Insecta" family="Cicadidae" genus="Guyalna" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="445" phylum="Arthropoda" rank="species" species="panamensis" status="comb. nov.">
<emphasis id="59E5BB7135112169FD85FF47754A6EC1" box="[535,770,223,248]" italics="true" pageId="15" pageNumber="445">Guyalna panamensis</emphasis>
(
<bibRefCitation id="0F001A9235112169FC80FF4775D46EC1" author="Davis, W. T." box="[786,924,223,248]" firstAuthor="Davis" journalOrPublisher="Journal of the New York Entomological Society" pageId="15" pageNumber="445" pagination="287 - 302" part="47" refId="ref12817" refString="Davis, W. T. (1939) New cicadas from North Carolina and the West Indies. Journal of the New York Entomological Society, 47, 287 - 302." title="New cicadas from North Carolina and the West Indies" type="journal article" year="1939">Davis, 1939</bibRefCitation>
)
</taxonomicName>
<emphasis id="59E5BB7135112169FC39FF4772596EC1" bold="true" box="[939,1041,223,248]" pageId="15" pageNumber="445">
<taxonomicNameLabel id="42D6060A35112169FC39FF4772446EC1" box="[939,1036,223,248]" pageId="15" pageNumber="445">n. comb</taxonomicNameLabel>
.
</emphasis>
Body length 
<quantity id="AC69CA8635112169FB3BFF78736C6ECE" box="[1193,1316,224,248]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.3" metricValueMin="2.2" pageId="15" pageNumber="445" unit="mm" value="22.5" valueMax="23.0" valueMin="22.0">22–23 mm</quantity>
, fore wing length 
<quantity id="AC69CA8635112169FF76FE9C77286F25" box="[228,352,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" metricValueMax="3.4" metricValueMin="3.2" pageId="15" pageNumber="445" unit="mm" value="33.0" valueMax="34.0" valueMin="32.0">32–34 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="238B34E835112169FF55FEBF72A76F79" box="[199,1263,295,320]" pageId="15" pageNumber="445" type="distribution">
<paragraph id="6B2E676335112169FF55FEBF72A76F79" blockId="15.[151,1436,151,320]" box="[199,1263,295,320]" pageId="15" pageNumber="445">
<emphasis id="59E5BB7135112169FF55FEBF77286F79" bold="true" box="[199,352,295,320]" pageId="15" pageNumber="445">Distribution.</emphasis>
The species has been reported only from 
<collectingCountry id="138627F335112169FCA6FEB075C56F06" box="[820,909,296,319]" name="Panama" pageId="15" pageNumber="445">Panama</collectingCountry>
(
<bibRefCitation id="0F001A9235112169FC09FEBF727E6F79" author="Metcalf, Z. P." box="[923,1078,295,320]" firstAuthor="Metcalf" journalOrPublisher="North Carolina State College Contribution" pageId="15" pageNumber="445" pagination="1 - 585" part="1502" refId="ref13609" refString="Metcalf, Z. P. (1963) General catalogue of the Homoptera, Fascicle VIII. Cicadoidea. Part 1. Cicadidae. Section I. Tibiceninae. North Carolina State College Contribution, 1502, i - vii, 1 - 585." title="General catalogue of the Homoptera, Fascicle VIII. Cicadoidea. Part 1. Cicadidae. Section I. Tibiceninae" type="journal article" year="1963">Metcalf 1963</bibRefCitation>
; 
<bibRefCitation id="0F001A9235112169FBD0FEBF72AB6F79" author="Sanborn, A. F." bookContentInfo="1001 pp." box="[1090,1251,295,320]" firstAuthor="Sanborn" journalOrPublisher="Elsevier / Academic Press, San Diego" pageId="15" pageNumber="445" refId="ref14140" refString="Sanborn, A. F. (2013) Catalogue of the Cicadoidea (Hemiptera: Cicadoidea). With contributions to the bibliography by Martin H. Villet. Elsevier / Academic Press, San Diego, 1001 pp." title="Catalogue of the Cicadoidea (Hemiptera: Cicadoidea). With contributions to the bibliography by Martin H. Villet" type="book" year="2013">Sanborn 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>