<document ID-CLB-Dataset="20126" ID-DOI="http://dx.doi.org/10.3897/afrinvertebr.63.76309" ID-GBIF-Dataset="4b66695a-c3d4-4711-8c87-00edd3951bf4" ID-Pensoft-Pub="2305-2562-1-19" ID-Pensoft-UUID="B0655A9F7BBC582691EC0F00E156BA38" ID-ZooBank="F849C700225A4923AE1962882F933E83" ModsDocID="2305-2562-63-1-19" checkinTime="1648229925038" checkinUser="pensoft" docAuthor="Boschert, Claire &amp; Dikow, Torsten" docDate="2022" docId="D5E81E9941B1515C813B4C0396816832" docLanguage="en" docName="AfrInvert 63(1): 19-75" docOrigin="African Invertebrates 63 (1)" docPubDate="2022-03-25" docSource="http://dx.doi.org/10.3897/afrinvertebr.63.76309" docTitle="Haplomydas Bezzi 1924" docType="treatment" docVersion="9" id="B0655A9F7BBC582691EC0F00E156BA38" lastPageNumber="19" masterDocId="B0655A9F7BBC582691EC0F00E156BA38" masterDocTitle="Taxonomic revision of the mydas-fly genera Eremohaplomydas Bequaert, 1959, Haplomydas Bezzi, 1924, and Lachnocorynus Hesse, 1969 (Insecta, Diptera, Mydidae)" masterLastPageNumber="75" masterPageNumber="19" pageNumber="19" updateTime="1732937927976" updateUser="ExternalLinkService">
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<mods:title id="5168CC02FDA563D4A5FCAED3EEAF6B62">Taxonomic revision of the mydas-fly genera Eremohaplomydas Bequaert, 1959, Haplomydas Bezzi, 1924, and Lachnocorynus Hesse, 1969 (Insecta, Diptera, Mydidae)</mods:title>
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<mods:namePart id="C9F3C20CBDBFCBF1FA74500F35CA46A4">Boschert, Claire</mods:namePart>
<mods:affiliation id="028FBC95E791C14CF39426249256BCBE">Rice University, Houston, TX, USA</mods:affiliation>
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<mods:namePart id="02636F2BDAFEF84396A9267D98DF46EA">Dikow, Torsten</mods:namePart>
<mods:nameIdentifier id="5784785A83849DC2003D2EE30AD02F13" type="ORCID">https://orcid.org/0000-0003-4816-2909</mods:nameIdentifier>
<mods:affiliation id="890CC77BA542CCBBC9E6EEDBF379F10B">Department of Entomology, National Museum of Natural History, Smithsonian Institution, 10 th Street and Constitution Avenue NW, Washington, DC 20560, USA</mods:affiliation>
<mods:nameIdentifier id="6012653D16485CDEBE76B077752E9E2F" type="email">dikowt@si.edu</mods:nameIdentifier>
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<mods:titleInfo id="2EC16B96206E74D95F25436DB0E96A39">
<mods:title id="E44BEB5F34766B2EA7CC112845531C06">African Invertebrates</mods:title>
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<mods:part id="0D6802708B8D40E5896A7D7DAA7ACACD">
<mods:date id="CFCA38AE39C8B6DF1E9ED18C43644EB1">2022</mods:date>
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<mods:number id="B3F995D09292A319F5E554A7E931862A">63</mods:number>
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<treatment id="D5E81E9941B1515C813B4C0396816832" ID-DOI="http://doi.org/10.5281/zenodo.7577352" ID-GBIF-Taxon="194257696" ID-Zenodo-Dep="7577352" LSID="urn:lsid:zoobank.org:act:AA86F72F-7319-43C5-9104-A618FA521E5E" httpUri="http://treatment.plazi.org/id/D5E81E9941B1515C813B4C0396816832" lastPageNumber="19" pageId="0" pageNumber="19" scope_class="Insecta" scope_family="Mydidae" scope_order="Diptera">
<subSubSection id="58FB9111F3054803ABD150534DB894DC" pageId="0" pageNumber="19" type="nomenclature">
<paragraph id="15AFAA3A3F19EFFD3A3C036C8DA1A9E2" pageId="0" pageNumber="19">
<taxonomicName id="17F5E81AE0632927198CE061CE023592" ID-CoL="4TGN" ID-ENA="2611090" LSID="http://zoobank.org/AA86F72F-7319-43C5-9104-A618FA521E5E" authority="Bezzi, 1924" authorityName="Bezzi" authorityYear="1924" class="Insecta" family="Mydidae" genus="Haplomydas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplomydas" order="Diptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">Haplomydas Bezzi, 1924</taxonomicName>
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<paragraph id="A3F9418B5210E4CA63D443F8AA04ED20">GBIF: https://www.gbif.org/species/1591511</paragraph>
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<treatmentCitation id="3091AA8790E82A895100007F5DA207BE" httpUri="https://www.biodiversitylibrary.org/page/40677714">
<taxonomicName id="39F8645AF7C98AD60C065ED716276636" ID-CoL="4TGN" authority="Bezzi, 1924: 199" authorityName="Bezzi" authorityPageNumber="199" authorityYear="1924" class="Insecta" family="Mydidae" genus="Haplomydas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplomydas" order="Diptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">Haplomydas Bezzi, 1924: 199</taxonomicName>
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.
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<paragraph id="C4214899CEB3B90BCFC32091AEEB2EF5" pageId="0" pageNumber="19">
<taxonomicName id="684DABCCBB8F88103AA6B5A871F8C6FB" authority="Seguy, 1929" authorityName="Seguy" authorityYear="1929" class="Insecta" family="Mydidae" genus="Heleomydas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heleomydas" order="Diptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
Heleomydas 
<normalizedToken id="E45BB5E79CE5C67D5F098DB7DB31FBD8" originalValue="Séguy">Seguy</normalizedToken>
, 1929
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- junior synonym; ZooBank http://zoobank.org/48330D1D-A176-4042-9F3C-97EC14FCD173
</paragraph>
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<paragraph id="7B08379405098325DCCD7862E63C8CE2" pageId="0" pageNumber="19">
Type-species: 
<taxonomicName id="8AA7DC4D9CA812B9BB978044BA5E79B9" authority="Bezzi, 1924" authorityName="Bezzi" authorityYear="1924" class="Insecta" family="Mydidae" genus="Haplomydas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplomydas crassipes" order="Diptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="crassipes">Haplomydas crassipes Bezzi, 1924</taxonomicName>
, by original designation.
</paragraph>
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<subSubSection id="58367FEAEC603BC8CBF5C1E73E17CB11" pageId="0" pageNumber="19" type="diagnosis">
<paragraph id="2CFE260D836A7D88357D0E530DCB0957" pageId="0" pageNumber="19">Diagnosis.</paragraph>
<paragraph id="6E759F02028F7A2D65AA697C819548F8" pageId="0" pageNumber="19">The genus can be delineated by the greatly expanded metathoracic femora, the distinct ventral keel terminating into a spur on the metathoracic tibiae, the presence of setae on the posterior anepisternum, the yellow to light brown colouration, and the absence of M3+M4 terminating into the costa.</paragraph>
</subSubSection>
<subSubSection id="ED0B9D5B4E74A51DC4CAB49EB6F9F64E" pageId="0" pageNumber="19" type="distribution">
<paragraph id="0160C75B018FF42AD62A5FDBA6CBEC7E" pageId="0" pageNumber="19">Distribution, biodiversity hotspots, phenology, and biology.</paragraph>
<paragraph id="1797E07A60D3FBE635FF9C303BEC73C0" pageId="0" pageNumber="19">
Known from diverse localities in Botswana, Mozambique, Namibia, and Zimbabwe (Fig. 
<figureCitation id="D7E868B93B665D10D063213367E7BF95" captionStart="Figure 57" captionStartId="F16" captionText="Figure 57. Map of southern Africa with elevational relief and biodiversity hotspots (sensu Conservation International in grey) and distribution of Eremohaplomydas stomachoris sp. nov., Eremohaplomydas whartoni sp. nov. (both localities only 5 km apart), and Haplomydas crassipes (SimpleMappr https: // www. simplemappr. net / map / 14090). Distribution and occurrence data available in Google Earth KML file https: // www. simplemappr. net / map / 14090. kml and also through GBIF (data-set https: // www. gbif. org / dataset / 993875 DD- 5915 - 4107 - 8707 - 835 D 5 A 8 D 1022, DOI https: // doi. org / 10.15468 / awpjz 9)." figureDoi="10.3897/afrinvertebr.63.76309.figure57" httpUri="https://binary.pensoft.net/fig/663594" pageId="0" pageNumber="19">57</figureCitation>
). A relatively commonly collected genus with collecting events between 1917 and 1999 (Table 
<tableCitation id="7016B084AD74E8071E3268096B648550" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Collecting event summary for Eremohaplomydas, Haplomydas, and Lachnocorynus species." httpUri="http://table.plazi.org/id/8DF9F799C695B7BAA951697DD1292320" pageId="0" pageNumber="19" tableUuid="8DF9F799C695B7BAA951697DD1292320">1</tableCitation>
). The genus occurs in the Eastern Afromontane biodiversity hotspot in eastern-most Zimbabwe (Fig. 
<figureCitation id="85D8AF3DFC54DC483CD940306B0618C5" captionStart="Figure 57" captionStartId="F16" captionText="Figure 57. Map of southern Africa with elevational relief and biodiversity hotspots (sensu Conservation International in grey) and distribution of Eremohaplomydas stomachoris sp. nov., Eremohaplomydas whartoni sp. nov. (both localities only 5 km apart), and Haplomydas crassipes (SimpleMappr https: // www. simplemappr. net / map / 14090). Distribution and occurrence data available in Google Earth KML file https: // www. simplemappr. net / map / 14090. kml and also through GBIF (data-set https: // www. gbif. org / dataset / 993875 DD- 5915 - 4107 - 8707 - 835 D 5 A 8 D 1022, DOI https: // doi. org / 10.15468 / awpjz 9)." figureDoi="10.3897/afrinvertebr.63.76309.figure57" httpUri="https://binary.pensoft.net/fig/663594" pageId="0" pageNumber="19">57</figureCitation>
). Adult flies are active in late summer to autumn (Table 
<tableCitation id="A646E1AF88939361866ECE1F5328BA09" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Seasonal imago flight activity of Eremohaplomydas, Haplomydas, and Lachnocorynus species through number of specimens collected and unique collecting events in each month (data given as  specimens /  collecting events). Months abbreviated starting with July. * = iNaturalist observation." httpUri="http://table.plazi.org/id/FD020CB799315F8304F2901FC17DB00E" pageId="0" pageNumber="19" tableUuid="FD020CB799315F8304F2901FC17DB00E">2</tableCitation>
). Nothing is known of the biology.
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