Type series | Additional specimens |
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Characters | Holotype | Males (n = 15) | Females (n = 5) | Males (n = 3) | Females (n = 3) | Males (n = 38) |
Snout-vent length | 13.0 | 13.5 ± 0.4 (12.8–14.3) | 15.4 ± 0.6 (14.7–16.0) | 13.4 ± 0.3 (13.1–13.6) | 15.0 ± 0.3 (14.7–15.3) | 13.6 ± 0.7 (12.2–14.5) |
Head length | 4.1 | 3.9 ± 0.2 (3.6–4.3) | 4.4 ± 0.2 (4.2–4.6) | 3.8 ± 0.2 (3.7–4.0) | 4.4 ± 0.2 (4.2–4.5) | 3.5 ± 0.2 (3.2–4.0) |
Head width | 4.7 | 4.8 ± 0.1 (4.5–4.9) | 5.4 ± 0.2 (5.2–5.5) | 4.6 ± 0.2 (4.5–4.8) | 5.1 ± 0.2 (4.9–5.2) | 4.6 ± 0.2 (4.4–4.9) |
Eye diameter | 1.6 | 1.5 ± 0.1 (1.4–1.6) | 1.6 ± 0.1 (1.5–1.7) | 1.5 ± 0.0 (1.5–1.5) | 1.7 ± 0.1 (1.6–1.7) | 1.5 ± 0.1 (1.4–1.7) |
Interorbital distance | 1.5 | 1.4 ± 0.1 (1.2–1.7) | 1.7 ± 0.1 (1.6–1.8) | 1.5 ± 0.1 (1.4–1.5) | 1.6 ± 0.1 (1.5–1.6) | 1.4 ± 0.1 (1.2–1.5) |
Eye-nostril distance | 1.1 | 1.1 ± 0.1 (0.9–1.4) | 1.2 ± 0.2 (1.0–1.4) | 1.2 ± 0.1 (1.1–1.2) | 1.3 ± 0.1 (1.2–1.4) | 1.1 ± 0.1 (0.9–1.2) |
Snout length | 2.1 | 2.0 ± 0.1 (1.7–2.2) | 2.3 ± 0.2 (2.1–2.5) | 2.0 ± 0.1 (2.0–2.1) | 2.2 ± 0.1 (2.1–2.3) | 2.0 ± 0.1 (1.8–2.2) |
Internarial distance | 1.3 | 1.2 ± 0.1 (0.9–1.4) | 1.3 ± 0.1 (1.2–1.5) | 1.2 ± 0.1 (1.1–1.2) | 1.3 ± 0.1 (1.3–1.4) | 1.0 ± 0.1 (0.9–1.1) |
Hand length | 3.7 | 3.7 ± 0.2 (3.3–4.0) | 4.2 ± 0.3 (3.7–4.4) | 3.7 ± 0.3 (3.5–4.0) | 4.2 ± 0.2 (4.0–4.3) | 3.7 ± 0.2 (3.2–4.0) |
Thigh Length | 6.7 | 6.3 ± 0.2 (5.9–6.7) | 7.0 ± 0.5 (6.2–7.6) | 6.2 ± 0.1 (6.1–6.2) | 7.1 ± 0.3 (6.8–7.4) | 6.5 ± 0.3 (6.0–7.1) |
Tibia length | 7.2 | 6.9 ± 0.4 (6.2–7.5) | 7.8 ± 0.2 (7.5–8.0) | 6.6 ± 0.2 (6.4–6.8) | 7.7 ± 0.3 (7.4–8.0) | 7.2 ± 0.3 (6.6–7.7) |
Tarsus length | 3.9 | 3.7 ± 0.3 (3.4–4.1) | 4.4 ± 0.2 (4.1–4.6) | 3.8 ± 0.1 (3.7–3.9) | 4.4 ± 0.4 (4.0–4.8) | 3.7 ± 0.2 (3.3–4.2) |
Foot length | 7.8 | 7.2 ± 0.3 (6.4–7.7) | 8.1 ± 0.4 (7.6–8.5) | 7.3 ± 0.1 (7.2–7.4) | 8.3 ± 0.4 (7.9–8.6) | 7.3 ± 0.3 (6.5–7.8) |
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Traits | Type locality n = 23 (460) | Other localities n = 8 (160) | Type locality n = 9 (180) |
Call duration (s) | 26.1 ± 18.7 (3.6–66.2) | 44.4 ± 34.9 (19.7–69.1) | – |
Series of notes duration (s) | 9.4 ± 6.5 (0.2–34.0) | 23.7 ± 14.5 (7.4–46.0) | 14.5 ± 15.7 (0.4–44.3) |
Interseries interval (s) | 0.6 ± 0.5 (0.2–2.0) | 0.6 ± 0.3 (0.3–0.8) | 1.9 ± 0.0 (0.3–10.9) |
Series per call | 3.4 ± 2.0 (1–9) | 2.5 ± 0.7 (1–3) | - |
Note duration (ms) | 37 ± 3 (27–46) | 41 ± 4 (34–52) | 69 ± 8 (50–114) |
Internote interval (ms) | 29 ± 6 (17–72) | 49 ± 9 (35–76) | 74 ± 20 (42–159) |
Pulse duration (ms) | 9 ± 2 (4–15) | 6 ± 1 (3–10) | 10 ± 3 (4–22) |
Interpulse interval (ms) | 18 ± 3 (11–31) | 30 ± 4 (22–40) | 47 ± 10 (5–94) |
Notes per minute | 928.6 ± 85.1 (738.0–1101.4) | 655.2 ± 75.4 (541.9–736.6) | 427.1 ± 64.2 (282.1–485.1) |
Pulse rate | 54.7 ± 5.9 (43.5–74.1) | 48.8 ± 4.4 (38.5–58.8) | 29.8 ± 3.0 (17.5–117.6) |
Pulses per note | 2.0 ± 0.0 (2–2) | 2.0 ± 0.0 (2–2) | 2.0 ± 0.1 (2–3) |
Dominant frequency (kHz) | 4.5 ± 0.2 (3.9–5.1) | 4.7 ± 0.1 (4.6–5.0) | 3.8 ± 0.2 (3.6–4.3) |
Minimum frequency (kHz) | 4.0 ± 0.3 (2.9–4.6) | 4.1 ± 0.3 (1.8–4.4) | 3.2 ± 0.2 (2.3–3.9) |
Maximum frequency (kHz) | 4.9 ± 0.2 (4.4–5.5) | 5.1 ± 0.1 (4.9–5.2) | 4.2 ± 0.1 (4.0–4.5) |
Air temperature (°C) | 24.0–25.2 | 20.5–23.0 | 17.0–24.0 |
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|
class.error | |
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15 |
|
4 |
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0.21 |
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|
1 |
|
37 |
|
0.03 | 0.00 |
Inter (SD) | |||||||||||||||||||
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Intra (SD) | Pmat | Pmin | Psal | Pmur | Pjar | Ppoc | Pres | Pflo | Pfal | Pmot | Pbol | Pame | Pter | Pfac | Patr | Pcan | Pbar | Pmys | |
Pmat | 0.001 (0.001) | 0.007 | 0.009 | 0.009 | 0.01 | 0.009 | 0.009 | 0.01 | 0.011 | 0.015 | 0.014 | 0.014 | 0.014 | 0.014 | 0.014 | 0.015 | 0.015 | 0.014 | |
Pmin | 0.003 (0.002) |
|
0.01 | 0.009 | 0.01 | 0.007 | 0.008 | 0.008 | 0.01 | 0.014 | 0.013 | 0.014 | 0.013 | 0.014 | 0.014 | 0.013 | 0.014 | 0.014 | |
Psal | 0.003 (0.001) |
|
0.056 | 0.006 | 0.007 | 0.009 | 0.01 | 0.011 | 0.012 | 0.014 | 0.014 | 0.014 | 0.012 | 0.013 | 0.012 | 0.012 | 0.013 | 0.013 | |
Pmur | 0.002 (0.001) |
|
0.048 | 0.025 | 0.006 | 0.009 | 0.01 | 0.01 | 0.011 | 0.014 | 0.013 | 0.013 | 0.012 | 0.012 | 0.012 | 0.012 | 0.012 | 0.013 | |
Pjar | 0 (0) |
|
0.056 | 0.026 | 0.022 | 0.01 | 0.01 | 0.011 | 0.012 | 0.014 | 0.014 | 0.014 | 0.013 | 0.015 | 0.014 | 0.014 | 0.014 | 0.015 | |
Ppoc | 0.01 (0.003) |
|
0.045 | 0.06 | 0.054 | 0.064 | 0.006 | 0.008 | 0.01 | 0.014 | 0.014 | 0.014 | 0.013 | 0.014 | 0.014 | 0.013 | 0.013 | 0.014 | |
Pres | 0.002 (0.002) |
|
0.046 | 0.062 | 0.056 | 0.068 | 0.035 | 0.007 | 0.011 | 0.013 | 0.013 | 0.013 | 0.013 | 0.014 | 0.013 | 0.013 | 0.013 | 0.014 | |
Pflo | 0.001 (0.001) |
|
0.046 | 0.066 | 0.06 | 0.072 | 0.039 | 0.031 | 0.011 | 0.013 | 0.013 | 0.014 | 0.014 | 0.014 | 0.013 | 0.013 | 0.014 | 0.014 | |
Pfal | 0.004 (0.002) |
|
0.072 | 0.088 | 0.07 | 0.087 | 0.075 | 0.076 | 0.08 | 0.015 | 0.014 | 0.014 | 0.014 | 0.015 | 0.014 | 0.014 | 0.015 | 0.016 | |
Pmot | 0.002 (0.001) |
|
0.134 | 0.142 | 0.127 | 0.134 | 0.139 | 0.131 | 0.131 | 0.134 | 0.01 | 0.014 | 0.014 | 0.013 | 0.015 | 0.015 | 0.014 | 0.013 | |
Pbol | 0.005 (0.003) |
|
0.118 | 0.131 | 0.12 | 0.137 | 0.125 | 0.125 | 0.12 | 0.12 | 0.07 | 0.013 | 0.013 | 0.014 | 0.014 | 0.014 | 0.013 | 0.014 | |
Pame | 0.003 (0.002) |
|
0.119 | 0.123 | 0.112 | 0.121 | 0.13 | 0.127 | 0.125 | 0.118 | 0.11 | 0.127 | 0.005 | 0.013 | 0.012 | 0.011 | 0.011 | 0.012 | |
Pter | 0.004 (0.001) |
|
0.123 | 0.124 | 0.114 | 0.125 | 0.133 | 0.125 | 0.127 | 0.12 | 0.112 | 0.128 | 0.015 | 0.013 | 0.013 | 0.011 | 0.012 | 0.012 | |
Pfac | 0 (0) |
|
0.17 | 0.152 | 0.147 | 0.163 | 0.167 | 0.162 | 0.168 | 0.154 | 0.153 | 0.163 | 0.104 | 0.108 | 0.007 | 0.011 | 0.011 | 0.011 | |
Patr | 0 (0) |
|
0.167 | 0.147 | 0.144 | 0.161 | 0.166 | 0.162 | 0.168 | 0.158 | 0.163 | 0.171 | 0.102 | 0.106 | 0.032 | 0.01 | 0.009 | 0.01 | |
Pcan | 0 (0) |
|
0.151 | 0.14 | 0.128 | 0.151 | 0.152 | 0.145 | 0.151 | 0.129 | 0.147 | 0.152 | 0.089 | 0.085 | 0.083 | 0.079 | 0.006 | 0.011 | |
Pbar | 0.001 (0.001) |
|
0.15 | 0.136 | 0.125 | 0.148 | 0.146 | 0.148 | 0.152 | 0.13 | 0.145 | 0.15 | 0.082 | 0.081 | 0.076 | 0.072 | 0.018 | 0.011 | |
Pmys | 0.015 (0.003) |
|
0.146 | 0.143 | 0.133 | 0.153 | 0.162 | 0.157 | 0.15 | 0.147 | 0.148 | 0.157 | 0.096 | 0.098 | 0.095 | 0.09 | 0.08 | 0.078 |