Revision of the Psilota Meigen, 1822 flower flies (Diptera: Syrphidae) of Australia
Author
Young, Andrew D.
Author
Skevington, Jeffrey H.
Author
Steenis, Wouter Van
text
Zootaxa
2020
2020-02-17
4737
1
1
127
journal article
24026
10.11646/zootaxa.4737.1.1
91c2f25f-409c-4319-bc7c-feb9e4210de1
1175-5326
3669490
C38FB007-98AE-4E4C-8A18-B1C75588815E
Psilota cuprea
Macquart, 1850
(Species plate:
Fig. 17
; Map:
Fig. 55B
)
Chrysogaster cupreus
Macquart, 1850: 317
.
Type
locality:
Tasmania
,
Australia
Diagnosis:
Face black, pale pollinose, with pale pile. Frons and vertex black pilose. Thorax black. Thoracic pile white, or mixed black and white. Hind leg simple. Calypter variable in colour. Halter variable in colour. Abdominal tergites 2 to 4 mostly to entirely red. Tergite pile variable in colour. Body length:
6.8–11.4 mm
.
FIGURE 17.
P
.
cuprea
Macquart. A
) male dorsal habitus, B) male anterolateral face, C) male lateral habitus. Scale bar =1 mm.
Description: MALE:
Head
:
Head black, face white pollinose. Face white, frons and vertex black pilose. Antenna orange to dark brown. Postpedicel variable in shape. Eye densely brown pilose.
Thorax
:
Thorax black. Legs entirely orange to entirely dark brown. Hind femur simple. Hind tibia simple. Pleuron lightly white pollinose or bare and shining. Pile pleuron entirely white or with mixed black and white pile. Pile on scutum and scutellum entirely white or with mixe black and white pile. Wing colourless. Wing microtrichia variable. Alula completely microtrichose. Halter variable in colour. Calypter variable in colour.
Abdomen
:
Tergite 1 black. Tergite 2 entirely red or with a large, anterior black area. Tergite 3 entirely red or with a small medial black spot. Tergite 4 entirely red. Pilosity of tergites variable in colour. Sternites 2 to 4 reddish, with variably coloured pilosity.
Genitalia
:
variable (see below).
FEMALE:
Identical except for normal sexual dimorphism.
Specimens examined:
AUSTRALIA
.
Australian Capital Territory
: Black Mountain,
-35.273675
149.097583
,
21.x.1966
;
23.x.1969
, I.F.B. Common, light trap, ANIC29034958; ANIC29034939; CNC_
Diptera
253814 (
3♀♀
, ANIC); Blundell’s [Cabin],
-35.292886
149.141800
,
20.i.1935
;
11.x.1950
;
24.x.1950
, S.J. Paramonov; Fuller, ANIC29034941; ANIC29034951; ANIC29034953-29034954; CNC_
Diptera
253704; CNC_
Diptera
253804 (
2♂♂
,
3♀♀
, ANIC); Blundells Ck.,
-35.220000
148.500000
,
ii.1987
, D.H. Colless, Malaise trap, CNC_Dip- tera253794 (
1♀
, ANIC); Canberra—CSIRO Botanical Garden,
-35.278878
149.108917
,
26.x.1988
, F.C. Thomp- son, USNM_ENT_00034137; USNM_ENT_00034139-00034141 (
4♀♀
, USNM); Canberra, Terr. 1,
-35.282000
149.128686
,
20-30.xi.1953
, M. Hull, CNC_
Diptera
254972 (
1♂
, CNC); Canberra,
-35.282000
149.128686
,
2.xi.1947
, S.J. Paramonov, ANIC29034940 (
1♂
, ANIC); Lees Creek Brindabella Rge.,
-35.338461
148.867925
,
1.i.1975
, G. Daniels, GDCBReg_19737 (
1♀
, GDCB); Mount Ainslie,
-35.266667
149.166667
,
28.xi.1996
, C.J. Burwell, QM_REG._NO._T223209 (
1♂
, QM);
New South Wales
: ‘Lugen Park’ Nullo Mountain Northeast of Rylstone,
-32.649278
150.217650
,
22.x.1953
, T.G. Campbell, ANIC29034942 (
1♂
, ANIC);
20 miles
South of Mil- ton,
-35.319178
150.439158
,
30.ix.1952
, S.J. Paramonov, ANIC29034943; ANIC29034959; CNC_
Diptera
253706; CNC_
Diptera
253815 (
4♀♀
, ANIC);
3 km
North of Lansdowne via Taree,
-31.758753
152.534258
, on house steps, wet sclerophyll forest, rainforest margin,
25.xii.1988
, G. Williams, AustralianMuseumK410625 (
1♀
, AM);
3 km
South of Mount Wilson, Blue Mountains,
-33.516011
150.378642
,
29.xi.1984
, D.K. McAlpine, B.J. Day, AustralianMuseumK407964 (
1♂
, AM;
4 miles
North of Bateman’s Bay,
-35.658456
150.177744
,
14.x.1952
;
12.x.1956
;
22.x.1952
,S.J. Paramonov,ANIC29034956;ANIC29034950; CNC_
Diptera
253705; CNC_
Diptera
253709 (
1♂
3♀♀
, ANIC); 4 mls W. of Amosfield,
-28.658686
152.016650
,
17.x.1971
, E.M. Exley, UQIC_Reg94770 (
1♀
, QM);
6 km
Northeast Bilpin, Blue Mountains,
-33.462750
150.572836
,
23.ix.1992
, N.W. Rodd, AustralianMuseumK410278 (
1♀
, AM); Back Yamma State Forest,
-33.339906
148.223703
,
11.xi.1964
, D.H. Colless, CNC_
Diptera
253708 (
1♀
, ANIC); Bald Rock N.P.,
-28.8525
152.055556
;
-28.852500
152.055558
, on
Leptospermum minutifolium
blossom,
6-7.xi.1984
, D.K. Yeates, UQIC_Reg94630; UQIC_Reg94631-94633; UQIC_Reg94637-94638; UQIC_Reg94744- UQIC94745; UQIC_Reg94675 (
4♂
m, #
5♀♀
, QM); Bawley Point,
-35.500000
150.400000
,
21-23.ix.2000
, Rentz, CNC_
Diptera
253696 (
1♂
, ANIC); Bilpin, Blue Mountains,
-33.500259
150.532989
,
21.ix.1978
, N.W. Rodd, Aus- tralianMuseumK410429 (
1♀
, AM); Blackheath,
-33.635539
150.283625
,
24.xii.1921
, CNC_
Diptera
253806 (
1♀
, ANIC); Boonoo Boonoo Falls,
-28.801247
152.165061
,
12.xi.1981
, G. & A. Daniels, GDCBReg_19687 (
1♀
, GDCB); Cherrybrook, electricity tower access track, end of Trevor’s Lane,
-33.710139
151.059500
,
13.ix.2008
, S. Ginn, AustralianMuseumK410146 (
1♂
, AM); Clyde Mountain, eastern foothills,
-35.546072
149.950333
,
1219m
, ex brackett fungus:
Stereum semilugens
,
15.viii.1967
, Z. Liepa, Emerged
25.VIII.1967
, ANIC29034945; ANIC29034952; CNC_
Diptera
253818 (
3♂♂
, ANIC); Cowan,
-33.590714
151.169561
,
7.xii.1931
, F.J. Gay, ANIC29034961 (
1♀
, ANIC); Dobroyd Point, Sydney,
-33.926375
151.125936
,
27.xi.1978
, N.W. Rodd, Australian- MuseumK410619 (
1♀
, AM); Durras Bay,
-35.645003
150.291439
,
12.xi.1953
, G. Daniels, CNC_
Diptera
253846 (
1♀
, CNC); Durras,
-35.646775
150.296939
,
29.ix.1959
, Z. Liepa, ANIC29034969 (
1♀
, ANIC); Gibraltar Range National Park, Waratah track, -29.5 152.333333, on
Leptospermum
sp. Blossom,
10-12.xi.1984
, D.K. Yeates, UQIC_Reg94634 (
1♀
, QM); Goondera Ridge, Royal National Park,
-34.114342
151.046586
,
2.x.1978
, G. Daniels, GDCBReg_19533; GDCBReg_19534 (
2♂♂
, GDCB);
22.viii.1976
, G. and A. Daniels, GDCBReg_19535 (
1♂
, GDCB); Heathcote NP, Kingfisher Pool,
-34.117042
150.952122
, dry eucalypt forest,
16.x.1994
, B.J.Sinclair, CNC_
Diptera
106594 (
1♀
, CNC); Jervis Bay,
-35.035431
150.668869
,
4.ix.1948
, E.F. Riek, ANIC29034949 (
1♂
, ANIC); Killara,
-33.768639
151.163469
,
13.x.1934
, D.F.W., ANIC29034955 (
1♀
, ANIC); Lorien Wildlife Refuge,
3 km
North of Lansdowne,
-31.758753
152.534258
,
7.xi.1993
, G. & T. Williams, AustralianMuseumK411046 (
1♂
, AM); Mooney Mooney Creek, near Gosford,
-33.437894
151.246467
,
1.xii.1989
, D.K. McAlpine, B.J. Day, AustralianMuseumK408009 (
1♀
, AM); Mount Canobolas State Recreation Area, summit Mount Canobolas,
12.8 km
Southwest of Orange,
-33.343889
148.982500
,
1395m
,
18.x.2002
, C. Lambkin, N. Starick, hand collected, ANIC29035762 (
1♂
, ANIC); Mount Gibralter, Comboyne Plateau,
-31.644328
152.422989
, shrub complex,
21.i.1985
, G. Williams, AustralianMuseumK410621 (
1♀
, AM); Mount Tomah, Blue Mountains,
-33.549614
150.416033
,
26.ix.1978
;
26.x.1978
;
28.xii.1985
, N.W. Rodd, N.W. Rodd Collection, AustralianMuseumK410430; AustralianMuseumK410252; AustralianMuseumK410154; AustralianMuseumK410163; AustralianMuseumK410175; AustralianMuseumK410182 (
2♂♂
,
3♀♀
, AM); Mt Canobolas State Recreation Area, summit Mt Canobolas
12.8 km
SW Orange,
-33.343889
148.982500
,
1395m
,
18.x.2002
, C.L. Lambkin, N.T. Starick, hand net, CNC_
Diptera
253847 (
1♀
, ANIC); Nadgee NR, near Eden,
-37.399378
149.908592
, wet sclerophyll forest,
21.xii.1985
;
14-15.ii.1986
, E.A. Sugden; D.J. Bickel, AustralianMuseumK410515 (
1♀
, AM); Nadgee Reserve
4.7km
S Merrica Hut,
-37.399378
149.908592
,
21.xii.1985
;
21.xii.1986
, E.A. Sugden, CNC_
Diptera
254983; Jeff_ Skevington_Specimen43807; Jeff_Skevington_Specimen43810-43811 (
1♂
,
3♀♀
, CNC; QM); New England Nat. Pk. Via Ebor,
-30.476022
152.660183
,
22–23.i.1966
, B. Cantrell, UQIC_Reg94705 UQIC_Reg94636; UQIC_ Reg94635 (
3♀♀
, QM); New England Nat. Pk. near Wright’s Lookout,
-30.476022
152.660183
,
14.x.1961
;
29– 31.x.2002
, C.W. Frazier; G.,M.Wood, CNC_
Diptera
253803; CNC_
Diptera
106427 (
2♀
, ANIC; CNC); Pigeon House Mtn., Ulladulla,
-35.349175
150.265064
,
13.xi.1976
, G. Daniels, GDCBReg_19767 (
1♀
, GDCB); Royal National Park, Flat Rock Creek, near Sydney,
-33.926375
151.125936
, From the rotten caudex of
Xanthorrhoea resinosa
,
10.viii.1973
, D.K. McAlpine, S.P. Kim, emergence on
4.IX.1973
, AustralianMuseumK410255 (
1♀
, AM); Rutherford Ck Brown Mtn. nr. Nimmitabel,
-36.601300
149.381311
,
26.xii.1974
, G. Daniels, GDCBReg_19536 (
1♂
, GDCB); Rutherford Creek, Brown Mtn. nr Nimmitabel,
-36.581667
149.551111
,
14.xi.1976
, G. Daniels, GDCBReg_19684 (
1♀
, GDCB); Siding Springs Observatory, Summit,
-31.255558
149.061111
,
31°15’20’’S
149°3’40’’E
,
17.x.1997
, J. Skevington, C. Lambkin, S. Winterton, UQIC_Reg94532–94536 (
5♂♂
, QM); Stanwell Tops,
-34.219903
150.991808
,
9.x.1977
, G. Daniels, GDCBReg_19528–19531 (
4♂
, GDCB); Tindery,
-35.754172
149.280764
,
1.i.1938
, E.H. Kipps, ANIC29034968 (
1♀
, ANIC); Wahroonga, Cook Trig, trail off Grosvenor Street,
-33.699450
151.126500
,
28.ix.2007
, D. Britton, S. Ginn, L. Holme, LHP002/01, AustralianMuseumK410478 (
1♂
, AM); Warrumbungle National Park Area,
-31.272222
149.061111
;
-31.279689
149.046797
, observatory hilltop; Observatory hilltop,
17.x.1997
, J. Skevington, UQIC_Reg94543–94545 (
3♂
, QM); Waste Point, Kosciusko Na- tional Park,
-36.427114
148.309022
,
24.i.1984
, David W. Inouye, AustralianMuseumK410600 (
1♀
, AM); West of Mount Mackenzie,
9 km
West of Tenterfield,
-29.066667
151.966667
, paddock with bracken fern,
9.xii.2003
, G. Daniels, AustralianMuseumK411060 (
1♀
, AM); Woolgoolga,
-30.113036
153.183792
,
28.i.1923
, Health Dept., CNC_
Diptera
253805 (
1♀
, ANIC); Woy woy,
-33.489839
151.319942
,
3.x.1925
, Mackerras, CNC_
Diptera
253809 (
1♀
, ANIC);
Queensland
:
7 km
North of Maryborough,
-25.470267
152.695200
,
23.viii.1977
, K.J. Lamkin, QM_ REG._NO._T223216 (
1♀
, QM); Agnes Water,
-33.710139
151.059500
,
3.ix.1987
, N.W. Rodd, AustralianMu- seumK410174 (
1♂
, AM); Bald Rock National Park, Boundary Swamp Creek,
-28.816667
152.066667
, Veg. over water,
11.x.1998
, J. Skevington, sweeping, UQIC_Reg94746 (
1♂
, QM); Bluff Range, West Slope, Biggenden,
- 25.600000
152.050000
,
21.viii.1975
, H. Frauca, ANIC29034946; CNC_
Diptera
253816 (
2♀
, ANIC); Brisbane,
- 27.526975
153.040322
, 1911, H. Hacker, QM_REG._NO._T223208 (
1♂
, QM);
x.1925
, G.H. Hardy, Australian- MuseumK407091 (
1♀
, AM); Brown Lake, N Stradbroke Is., S.E.Q.,
-27.490105
153.432379
, on
Leptospermum flavescens
blossom,
21–24.ix.1984
, D.K. Yeates, UQIC_Reg94674 (
1♂
, QM); Bundaberg,
-24.866972
152.350972
, viii-ix.1971, H. Frauca, ANIC29034970 (
1♀
, ANIC); Collaroy, Sydney,
-33.926375
151.125936
, 1921, Health Dept. Date is ambiguous reads ”2.9.21”, ANIC29034947 (
1♂
, ANIC); Dayboro,
0.5 km
South,
-27.200000
152.816667
,
18–19.ix.1999
, S.G. Evans, QM_REG._NO._T223227 (
1♀
, QM); Flinders Peak nr. Boonah,
- 27.800000
152.816667
, summit,
13.ix.1995
, D.K. Yeates, UQIC_Reg94538-94539 (
2♂♂
, QM); Flinders Peak, via Peak Crossing,
-27.816667
152.816667
,
13.ix.1995
, C.J. Burwell, D.K. Yeates, QM_REG._NO._T223211-223215 (
5♂♂
, QM); Girraween National Park, Hilltop
1.2km
NE of Bald Rock Campground,
-28.821942
151.937850
,
31.xii.2014
;
1.i.2015
, J.H. Skevington, CNC384663-384670; CNC384926-384927; CNC384624-38430; CNC451705–451706; (
20♂♂
, CNC); Kenniff’s Lookout, Mt. Moffatt sect., Carnarvon Nat. Pk.,
-25.165167
147.840419
,
25.ix.1986
, D.K. Yeates, UQIC_Reg94624 (
1♂
, QM); Lamington N.P.,
-28.045967
153.127983
,
29.x.1955
, F.A. Perkins, UQIC_Reg94905 (
1♀
, QM); Miriam Vale,
-24.330397
151.557592
, on
Angrophora costata
,
5.xii.1977
, T. Sears, UQIC_Reg94623 (
1♂
, QM); Mount Marlay Summit, Stanthorpe,
-28.666667
151.933333
,
20.x.1997
;
22.x.2005
, J. Skevington, C. Lambkin, S. Winterton; D. K. Yeates, UQIC_Reg94539–94541; ANIC29035763–29035764 (
5♂♂
, QM);
-28.653583
151.946906
;
-28.653583
151.946908
, on
Leptospermum
,
22.ix.1985
, D.K. Yeates, UQIC_Reg94626;, UQIC_Reg94639; UQIC_Reg94668; UQIC_Reg94735 (
4♂♂
, QM); Mt. Beerburrum, Glasshouse Mts.,
-26.950000
152.933333
,
280m
,
29.x.1991
, R. Eastwood, GDCBReg_19735 (
1♂
, QM); Mt. Norman Area, Via Wallangarra,
-28.873267
151.973822
,
2.x.1971
, G.B. Monteith, UQIC_Reg94769 (
1♀
, QM); Near Rosedale,
-24.631128
151.913664
,
27.x.1974
, H. Frauca, ANIC29034948; ANIC29034965 (
1♂
,
1♀
, ANIC); Six Mile Creek, W of Elimbah,
-27.016667
152.933333
,
15.ix.1985
, G. and A. Daniels, GDCBReg_ 19569 (
1♀
, QM); Stanthorpe,
-28.657928
151.930344
,
22.ix.1979
, S.J. Johnson, GDCBReg_19532 (
1♂
, GDCB); Summit of Mount Greville,
-28.074028
152.502008
,
393m
,
3.i.2015
, J.H., A.M. and A.W. Skevington, CNC451749 (
1♂
, CNC); Tambourine Mountain,
-27.923086
153.171717
,
28.x.1912
, H. Hacker, QM_REG._NO._T223184 (
1♀
, QM); The Blunder, Brisbane,
-27.526975
153.040322
,
9.x.1966
;
18.ix.1966
;
25–26.ix.1966
, C.F. Ashby, ANIC29034944; ANIC29034964; ANIC29034966; CNC_
Diptera
253707 (
1♂
,
3♀♀
, ANIC); Woombye, near Nam- bour,
-26.666911
152.966606
,
11–16.x.1965
, D.H. Colless, ANIC29034963 (
1♀
, ANIC); along Rd to 18 Mile Swamp, N. Stradbroke Island,
-27.516667
153.483333
,
17.ix.1989
, M.A. Schneider, UQIC_Reg94771 (
1♀
, QM);
South Australia
: Adelaide Area, Mount Lofty,
-34.984972
138.708333
,
4.x.1997
, J. Skevington, CNC_
Diptera
254973 (
1♂
, QM); Adelaide, Belair National Park,
-35.000000
138.633333
, Hilltopping,
4.x.1997
, J. & A. Skevington, C. Lambkin, S. Winterton, UQIC_Reg94522–94531 (
8♂♂
,
2♀♀
, QM); Dutchmans Stern Conserva- tion Park,
-32.296944
137.983058
, summit,
7.x.1997
, S. Winterton, J.&A. Skevington, C. Lambkin, UQIC_ Reg94537 (
1♂
, QM); Para Wirra Recreational Park,
-34.701739
138.824928
,
8.x.2014
, A. Marquart, SLAM trap, CNC587054 (
1♀
, CNC); Sandy Creek Conservation Park,
8km
NE Gawler,
-34.603333
138.855833
,
29.ix–4.x.1997
, S. Winterton, J.&A. Skevington, C. Lambkin, Malaise trap, malaise in heath, UQIC_Reg94542 (
1♂
, QM); South Para Reserve,
-34.676314
138.876983
,
20.x.2015
, A. Marquart, SLAM trap, CNC587060 (
1♀
, CNC);
Tasmania
:
1km
SEE Gladstone,
-40.580000
148.010000
,
29.i.1963
, I.D. Naumann & J.C. Cardale, CNC_
Diptera
253802 (
1♀
, ANIC);
9 km
SE Miena,
-42.000000
146.766667
,
1060m
,
11.i.1992
, G. and A. Daniels, GDCBReg_19616 (
1♀
, QM); Eaglehawk Neck,
-43.019722
147.925000
,
24.iv.1916
, G.H. Hardy, 12, CNC_
Diptera
253801 (
1♀
, ANIC); Erriba,
-41.425350
146.137136
,
30.i.1949
, E.F. Riek, ANIC29034960; ANIC29034967; CNC_
Diptera
253710 (
3♀♀
, ANIC); Franklin-Gordon Wild River National Park, Donaghy’s Hill,
-42.197778
145.931944
,
10–11.i.2015
, J.H., A.M. and A.W. Skevington, CNC385297; CNC385305; CNC385318; CNC451889–451892; CNC451894; CNC457453 (
5♂♂
,
4♀♀
, CNC); Hobart,
-42.882139
147.327194
,
16.xii.1917
, G.H. Hardy, 45, CNC_Dip- tera253800 (
1♀
, ANIC); Lyell Highway, Frankin R nature trail,
-42.215700
146.019800
,
11.i.2015
, S.J. Grove, hand collected, TMAG-F11616 (
1♂
, TMAG); N Bruny Island; Waterview Hill,
-43.093056
147.351111
, E Hilltop,
31.xii.1998
, J. & A. Skevington, M. Mathieson, UQIC_Reg94797 (
1♂
, QM); North of Corinna,
-41.588514
145.189764
,
12.i.2015
, J.H., A.M. and A.W. Skevington, CNC457433 (
1♀
, CNC); Oonah,
-41.224222
145.618608
,
30.i.1949
, E.F. Riek, ANIC29034971 (
1♀
, ANIC); Pump house Bay Camping Ground Arthurs Lake,
-41.983333
146.866667
,
12.ii.1992
, D.S. Horning, Jr., in flight, CNC_
Diptera
253698 (
1♀
, CNC); Samford, 1959, C.F. Ashby, Date is ambiguous reads ”6.12.59”, ANIC29034972 (
1♀
, ANIC); Warra Long Term Ecological Research Site 147, -43.1 146.683333,
17.xi–16.xii.1998
, malaise trap, UQIC_Reg94607–94608 (
2♀♀
, QM);
Victoria
: Ararat,
- 37.284372
142.932553
, H.W. Davey, CNC_
Diptera
253811 (
1♀
, ANIC); Grampians National Park, Reed’s Look- out,
-37.148111
142.448261
,
23.x.2014
, J.H., A.M. and A.W. Skevington, CNC384097; CNC457381 (
2♂♂
, CNC); Heathmont,
-37.829989
145.242019
, CNC_
Diptera
253778 (
1♀
, CNC); Mt. Buffalo N.P., Dicksons Falls,
-36.763417
146.788092
,
1440m
,
29.i.2006
, Jeff_Skevington_Specimen43805–43806 (
2♂♂
, QM); Yarra Ranges N.P., Mt. Don- na Buang track,
-37.709467
145.676642
,
4.ii.2006
, M.P. van Zuijen, Jeff_Skevington_Specimen43808 (
1♀
, QM);
Western Australia
:
2 km
North of Pemberton,
-34.426158
116.039058
,
29.ix.1987
, D.K. McAlpine, AustralianMuseumK407981 (
1♀
, AM); Geraldton,
-28.777372
114.614972
,
ix.1926
, E.W. Ferguson, CNC_
Diptera
253812 (
1♀
, ANIC); Barrington, Allyn Range,
-41.356572
146.279756
,
Leptospermum
,
ii.1925
, Label unclear: ”Tps Feb ‘25 SU ZooExp”, CNC_
Diptera
253810 (
1♀
, ANIC); Mt. Wellington,
-42.890000
147.230000
,
2.xii.2016
, G.H. Hardy, Date ambiguous: ”2.12.16 /48”, CNC_
Diptera
253798 (
1♂
, ANIC).
Discussion:
Psilota cuprea
doubtlessly encompasses multiple species as described here. Initial DNA barcode analysis indicates that at least nine species are represented (
Figure 43
). Further DNA barcoding and male genitalia dissection are required to adequately separate the full range of specimens into species and will be tackled in a future revision. Furthermore, there are several other species of
Psilota
described from
Australia
that may or may not be synonyms of
P. cuprea
,
including
P. erythrogaster
,
P. rubra
, and
P. rubriventris
(See ‘species not here recognized’ below). Because it is impossible to tell what name should properly be assigned to different barcode clusters without a full revision of this complex, these species have not been treated individually.As mentioned in the introduction, an individual from this complex was reared from the “rotten caudex of
Xanthorrhoea resinosa
”. The
holotype
of this species was not available for study.