<document id="05B36F0C1EE583997CEEA268A52E93D2" ID-DOI="10.11646/zootaxa.1182.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5064071" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625278111192" checkinUser="felipe" docAuthor="Mermudes, José Ricardo M. &amp; Napp, Dilma Solange" docDate="2006" docId="A02F87F5FF85F9781730FAEAFEFDFA58" docLanguage="en" docName="zt01182p130.pdf" docOrigin="Zootaxa 1182 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1182.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Ptychoderes brevis Jordan 1894" docType="treatment" docVersion="3" lastPageNumber="28" masterDocId="5C16FF8DFF9CF9631638FFB7FF8DFFC1" masterDocTitle="Revision and cladistic analysis of the genus: Ptychoderes Schoenherr, 1823 (Coleoptera, Anthribidae, Anthribinae, Ptychoderini)" masterLastPageNumber="130" masterPageNumber="1" pageNumber="26" updateTime="1698984982767" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="C6CCB729CEB494341B88141D84E2CC5D">Revision and cladistic analysis of the genus: Ptychoderes Schoenherr, 1823 (Coleoptera, Anthribidae, Anthribinae, Ptychoderini)</mods:title>
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<mods:namePart id="2E3B7D768F9C6398600938D968B44FEF">Mermudes, José Ricardo M.</mods:namePart>
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<heading id="7371818FFF85F97A1730FAEAFD19FAB7" bold="true" box="[264,660,1372,1399]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
<taxonomicName id="EF864D60FF85F97A1730FAEAFD19FAB7" ID-CoL="78NJ3" authority="Jordan, 1894" authorityName="Jordan" authorityYear="1894" box="[264,660,1372,1399]" class="Insecta" family="Anthribidae" genus="Ptychoderes" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="1AF2EAF1FF85F97A1730FAEAFD19FAB7" bold="true" box="[264,660,1372,1399]" pageId="25" pageNumber="26">
<emphasis id="1AF2EAF1FF85F97A1730FAEAFE65FAB6" bold="true" box="[264,488,1373,1399]" italics="true" pageId="25" pageNumber="26">Ptychoderes brevis</emphasis>
<bibRefCitation id="4C174B12FF85F97A17C9FAEBFD19FAB7" author="Jordan, K." box="[497,660,1372,1398]" pageId="25" pageNumber="26" pagination="591 - 651" refId="ref63636" refString="Jordan, K. (1894) On Anthribidae in the museum of Honourable Walter Rothschild. Novitates Zoologicae 1 (4), 591 - 651." type="journal article" year="1894">
<collectingCountry id="50917673FF85F97A17C9FAEBFDC1FAB7" box="[497,588,1372,1398]" name="Jordan" pageId="25" pageNumber="26">Jordan</collectingCountry>
, 1894
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</emphasis>
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</heading>
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<subSubSection id="609C6568FF85F97A1730FAC8FDA9FA58" box="[264,548,1407,1433]" pageId="25" pageNumber="26" type="description">
<paragraph id="283936E3FF85F97A1730FAC8FDA9FA58" blockId="25.[264,660,1372,1433]" box="[264,548,1407,1433]" pageId="25" pageNumber="26">
<figureCitation id="B0BD2A66FF85F97A1730FAC8FEF5FA58" box="[264,376,1407,1433]" captionStart="FIGURES 208–214" captionStartId="117.[264,382,1545,1569]" captionTargetBox="[293,1285,343,1521]" captionTargetId="figure-64@117.[292,1294,326,1521]" captionTargetPageId="117" captionText="FIGURES 208–214. 208, Ptychoderes crustatus Frieser, 1988, male; 209, Ptychoderes brevis Jordan, 1894, female holotype; 210, Ptychoderes jekeli Mermudes &amp; Napp, 2004, female holotype; 211, Ptychoderes longicollis Jordan, 1894, male. 212–214, Ptychoderes jordani Frieser, 1959. 212, male holotype; 213, female paratype; 214, pale form, male." figureDoi="http://doi.org/10.5281/zenodo.5064125" httpUri="https://zenodo.org/record/5064125/files/figure.png" pageId="25" pageNumber="26">Figs. 209</figureCitation>
, 
<figureCitation id="B0BD2A66FF85F97A17BBFAC8FE3CFA58" box="[387,433,1407,1433]" captionStart="FIGURES 228–238" captionStartId="120.[264,383,1517,1541]" captionTargetBox="[280,1307,348,1493]" captionTargetId="figure-83@120.[280,1307,326,1493]" captionTargetPageId="120" captionText="FIGURES 228–238. 228, Ptychoderes bivittatus Jordan, 1894, male; 229, P. nebulosus (Olivier, 1795), male; 230, P. brevis Jordan, 1894, female holotype. 231–234 detail of elytral apex. 231, Unanthribus maximus; 232, Ptychoderes mixtus; 233, Ptychoderes elongatus; 234, Ptychoderes callosus. 235, pygidium and ventrite V, Ptychoderes nebulosus. 236–238, ventrite I, male. 236, Ptychoderes mixtus; 237, Ptychoderes rugicollis; 238, Ptychoderes nebulosus." figureDoi="http://doi.org/10.5281/zenodo.5064131" httpUri="https://zenodo.org/record/5064131/files/figure.png" pageId="25" pageNumber="26">230</figureCitation>
, 
<figureCitation id="B0BD2A66FF85F97A1786FAC8FE60FA58" box="[446,493,1407,1433]" captionStart="FIGURES 239–250" captionStartId="121.[264,383,1756,1780]" captionTargetBox="[309,1279,326,1725]" captionTargetId="figure-77@121.[309,1279,326,1725]" captionTargetPageId="121" captionText="FIGURES 239–250. Pronotum. 239, Tribotropis prasinata; 240, Unanthribus grandis; 241, Ptychoderes crustatus; 242, P. brevis, female holotype; 243, P. longicollis, holotype; 244, P. jordani, male holotype; 245, P. magnus, male holotype; 246, P. depressus; 247, P. virgatus; 248, P. bivittatus, male syntype; 249, P.rugicollis; 250, P. elongatus." figureDoi="http://doi.org/10.5281/zenodo.5064133" httpUri="https://zenodo.org/record/5064133/files/figure.png" pageId="25" pageNumber="26">242</figureCitation>
, 
<figureCitation id="B0BD2A66FF85F97A17C0FAC8FDA9FA58" box="[504,548,1407,1433]" captionStart="FIGURES 263–264. 263" captionStartId="123.[264,383,1767,1791]" captionTargetBox="[383,1204,328,1733]" captionTargetId="figure-62@123.[383,1205,318,1763]" captionTargetPageId="123" captionText="FIGURES 263–264. 263. Geographical distribution of P. longicollis (black circles), P. crustatus (white triangles), P. brevis (black squares), P. jekeli (black triangles). 264. Geographical distribution of P. jordani (black squares), P.depressus (black triangles), P. virgatus (white circles)." figureDoi="http://doi.org/10.5281/zenodo.5064137" httpUri="https://zenodo.org/record/5064137/files/figure.png" pageId="25" pageNumber="26">263</figureCitation>
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<taxonomicName id="EF864D60FF85F97A1730FA67FD27FA27" ID-CoL="78NJ3" authority="Jordan, 1894: 592" authorityName="Jordan" authorityPageNumber="592" authorityYear="1894" box="[264,682,1487,1511]" class="Insecta" family="Anthribidae" genus="Ptychoderes" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="1AF2EAF1FF85F97A1730FA67FE58FA26" box="[264,469,1488,1511]" italics="true" pageId="25" pageNumber="26">Ptychoderes brevis</emphasis>
<treatmentCitation id="A92710F2FF85F97A17D8FA78FD27FA27" author="Jordan, K." box="[480,682,1487,1510]" page="592" pageId="25" pageNumber="26" year="1894">
<bibRefCitation id="4C174B12FF85F97A17D8FA78FD27FA27" author="Jordan, K." box="[480,682,1487,1510]" pageId="25" pageNumber="26" pagination="591 - 651" refId="ref63636" refString="Jordan, K. (1894) On Anthribidae in the museum of Honourable Walter Rothschild. Novitates Zoologicae 1 (4), 591 - 651." type="journal article" year="1894">
<collectingCountry id="50917673FF85F97A17D8FA78FDA7FA27" box="[480,554,1487,1510]" name="Jordan" pageId="25" pageNumber="26">Jordan</collectingCountry>
, 1894: 592
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; 
<treatmentCitation id="A92710F2FF85F97A1480FA78FCF5FA27" author="Bovie, A." box="[696,888,1487,1510]" page="219" pageId="25" pageNumber="26" year="1906">
<bibRefCitation id="4C174B12FF85F97A1480FA78FCF5FA27" author="Bovie, A." box="[696,888,1487,1510]" pageId="25" pageNumber="26" pagination="218 - 334" refId="ref62859" refString="Bovie, A. (1906) Catalogue des Anthribides. Annales de la Societe Entomologique de Belgique, 49, 218 - 334." type="journal article" year="1906">Bovie, 1906: 219</bibRefCitation>
</treatmentCitation>
(cat.); 
<treatmentCitation id="A92710F2FF85F97A15F4FA78FB1DFA27" author="Wolfrum, P." box="[972,1168,1487,1510]" page="6" pageId="25" pageNumber="26" year="1929">
<bibRefCitation id="4C174B12FF85F97A15F4FA78FB1DFA27" author="Wolfrum, P." box="[972,1168,1487,1510]" pageId="25" pageNumber="26" refId="ref65348" refString="Wolfrum, P. (1929) Coleopterorum Catalogus, pars 102, Anthribidae. W. Junk, Berlin, 145 p." type="book" year="1929">Wolfrum, 1929: 6</bibRefCitation>
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(cat.); 
<treatmentCitation id="A92710F2FF85F97A12DCFA78FE75F9C7" author="Blackwelder, R. E." page="765" pageId="25" pageNumber="26" year="1947">
<bibRefCitation id="4C174B12FF85F97A12DCFA78FE75F9C7" author="Blackwelder, R. E." pageId="25" pageNumber="26" pagination="765 - 925" refId="ref62793" refString="Blackwelder, R. E. (1947) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Bulletin of United State National Museum 185 (5), 765 - 925." type="journal article" year="1947">Blackwelder, 1947: 765</bibRefCitation>
</treatmentCitation>
(cat.); 
<treatmentCitation id="A92710F2FF85F97A147DFA58FCB4F9C7" author="Rheinheimer, J." box="[581,825,1519,1542]" page="10" pageId="25" pageNumber="26" year="2004">
<bibRefCitation id="4C174B12FF85F97A147DFA58FCB4F9C7" author="Rheinheimer, J." box="[581,825,1519,1542]" pageId="25" pageNumber="26" pagination="3 - 242" refId="ref64865" refString="Rheinheimer, J. (2004) Illustrater Katalog und Bibliographie der Anthribidae der Welt (Insecta: Coleoptera). Mitteilungen des Entomologischen Vereins Stuttgart, 39 (1 / 2), 3 - 242." type="journal article" year="2004">Rheinheimer, 2004: 10</bibRefCitation>
</treatmentCitation>
(cat.).
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</paragraph>
<paragraph id="283936E3FF85F9791730F980FD27FEF7" blockId="25.[264,1325,1591,1897]" lastBlockId="26.[264,1325,284,1870]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
Female. Vestiture (
<figureCitation id="B0BD2A66FF85F97A17DDF980FDC1F990" box="[485,588,1591,1617]" captionStart="FIGURES 208–214" captionStartId="117.[264,382,1545,1569]" captionTargetBox="[293,1285,343,1521]" captionTargetId="figure-64@117.[292,1294,326,1521]" captionTargetPageId="117" captionText="FIGURES 208–214. 208, Ptychoderes crustatus Frieser, 1988, male; 209, Ptychoderes brevis Jordan, 1894, female holotype; 210, Ptychoderes jekeli Mermudes &amp; Napp, 2004, female holotype; 211, Ptychoderes longicollis Jordan, 1894, male. 212–214, Ptychoderes jordani Frieser, 1959. 212, male holotype; 213, female paratype; 214, pale form, male." figureDoi="http://doi.org/10.5281/zenodo.5064125" httpUri="https://zenodo.org/record/5064125/files/figure.png" pageId="25" pageNumber="26">Fig. 209</figureCitation>
): pale dorsal vitta with dense whitish scales, intermingled with olivaceous­green scales, decumbent and denser at sides of pronotum, elytral interstriae 1, 2 and 4 and apical declivity. Pygidium with predominance of olivaceous­green scales. Ventral body surface and legs with appressed, dense, yellowish­white scales; sides of prosternum, below lateral carina, with a vitta of scales concolorous with those of pronotum (
<figureCitation id="B0BD2A66FF85F97A1729F948FEF7F8D8" box="[273,378,1791,1817]" captionStart="FIGURES 228–238" captionStartId="120.[264,383,1517,1541]" captionTargetBox="[280,1307,348,1493]" captionTargetId="figure-83@120.[280,1307,326,1493]" captionTargetPageId="120" captionText="FIGURES 228–238. 228, Ptychoderes bivittatus Jordan, 1894, male; 229, P. nebulosus (Olivier, 1795), male; 230, P. brevis Jordan, 1894, female holotype. 231–234 detail of elytral apex. 231, Unanthribus maximus; 232, Ptychoderes mixtus; 233, Ptychoderes elongatus; 234, Ptychoderes callosus. 235, pygidium and ventrite V, Ptychoderes nebulosus. 236–238, ventrite I, male. 236, Ptychoderes mixtus; 237, Ptychoderes rugicollis; 238, Ptychoderes nebulosus." figureDoi="http://doi.org/10.5281/zenodo.5064131" httpUri="https://zenodo.org/record/5064131/files/figure.png" pageId="25" pageNumber="26">Fig. 230</figureCitation>
). Sides of mesosternum with a wide, glabrous vitta. Mesepimera, mesepisterna, metepisterna, sides of both metasternum and ventrites with shallow setigerous punctures, each provided with a pale, yellowish scale. Outer surface of meso­ and metafemora with two spots of blackish­brown scales.
</paragraph>
</subSubSection>
<subSubSection id="609C6568FF86F9781700FEF3FEE3FDE1" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="description">
<paragraph id="283936E3FF86F9791700FEF3FDADFDB7" blockId="26.[264,1325,284,1870]" pageId="26" pageNumber="27">Rostrum at least 1.2 times longer than wide across base. Distance between scrobe and eyes about half maximum width of eye. Sides looking tricarinate due to the careniform upper margin of scrobe; carinae long and thick fused with the scrobal margin; with a transverse carina between the margin of scrobe and lower carina. Scrobes careniform at dorsal and basal margins; extension deep, narrow with a deep, distal pit, slightly wider than extension. Mentum finely, sparsely punctate; strongly impressed at sides, shallowly grooved at apical margin. Gula with moderately coarse, shallow, sparse punctures. Frons with prominent carinae.</paragraph>
<paragraph id="283936E3FF86F9791700FD33FC00FCFF" blockId="26.[264,1325,284,1870]" pageId="26" pageNumber="27">Antennae short, reaching antebasal carina of prothorax. Segments III–IV slightly longer than II; V slightly longer than VI–VIII, subequal; IX one third longer than VIII; X the shortest; IX–X weakly expanded at apex; XI one third longer than X (II = 0.56–0.65; III = 0.70–0.72; IV = 0.72–0.75; V = 0.60–0.65; VI = 0.52–0.55; VII = 0.52–0.55; VIII = 0.48–0.50; IX = 0.75–0.76; X = 0.48–0.55; XI = 0.80).</paragraph>
<paragraph id="283936E3FF86F9791700FCFBFB14FA7F" blockId="26.[264,1325,284,1870]" pageId="26" pageNumber="27">
Prothorax (
<figureCitation id="B0BD2A66FF86F9791784FCFBFDA0FCA7" box="[444,557,844,870]" captionStart="FIGURES 208–214" captionStartId="117.[264,382,1545,1569]" captionTargetBox="[293,1285,343,1521]" captionTargetId="figure-64@117.[292,1294,326,1521]" captionTargetPageId="117" captionText="FIGURES 208–214. 208, Ptychoderes crustatus Frieser, 1988, male; 209, Ptychoderes brevis Jordan, 1894, female holotype; 210, Ptychoderes jekeli Mermudes &amp; Napp, 2004, female holotype; 211, Ptychoderes longicollis Jordan, 1894, male. 212–214, Ptychoderes jordani Frieser, 1959. 212, male holotype; 213, female paratype; 214, pale form, male." figureDoi="http://doi.org/10.5281/zenodo.5064125" httpUri="https://zenodo.org/record/5064125/files/figure.png" pageId="26" pageNumber="27">Figs. 209</figureCitation>
, 
<figureCitation id="B0BD2A66FF86F9791401FCFBFDEBFCA7" box="[569,614,844,870]" captionStart="FIGURES 239–250" captionStartId="121.[264,383,1756,1780]" captionTargetBox="[309,1279,326,1725]" captionTargetId="figure-77@121.[309,1279,326,1725]" captionTargetPageId="121" captionText="FIGURES 239–250. Pronotum. 239, Tribotropis prasinata; 240, Unanthribus grandis; 241, Ptychoderes crustatus; 242, P. brevis, female holotype; 243, P. longicollis, holotype; 244, P. jordani, male holotype; 245, P. magnus, male holotype; 246, P. depressus; 247, P. virgatus; 248, P. bivittatus, male syntype; 249, P.rugicollis; 250, P. elongatus." figureDoi="http://doi.org/10.5281/zenodo.5064133" httpUri="https://zenodo.org/record/5064133/files/figure.png" pageId="26" pageNumber="27">242</figureCitation>
) slightly longer than wide; sides converging anteriorly from antebasal carina. Pronotal depression deep, with a almost rounded, weakly prominent, median tubercle, not reaching lateral margins of depression. Pronotum evidently rugose throughout, the rugae prominent and long. Antebasal carina interrupted at middle (almost twice width of carina), oblique towards lateral carina. Lateral carina straight (
<figureCitation id="B0BD2A66FF86F979128CFC5BFA93FBC7" box="[1204,1310,1004,1030]" captionStart="FIGURES 228–238" captionStartId="120.[264,383,1517,1541]" captionTargetBox="[280,1307,348,1493]" captionTargetId="figure-83@120.[280,1307,326,1493]" captionTargetPageId="120" captionText="FIGURES 228–238. 228, Ptychoderes bivittatus Jordan, 1894, male; 229, P. nebulosus (Olivier, 1795), male; 230, P. brevis Jordan, 1894, female holotype. 231–234 detail of elytral apex. 231, Unanthribus maximus; 232, Ptychoderes mixtus; 233, Ptychoderes elongatus; 234, Ptychoderes callosus. 235, pygidium and ventrite V, Ptychoderes nebulosus. 236–238, ventrite I, male. 236, Ptychoderes mixtus; 237, Ptychoderes rugicollis; 238, Ptychoderes nebulosus." figureDoi="http://doi.org/10.5281/zenodo.5064131" httpUri="https://zenodo.org/record/5064131/files/figure.png" pageId="26" pageNumber="27">Fig. 230</figureCitation>
). Secondary carina sinuous, narrowly interrupted at middle, almost reaching lateral carina. Laterobasal carina reaching posterior margin of prothorax. Prosternum raised, with coarse, deep dense punctures; antecoxal furrow straight, deep and entire. Mesosternum with coarse, shallow, sparse punctures; near the intercoxal process with two median, almost rounded, contiguous, deep depressions and coarse punctures at sides. Intercoxal process of mesosternum evidently impressed at apex, swollen at lateral and apical margins, with lateroapical projections. Elytra 1.2–1.9 times longer than wide across humeri, somewhat convex, transversely depressed before middle; basal gibbosities prominent; surface rugose from base to behind middle, more so at base and near suture. Sides subparallel at basal half, slightly wider than humeri behind middle. Anteapical tubercles absent, interstriae not swollen. Outer apical glabrous area of profemora with a single deep puncture.
</paragraph>
<paragraph id="283936E3FF86F9791700FA7BFE1AF96F" blockId="26.[264,1325,284,1870]" pageId="26" pageNumber="27">
Abdomen. Ventrite I with scarce, fine punctures, partially obscured by vestiture; intercoxal process depressed, groove line shaped. Ventrite V one third longer than IV, with lateroapical projections somewhat evident, not acuminated; apex emarginate. Pygidium slightly longer than wide, subconvex, not impressed medially at apex; with a short groove to fit elytra; apex truncate. Ovipositor (apex exposed) with a toothed plate similar to that of 
<taxonomicName id="EF864D60FF86F9791730F923FE1CF96C" authorityName="Mermudes &amp; Napp" authorityYear="2006" box="[264,401,1684,1709]" class="Insecta" family="Anthribidae" genus="Ptychoderes" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="crustatus">
<emphasis id="1AF2EAF1FF86F9791730F923FE1CF96C" box="[264,401,1684,1709]" italics="true" pageId="26" pageNumber="27">P. crustatus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="283936E3FF86F9791700F90BFDC0F8E7" blockId="26.[264,1325,284,1870]" pageId="26" pageNumber="27">Measurements in mm, female. TL = 14.3–14.5; RL = 2.0–2.1; RAW = 1.7–1.9; RBW = 1.5–1.7; HW = 2.3–2.5; DSO = 0.7–0.7; MLO = 0.7–0.7; PL = 4.2–4.3; PW = 4.0–4.2; E L = 6.0–9.0; EW = 4.7–4.8.</paragraph>
<paragraph id="283936E3FF86F9791700F883FE73F88F" blockId="26.[264,1325,284,1870]" box="[312,510,1844,1870]" pageId="26" pageNumber="27">Male. Unknown.</paragraph>
<paragraph id="283936E3FF87F9781730FEABFE24FEF4" blockId="27.[264,1323,284,465]" box="[264,425,284,309]" pageId="27" pageNumber="28">
<emphasis id="1AF2EAF1FF87F9781730FEABFE24FEF4" box="[264,425,284,309]" italics="true" pageId="27" pageNumber="28">Type material</emphasis>
</paragraph>
<paragraph id="283936E3FF87F9781700FE88FDC3FE10" blockId="27.[264,1323,284,465]" pageId="27" pageNumber="28">
<materialsCitation id="98EE3CBEFF87F9781700FE88FDC7FE10" collectionCode="BMNH" collectorName="Para, N." country="Jordan" county="Karl" location="Type" municipality="Type" pageId="27" pageNumber="28" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
Female 
<typeStatus id="F73D8841FF87F97817A2FE88FD8DFE98" box="[410,512,319,345]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
(
<collectionCode id="4E97AE26FF87F978142CFE88FDE2FE98" box="[532,623,319,345]" pageId="27" pageNumber="28">BMNH</collectionCode>
) examined, labelled as follows: 1) 
<location id="2D596038FF87F978121DFE88FBD2FE98" LSID="urn:lsid:plazi:treatment:A02F87F5FF85F9781730FAEAFEFDFA58:2D596038FF87F978121DFE88FBD2FE98" box="[1061,1119,319,345]" country="Jordan" county="Karl" municipality="Type" name="Type" pageId="27" pageNumber="28">Type</location>
[rounded, white with red border, printed]; 2) 
<taxonomicName id="EF864D60FF87F9781459FED0FCB2FE41" authorityName="Jordan" authorityYear="1894" box="[609,831,359,384]" class="Insecta" family="Anthribidae" genus="Ptychoderes" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="1AF2EAF1FF87F9781459FED0FCB2FE41" box="[609,831,359,384]" italics="true" pageId="27" pageNumber="28">Ptychoderes brevis</emphasis>
</taxonomicName>
, 
<specimenCount id="3E80FD6AFF87F978156BFED1FCEAFEBB" box="[851,871,358,378]" pageId="27" pageNumber="28" type="female">♀</specimenCount>
, 
<collectingMunicipality id="C85DAC99FF87F978154EFED0FC38FE40" box="[886,949,359,385]" pageId="27" pageNumber="28">Type</collectingMunicipality>
: 
<collectingCountry id="50917673FF87F97815F9FED0FB82FE40" box="[961,1039,359,385]" name="Jordan" pageId="27" pageNumber="28">Jordan</collectingCountry>
[white, manuscript]; 3) 
<collectorName id="85735335FF87F9781730FE38FEE1FE68" box="[264,364,399,425]" pageId="27" pageNumber="28">Pará, N.</collectorName>
<collectingCountry id="50917673FF87F9781740FE38FE32FE68" box="[376,447,399,425]" name="Brazil" pageId="27" pageNumber="28">Brazil</collectingCountry>
[white, printed]; 4) 
<collectingCounty id="C1584E6FFF87F9781486FE38FD7FFE68" box="[702,754,399,425]" pageId="27" pageNumber="28">Karl</collectingCounty>
<collectingCountry id="50917673FF87F97814C6FE38FCC1FE68" box="[766,844,399,425]" name="Jordan" pageId="27" pageNumber="28">Jordan</collectingCountry>
coll., B.M. 1940–109 [white, printed]. Type locality: 
<collectingCountry id="50917673FF87F9781788FE00FE7AFE10" box="[432,503,439,465]" name="Brazil" pageId="27" pageNumber="28">Brazil</collectingCountry>
(Pará)
</materialsCitation>
.
</paragraph>
<paragraph id="283936E3FF87F9781730FDB0FEE3FDE1" blockId="27.[264,1324,519,1113]" box="[264,366,519,544]" pageId="27" pageNumber="28">
<emphasis id="1AF2EAF1FF87F9781730FDB0FEE3FDE1" box="[264,366,519,544]" italics="true" pageId="27" pageNumber="28">Remarks</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="609C6568FF87F9781700FD85FB01FB98" pageId="27" pageNumber="28" type="reference_group">
<paragraph id="283936E3FF87F9781700FD85FB7EFCD8" blockId="27.[264,1324,519,1113]" pageId="27" pageNumber="28">
<taxonomicName id="EF864D60FF87F9781700FD85FD99FD8A" ID-CoL="78NJ3" authorityName="Jordan" authorityYear="1894" box="[312,532,562,587]" class="Insecta" family="Anthribidae" genus="Ptychoderes" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="1AF2EAF1FF87F9781700FD85FD99FD8A" box="[312,532,562,587]" italics="true" pageId="27" pageNumber="28">Ptychoderes brevis</emphasis>
</taxonomicName>
belongs to the 
<taxonomicName id="EF864D60FF87F97814FDFD85FCC1FD8A" ID-CoL="8K9LP" authorityName="Mermudes &amp; Napp" authorityYear="2006" box="[709,844,562,587]" class="Insecta" family="Anthribidae" genus="Ptychoderes" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="crustatus">
<emphasis id="1AF2EAF1FF87F97814FDFD85FCC1FD8A" box="[709,844,562,587]" italics="true" pageId="27" pageNumber="28">P. crustatus</emphasis>
</taxonomicName>
<superScript id="DFF39BABFF87F9781574FD99FCD8FDFD" attach="left" box="[844,853,558,572]" fontSize="6" pageId="27" pageNumber="28">+</superScript>
group, being the sister group of 
<taxonomicName id="EF864D60FF87F97812F7FD85FDC0FDB5" authority="Mermudes and Napp, 2004" authorityName="Mermudes and Napp" authorityYear="2004" class="Insecta" family="Anthribidae" genus="Ptychoderes" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="jekeli">
<emphasis id="1AF2EAF1FF87F97812F7FD85FAA6FD8A" box="[1231,1323,562,587]" italics="true" pageId="27" pageNumber="28">P.jekeli</emphasis>
<bibRefCitation id="4C174B12FF87F9781730FDEDFDC0FDB5" author="Mermudes, J. R. M. &amp; Napp, D. S." box="[264,589,602,628]" pageId="27" pageNumber="28" pagination="27 - 30" refId="ref64325" refString="Mermudes, J. R. M. &amp; Napp, D. S. (2004). Duas novas especies de Ptychoderes Schoenherr (Coleoptera, Anthribidae, Anthribinae, Ptychoderini). Revista Brasileira de Entomologia, 48 (1), 27 - 30." type="journal article" year="2004">Mermudes and Napp, 2004</bibRefCitation>
</taxonomicName>
by two synapomorphies: pronotal depression with a median, somewhat rounded, weakly prominent tubercle (20 
<subScript id="B40234A6FF87F9781566FD25FCEBFD61" attach="none" box="[862,870,658,672]" fontSize="6" pageId="27" pageNumber="28">5</subScript>
), and glabrous area of profemora with a single coarse puncture (36 
<subScript id="B40234A6FF87F978145CFD0BFDE1FD0B" attach="right" box="[612,620,700,714]" fontSize="6" pageId="27" pageNumber="28">2</subScript>
). 
<taxonomicName id="EF864D60FF87F97814B1FD1AFD7CFD07" ID-CoL="78NJ3" authorityName="Jordan" authorityYear="1894" box="[649,753,685,710]" class="Insecta" family="Anthribidae" genus="Ptychoderes" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="1AF2EAF1FF87F97814B1FD1AFD7CFD07" box="[649,753,685,710]" italics="true" pageId="27" pageNumber="28">P. brevis</emphasis>
</taxonomicName>
differs from 
<emphasis id="1AF2EAF1FF87F97815A2FD1AFB8FFD07" box="[922,1026,685,710]" italics="true" pageId="27" pageNumber="28">
<taxonomicName id="EF864D60FF87F97815A2FD1AFC73FD07" authorityName="Mermudes and Napp" authorityYear="2004" box="[922,1022,685,710]" class="Insecta" family="Anthribidae" genus="Ptychoderes" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="jekeli">P. jekeli</taxonomicName>
,
</emphasis>
and also from all other species of 
<taxonomicName id="EF864D60FF87F97817BCFD6FFD99FD30" ID-CoL="8HQBP" authorityName="sensu Schoenherr" authorityYear="1833" box="[388,532,728,753]" class="Insecta" family="Anthribidae" genus="Ptychoderes" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="1AF2EAF1FF87F97817BCFD6FFD99FD30" box="[388,532,728,753]" italics="true" pageId="27" pageNumber="28">Ptychoderes</emphasis>
</taxonomicName>
, by the pygidium not impressed medially near apex, and ventrite V with projected, not acuminated, lateroapical angles, both autapomorphies of 
<taxonomicName id="EF864D60FF87F97812B1FCB7FB61FCD8" ID-CoL="78NJ3" authorityName="Jordan" authorityYear="1894" box="[1161,1260,768,793]" class="Insecta" family="Anthribidae" genus="Ptychoderes" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="1AF2EAF1FF87F97812B1FCB7FB61FCD8" box="[1161,1260,768,793]" italics="true" pageId="27" pageNumber="28">P. brevis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="283936E3FF87F9781700FC9FFB01FB98" blockId="27.[264,1324,519,1113]" pageId="27" pageNumber="28">
<taxonomicName id="EF864D60FF87F9781700FC9FFD94FC80" ID-CoL="78NJ3" authorityName="Jordan" authorityYear="1894" box="[312,537,808,833]" class="Insecta" family="Anthribidae" genus="Ptychoderes" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="1AF2EAF1FF87F9781700FC9FFD94FC80" box="[312,537,808,833]" italics="true" pageId="27" pageNumber="28">Ptychoderes brevis</emphasis>
</taxonomicName>
is similar to 
<taxonomicName id="EF864D60FF87F9781487FC9FFC5EFC80" ID-CoL="6WNJ6" authority="Jekel, 1855" authorityName="Jekel" authorityYear="1855" box="[703,979,807,833]" class="Insecta" family="Anthribidae" genus="Ptychoderes" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="callosus">
<emphasis id="1AF2EAF1FF87F9781487FC9FFCB2FC80" box="[703,831,808,833]" italics="true" pageId="27" pageNumber="28">P. callosus</emphasis>
<bibRefCitation id="4C174B12FF87F9781572FC90FC5EFC80" author="Jekel, H." box="[842,979,807,833]" pageId="27" pageNumber="28" pagination="1 - 2" refId="ref63597" refString="Jekel, H. (1855) Insecta Saundersiana or characters of undescribed insects in the collection of William Wilson Saunders (Coleoptera, Curculionidae). Part 1. 1 - 153, plates 1 - 2." type="book chapter" year="1855">Jekel, 1855</bibRefCitation>
</taxonomicName>
by the dorsal vestiture with olivaceous­green scales, differing by: 1) prosternum raised with coarse, deep, dense punctures; 2) prosternal furrow straight, deep and entire and 3) elytra with prominent basal gibbosities, lacking both anteapical tubercles and swollen interstriae. In 
<taxonomicName id="EF864D60FF87F978124CFC17FB78FC78" ID-CoL="6WNJ6" authorityName="Jekel" authorityYear="1855" box="[1140,1269,928,953]" class="Insecta" family="Anthribidae" genus="Ptychoderes" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="callosus">
<emphasis id="1AF2EAF1FF87F978124CFC17FB78FC78" box="[1140,1269,928,953]" italics="true" pageId="27" pageNumber="28">P. callosus</emphasis>
</taxonomicName>
, the prosternum is weakly raised, with moderately coarse, shallow, sparse punctures; the prosternal furrow is shallow, strongly curved in front of procoxae, and with a subtriangular prosternal projection at middle not interrupting the furrow; the elytra have weakly prominent basal gibbosities, anteapical tubercles and interstriae 3–9 swollen.
</paragraph>
</subSubSection>
<subSubSection id="609C6568FF87F9781730FB27FE6BFAE0" pageId="27" pageNumber="28" type="distribution">
<paragraph id="283936E3FF87F9781730FB27FCF8FB10" blockId="27.[264,885,1168,1233]" pageId="27" pageNumber="28">
<emphasis id="1AF2EAF1FF87F9781730FB27FD98FB68" box="[264,533,1168,1193]" italics="true" pageId="27" pageNumber="28">
Distribution (
<figureCitation id="B0BD2A66FF87F9781790FB27FD80FB68" box="[424,525,1168,1193]" captionStart="FIGURES 263–264. 263" captionStartId="123.[264,383,1767,1791]" captionTargetBox="[383,1204,328,1733]" captionTargetId="figure-62@123.[383,1205,318,1763]" captionTargetPageId="123" captionText="FIGURES 263–264. 263. Geographical distribution of P. longicollis (black circles), P. crustatus (white triangles), P. brevis (black squares), P. jekeli (black triangles). 264. Geographical distribution of P. jordani (black squares), P.depressus (black triangles), P. virgatus (white circles)." figureDoi="http://doi.org/10.5281/zenodo.5064137" httpUri="https://zenodo.org/record/5064137/files/figure.png" pageId="27" pageNumber="28">Fig. 263</figureCitation>
)
</emphasis>
<collectingCountry id="50917673FF87F9781700FB00FE58FB10" box="[312,469,1207,1233]" name="Colombia" pageId="27" pageNumber="28">COLOMBIA</collectingCountry>
and 
<collectingCountry id="50917673FF87F9781435FB00FDFAFB10" box="[525,631,1207,1233]" name="Brazil" pageId="27" pageNumber="28">BRAZIL</collectingCountry>
(Pará, 
<collectingRegion id="EA42F801FF87F97814F3FB00FCE4FB10" box="[715,873,1207,1233]" country="Brazil" name="Mato Grosso" pageId="27" pageNumber="28">Mato Grosso</collectingRegion>
).
</paragraph>
<paragraph id="283936E3FF87F9781730FABFFE6BFAE0" blockId="27.[264,1324,1288,1433]" box="[264,486,1288,1313]" pageId="27" pageNumber="28">
<emphasis id="1AF2EAF1FF87F9781730FABFFE6BFAE0" box="[264,486,1288,1313]" italics="true" pageId="27" pageNumber="28">Material examined</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="609C6568FF87F9781700FA98FEFDFA58" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="283936E3FF87F9781700FA98FEFDFA58" blockId="27.[264,1324,1288,1433]" pageId="27" pageNumber="28">
<materialsCitation id="98EE3CBEFF87F9781700FA98FEEFFAB0" collectionCode="XI" country="Colombia" location="Putumayo" pageId="27" pageNumber="28" specimenCount="1" stateProvince="Putumayo" typeStatus="holotype">
<collectingCountry id="50917673FF87F9781700FA98FE55FA88" box="[312,472,1327,1353]" name="Colombia" pageId="27" pageNumber="28">COLOMBIA</collectingCountry>
. 
<emphasis id="1AF2EAF1FF87F97817DDFA87FDD1FA88" box="[485,604,1328,1353]" italics="true" pageId="27" pageNumber="28">
<collectingRegion id="EA42F801FF87F97817DDFA87FDD1FA88" box="[485,604,1328,1353]" country="Colombia" name="Putumayo" pageId="27" pageNumber="28">Putumayo</collectingRegion>
</emphasis>
: 
<collectingRegion id="EA42F801FF87F9781457FA98FD4DFA88" box="[623,704,1327,1353]" country="Colombia" name="Narino" pageId="27" pageNumber="28">Nariño</collectingRegion>
(Villa Amazonica), 1 F (compared with 
<typeStatus id="F73D8841FF87F978128BFA98FA93FA88" box="[1203,1310,1327,1353]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
), 
<date id="5C381023FF87F9781730FAE0FEEFFAB0" box="[264,354,1367,1393]" pageId="27" pageNumber="28" value="1995-10-01" valueMax="1995-11-01" valueMin="1995-10-01">
1.X– 
<collectionCode id="4E97AE26FF87F9781707FAE0FEEFFAB0" box="[319,354,1367,1393]" pageId="27" pageNumber="28">XI</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="98EE3CBEFF87F978175AFAE0FD24FAB0" box="[354,681,1367,1393]" collectionCode="CPMT" collectorName="Mracek" country="Colombia" pageId="27" pageNumber="28" specimenCount="1" stateProvince="Narino" typeStatus="holotype">
.1995, 
<collectorName id="85735335FF87F9781789FAE0FD87FAB0" box="[433,522,1367,1393]" pageId="27" pageNumber="28">Mracek</collectorName>
col. (
<collectionCode id="4E97AE26FF87F9781473FAE0FD2CFAB0" box="[587,673,1367,1393]" pageId="27" pageNumber="28">CPMT</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="98EE3CBEFF87F978148CFAE0FEE6FA58" collectionCode="MZSP" collectorName="Isnard Rubim" country="Brazil" location="Mato Grosso" pageId="27" pageNumber="28" specimenCount="1" stateProvince="Mato Grosso" typeStatus="holotype">
<collectingCountry id="50917673FF87F978148CFAE0FCAFFAB0" box="[692,802,1367,1393]" name="Brazil" pageId="27" pageNumber="28">BRAZIL</collectingCountry>
. 
<emphasis id="1AF2EAF1FF87F9781515FAEFFC4BFAB0" box="[813,966,1368,1393]" italics="true" pageId="27" pageNumber="28">
<collectingRegion id="EA42F801FF87F9781515FAEFFC4BFAB0" box="[813,966,1368,1393]" country="Brazil" name="Mato Grosso" pageId="27" pageNumber="28">Mato Grosso</collectingRegion>
</emphasis>
: Sinop, 1 F, 
<collectorName id="85735335FF87F9781261FAE0FB74FAB0" box="[1113,1273,1367,1393]" pageId="27" pageNumber="28">Isnard Rubim</collectorName>
leg. (
<collectionCode id="4E97AE26FF87F9781728FAC8FEEEFA58" box="[272,355,1407,1433]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="27" pageNumber="28" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>