<document id="EDC312C97638816AD686CB1C7DC43BC3" ID-CLB-Dataset="21174" ID-DOI="10.11646/zootaxa.4629.3.1" ID-GBIF-Dataset="9419bc9b-dbe4-426c-a34e-f0034848ff99" ID-ISSN="1175-5326" ID-Zenodo-Dep="3995508" ID-ZooBank="1D47B452-0A40-4BC5-9BBA-C06DEB298C9A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1598059654494" checkinUser="felipe" docAuthor="Silva, Fernando A. B. &amp; Valois, Marcely" docDate="2019" docId="9D12E5573A4FFF8995F1FC3EFBFEC4E1" docLanguage="en" docName="zootaxa.4629.3.1.pdf" docOrigin="Zootaxa 4629 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scybalocanthon kelleri Pereira &amp; Martinez 1956" docType="treatment" docVersion="10" lastPageNumber="324" masterDocId="612B9D2F3A59FF9E9566F840FFF1C516" masterDocTitle="A taxonomic revision of the genus Scybalocanthon Martínez, 1948 (Coleoptera Scarabaeidae: Scarabaeinae: Deltochilini)" masterLastPageNumber="341" masterPageNumber="301" pageNumber="323" updateTime="1732598001296" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CF71BE161BD0C975CAAFA97A787AB7AB">A taxonomic revision of the genus Scybalocanthon Martínez, 1948 (Coleoptera Scarabaeidae: Scarabaeinae: Deltochilini)</mods:title>
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<mods:namePart id="A53E16CA1F12C5EF5C25065AD1F912E0">Silva, Fernando A. B.</mods:namePart>
<mods:affiliation id="0F4030E8E105CEC62927FCB5066EBB7B">Universidade Federal do Pará, Instituto de Ciências Biológicas, setor de Zoologia, Campus Belém. Rua Augusto Corrêa, 01. Guamá. Belém PA 66075 - 110, Brazil. E-mail: fernandoabsilva @ yahoo. com. br</mods:affiliation>
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<mods:namePart id="CC7556B24C1E0798B967FAA5FE8E5F73">Valois, Marcely</mods:namePart>
<mods:affiliation id="5B80854D1AEC2540FA2BA25FEDA1695F">Programa de Pós-graduação em Zoologia, Universidade Federal do Pará / Museu Paraense Emílio Goeldi. Rua Augusto Corrêa, 01. Guamá. Belém PA 66075 - 110, Brazil.</mods:affiliation>
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<treatment id="9D12E5573A4FFF8995F1FC3EFBFEC4E1" ID-DOI="http://doi.org/10.5281/zenodo.4335163" ID-GBIF-Taxon="167293220" ID-Zenodo-Dep="4335163" LSID="urn:lsid:plazi:treatment:9D12E5573A4FFF8995F1FC3EFBFEC4E1" httpUri="http://treatment.plazi.org/id/9D12E5573A4FFF8995F1FC3EFBFEC4E1" lastPageId="23" lastPageNumber="324" pageId="22" pageNumber="323" scope_family="Scarabaeidae" scope_order="Coleoptera" scope_subFamily="Scarabaeinae" scope_tribe="Deltochilini">
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<paragraph id="150454413A4FFF8895F1FC3EFD04C18E" blockId="22.[151,757,1149,1213]" box="[151,757,1149,1177]" pageId="22" pageNumber="323">
<heading id="4E4CE32D3A4FFF8895F1FC3EFD04C18E" bold="true" box="[151,757,1149,1177]" fontSize="11" level="1" pageId="22" pageNumber="323" reason="1">
<taxonomicName id="D2BB2FC23A4FFF8895F1FC3EFD04C18E" ID-CoL="4W3LL" authority="Pereira &amp; Martinez, 1956" authorityName="Pereira &amp; Martinez" authorityYear="1956" box="[151,757,1149,1177]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="kelleri">
<emphasis id="27CF88533A4FFF8895F1FC3EFD04C18E" bold="true" box="[151,757,1149,1177]" pageId="22" pageNumber="323">
<emphasis id="27CF88533A4FFF8895F1FC3EFE5CC18F" bold="true" box="[151,429,1150,1177]" italics="true" pageId="22" pageNumber="323">Scybalocanthon kelleri</emphasis>
<bibRefCitation id="712A29B03A4FFF8894D2FC3EFD04C18E" author="Pereira, F. S. &amp; Martinez, A." box="[436,757,1149,1176]" pageId="22" pageNumber="323" pagination="91 - 192" refId="ref47123" refString="Pereira, F. S. &amp; Martinez, A. (1956) Os generos de Canthonini Americanos (Col. Scarabaeidae). Revista Brasileira de Entomologia, 6, 91 - 192." type="journal article" year="1956">Pereira &amp; Martínez, 1956</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="5DA107CA3A4FFF8895F8FCE2FE6DC1AB" box="[158,412,1185,1213]" pageId="22" pageNumber="323" type="description">
<paragraph id="150454413A4FFF8895F8FCE2FE6DC1AB" blockId="22.[151,757,1149,1213]" box="[158,412,1185,1213]" pageId="22" pageNumber="323">
(
<figureCitation id="8D8048C43A4FFF8895C0FCE2FEF7C1AA" box="[166,262,1186,1212]" captionStart="FIGURE 1" captionStartId="8.[151,250,1804,1829]" captionTargetBox="[165,1428,599,1773]" captionTargetId="figure@8.[151,1436,594,1776]" captionTargetPageId="8" captionText="FIGURE 1. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. batesi; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N–O, S. maculatus; P, S. martinezi new species; Q–R; S. moniliatus; S. S. nigriceps; T. S. papaxibe new species. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3995510" httpUri="https://zenodo.org/record/3995510/files/figure.png" pageId="22" pageNumber="323">Figs. 1L</figureCitation>
, 
<figureCitation id="8D8048C43A4FFF889475FCE2FEC4C1AA" box="[275,309,1186,1212]" captionStart="FIGURE 4" captionStartId="14.[151,250,1663,1688]" captionTargetBox="[164,1424,190,1621]" captionTargetId="figure@14.[151,1436,181,1637]" captionTargetPageId="14" captionText="FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995518" httpUri="https://zenodo.org/record/3995518/files/figure.png" pageId="22" pageNumber="323">4L</figureCitation>
, 
<figureCitation id="8D8048C43A4FFF889426FCE1FE95C1AA" box="[320,356,1185,1212]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="22" pageNumber="323">5K</figureCitation>
, 
<figureCitation id="8D8048C43A4FFF889416FCE2FE65C1AB" box="[368,404,1186,1213]" captionStart="FIGURE 6" captionStartId="17.[151,250,1784,1809]" captionTargetBox="[179,1376,186,1734]" captionTargetId="figure@17.[174,1414,181,1759]" captionTargetPageId="17" captionText="FIGURE 6. Known distribution of species of Scybalocanthon." figureDoi="http://doi.org/10.5281/zenodo.3995522" httpUri="https://zenodo.org/record/3995522/files/figure.png" pageId="22" pageNumber="323">6A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5DA107CA3A4FFF8895F1FCA1FC81C043" pageId="22" pageNumber="323" type="reference_group">
<paragraph id="150454413A4FFF8895F1FCA1FF07C021" blockId="22.[151,1436,1248,1365]" pageId="22" pageNumber="323">
<treatmentCitationGroup id="35AB736F3A4FFF8895F1FCA1FF07C021" pageId="22" pageNumber="323">
<treatmentCitation id="941A72503A4FFF8895F1FCA1FD3CC1EF" author="Pereira, F. S. &amp; Martinez, A." box="[151,717,1248,1273]" page="114" pageId="22" pageNumber="323" year="1956">
<taxonomicName id="D2BB2FC23A4FFF8895F1FCA1FD3CC1EF" ID-CoL="4W3LL" authority="Pereira &amp; Martinez, 1956: 114" authorityName="Pereira &amp; Martinez" authorityPageNumber="114" authorityYear="1956" box="[151,717,1248,1273]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="kelleri">
<emphasis id="27CF88533A4FFF8895F1FCA1FE72C1EF" box="[151,387,1249,1273]" italics="true" pageId="22" pageNumber="323">Scybalocanthon kelleri</emphasis>
<bibRefCitation id="712A29B03A4FFF8894EDFCA0FD3CC1EF" author="Pereira, F. S. &amp; Martinez, A." box="[395,717,1248,1273]" pageId="22" pageNumber="323" pagination="91 - 192" refId="ref47123" refString="Pereira, F. S. &amp; Martinez, A. (1956) Os generos de Canthonini Americanos (Col. Scarabaeidae). Revista Brasileira de Entomologia, 6, 91 - 192." type="journal article" year="1956">Pereira &amp; Martínez, 1956: 114</bibRefCitation>
</taxonomicName>
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, 116 (original description and identification key); 
<treatmentCitation id="941A72503A4FFF8891BDFCA0FEDCC00E" author="Vulcano, M. A. &amp; Pereira, F. S." page="638" pageId="22" pageNumber="323" year="1964">
<bibRefCitation id="712A29B03A4FFF8891BDFCA0FEDCC00E" author="Vulcano, M. A. &amp; Pereira, F. S." pageId="22" pageNumber="323" pagination="570 - 685" refId="ref48402" refString="Vulcano, M. A. &amp; Pereira, F. S. (1964) Catalog of the Canthonini (Col. Scarab.) inhabiting the Western Hemisphere. Entomologischen Arbeiten aus dem Museum G. Frey, 15, 570 - 685." type="journal article" year="1964">Vulcano &amp; Pereira 1964: 638</bibRefCitation>
</treatmentCitation>
(catalog); 
<treatmentCitation id="941A72503A4FFF8894FCFCBFFD33C00E" author="Vulcano, M. A. &amp; Pereira, F. S." box="[410,706,1279,1304]" page="554" pageId="22" pageNumber="323" year="1967">
<bibRefCitation id="712A29B03A4FFF8894FCFCBFFD33C00E" author="Vulcano, M. A. &amp; Pereira, F. S." box="[410,706,1279,1304]" pageId="22" pageNumber="323" pagination="533 - 603" refId="ref48448" refString="Vulcano, M. A. &amp; Pereira, F. S. (1967) Sinopse dos Passalidae e Scarabaeidae s. str. da regiao Amazonica (Insecta, Coleoptera). Atas do Simposio sobre a Biota Amazonica, Zoologia, 5, 533 - 603." type="journal article" year="1967">Vulcano &amp; Pereira 1967: 554</bibRefCitation>
</treatmentCitation>
(distribution, identification key); 
<treatmentCitation id="941A72503A4FFF88917CFCBFFAB5C00E" author="Halffter, G. &amp; Martinez, A." box="[1050,1348,1279,1304]" page="67" pageId="22" pageNumber="323" year="1977">
<bibRefCitation id="712A29B03A4FFF88917CFCBFFAB5C00E" author="Halffter, G. &amp; Martinez, A." box="[1050,1348,1279,1304]" pageId="22" pageNumber="323" pagination="29 - 107" refId="ref45849" refString="Halffter, G. &amp; Martinez, A. (1977) Revision monografica de los Canthonina Americanos, IV parte. Clave para generos y subgeneros. Folia Entomologica Mexicana, 38, 29 - 107." type="journal article" year="1977">Halffter &amp; Martínez 1977: 67</bibRefCitation>
</treatmentCitation>
(species list).
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</paragraph>
<paragraph id="150454413A4FFF8895F1FD7DFC81C043" blockId="22.[151,1436,1248,1365]" box="[151,880,1340,1365]" pageId="22" pageNumber="323">
<treatmentCitationGroup id="35AB736F3A4FFF8895F1FD7DFC81C043" box="[151,880,1340,1365]" pageId="22" pageNumber="323">
<treatmentCitation id="941A72503A4FFF8895F1FD7DFE1FC043" author="Krajcik, M." box="[151,494,1340,1365]" page="28" pageId="22" pageNumber="323" year="2006">
<taxonomicName id="D2BB2FC23A4FFF8895F1FD7DFE1FC043" authority=": Krajcik 2006: 28" authorityName=": Krajcik" authorityPageNumber="28" authorityYear="2006" box="[151,494,1340,1365]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="kelleri">
<emphasis id="27CF88533A4FFF8895F1FD7DFEC7C043" box="[151,310,1341,1365]" italics="true" pageId="22" pageNumber="323">Canthon kelleri</emphasis>
: 
<bibRefCitation id="712A29B03A4FFF889425FD7CFE1FC043" author="Krajcik, M." box="[323,494,1340,1365]" pageId="22" pageNumber="323" pagination="1 - 189" refId="ref46404" refString="Krajcik, M. (2006) Checklist of Scarabaeoidea of the world. 1. Scarabaeinae (Coleoptera: Scarabaeidae: Scarabaeinae). Animma. X, Supplement 3, 1 - 189." type="journal article" year="2006">Krajcik 2006: 28</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(catalog); 
<treatmentCitation id="941A72503A4FFF88973AFD7CFCF9C043" author="Krajcik, M." box="[604,776,1340,1365]" page="63" pageId="22" pageNumber="323" year="2012">
<bibRefCitation id="712A29B03A4FFF88973AFD7CFCF9C043" author="Krajcik, M." box="[604,776,1340,1365]" pageId="22" pageNumber="323" pagination="1 - 278" refId="ref46440" refString="Krajcik, M. (2012) Checklist of the world Scarabaeoidea. Animma. X, Supplement 5, 1 - 278." type="journal article" year="2012">Krajcik 2012: 63</bibRefCitation>
</treatmentCitation>
(catalog).
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</paragraph>
</subSubSection>
<subSubSection id="5DA107CA3A4FFF8895F1FD3DFBAFC312" pageId="22" pageNumber="323" type="diagnosis">
<paragraph id="150454413A4FFF8895F1FD3DFBAFC312" blockId="22.[151,1437,1405,2044]" pageId="22" pageNumber="323">
<emphasis id="27CF88533A4FFF8895F1FD3DFEFBC08E" bold="true" box="[151,266,1405,1432]" pageId="22" pageNumber="323">Diagnosis</emphasis>
. Specimens of 
<taxonomicName id="D2BB2FC23A4FFF8894D0FD3FFDEBC08E" authorityName=": Krajcik" authorityYear="2006" box="[438,538,1406,1433]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="kelleri">
<emphasis id="27CF88533A4FFF8894D0FD3FFDEBC08E" box="[438,538,1406,1433]" italics="true" pageId="22" pageNumber="323">S. kelleri</emphasis>
</taxonomicName>
(
<figureCitation id="8D8048C43A4FFF88974EFD3EFD8AC08E" box="[552,635,1406,1432]" captionStart="FIGURE 1" captionStartId="8.[151,250,1804,1829]" captionTargetBox="[165,1428,599,1773]" captionTargetId="figure@8.[151,1436,594,1776]" captionTargetPageId="8" captionText="FIGURE 1. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. batesi; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N–O, S. maculatus; P, S. martinezi new species; Q–R; S. moniliatus; S. S. nigriceps; T. S. papaxibe new species. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3995510" httpUri="https://zenodo.org/record/3995510/files/figure.png" pageId="22" pageNumber="323">Fig. 1L</figureCitation>
) are similar to some specimens of 
<taxonomicName id="D2BB2FC23A4FFF889166FD3FFB66C08E" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1855" box="[1024,1175,1406,1433]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="sexspilotus">
<emphasis id="27CF88533A4FFF889166FD3FFB66C08E" box="[1024,1175,1406,1433]" italics="true" pageId="22" pageNumber="323">S. sexspilotus</emphasis>
</taxonomicName>
(
<figureCitation id="8D8048C43A4FFF8891C0FD3EFB0AC08E" box="[1190,1275,1406,1432]" captionStart="FIGURE 2" captionStartId="9.[151,250,1178,1203]" captionTargetBox="[165,1424,188,1136]" captionTargetId="figure@9.[151,1436,181,1152]" captionTargetPageId="9" captionText="FIGURE 2. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon pinopterus; B, S. pygidialis; C–G, S. sexpilotus; H–J, S. trimaculatus; K, S. uniplagiatus. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3995514" httpUri="https://zenodo.org/record/3995514/files/figure.png" pageId="22" pageNumber="323">Fig. 2C</figureCitation>
) in having the pronotum bicolored, with a transverse black spot; aedeagus with parameres elongate, subrectangular, not excavated ventrally (
<figureCitation id="8D8048C43A4FFF88946AFD86FE65C0F7" box="[268,404,1478,1505]" captionStart="FIGURE 4" captionStartId="14.[151,250,1663,1688]" captionTargetBox="[164,1424,190,1621]" captionTargetId="figure@14.[151,1436,181,1637]" captionTargetPageId="14" captionText="FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995518" httpUri="https://zenodo.org/record/3995518/files/figure.png" pageId="22" pageNumber="323">Figs. 4L, U</figureCitation>
); and endophallus lacking bristles or microbristles close to the FLP sclerite (
<figureCitation id="8D8048C43A4FFF889062FD86FA61C0F7" box="[1284,1424,1477,1505]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="22" pageNumber="323">Figs. 5K, U</figureCitation>
). However, they can be distinguished by head, pygidium and femora uniformly colored.
</paragraph>
</subSubSection>
<subSubSection id="5DA107CA3A4FFF8895A1FE4DFD81C286" pageId="22" pageNumber="323" type="description">
<paragraph id="150454413A4FFF8895A1FE4DFD81C286" blockId="22.[151,1437,1405,2044]" pageId="22" pageNumber="323">
<emphasis id="27CF88533A4FFF8895A1FE4DFE56C33E" bold="true" box="[199,423,1549,1576]" pageId="22" pageNumber="323">Description. Body.</emphasis>
Oval, lateral edges rounded. Surface completely microgranulate, with small and dense punctures randomly distributed. 
<emphasis id="27CF88533A4FFF889763FE71FDB8C35A" bold="true" box="[517,585,1585,1612]" pageId="22" pageNumber="323">Color</emphasis>
. Head and elytra opaque, pronotum with metallic sheen. Most of pronotum and elytra, lateral portion of hypomera, lateral portion of abdominal ventrites, and pygidium yellow or dark brown. Other parts black. 
<emphasis id="27CF88533A4FFF88940FFE3AFE37C382" bold="true" box="[361,454,1658,1684]" pageId="22" pageNumber="323">Length.</emphasis>
7.7–9.0 mm. 
<emphasis id="27CF88533A4FFF889702FE3AFD4CC382" bold="true" box="[612,701,1658,1684]" pageId="22" pageNumber="323">Thorax</emphasis>
. Pronotum with transverse black spot in a butterfly or bat shape. Anterior angles of pronotum acute (approximately 80°). Lateral margin regularly curved outward, not forming an angle at the middle portion. 
<emphasis id="27CF88533A4FFF88949BFE82FDB8C3CA" bold="true" box="[509,585,1730,1756]" pageId="22" pageNumber="323">Elytra</emphasis>
. Striae thin and shiny, weakly impressed, punctures almost inconspicuous. Eighth stria with a thin carina at the anterior portion. 
<emphasis id="27CF88533A4FFF889796FEA5FC9BC216" bold="true" box="[752,874,1765,1792]" pageId="22" pageNumber="323">Aedeagus.</emphasis>
Parameres slightly asymmetrical, with subrectangular shape, truncate apically (
<figureCitation id="8D8048C43A4FFF88948DFF4AFDB3C232" box="[491,578,1802,1828]" captionStart="FIGURE 4" captionStartId="14.[151,250,1663,1688]" captionTargetBox="[164,1424,190,1621]" captionTargetId="figure@14.[151,1436,181,1637]" captionTargetPageId="14" captionText="FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995518" httpUri="https://zenodo.org/record/3995518/files/figure.png" pageId="22" pageNumber="323">Fig. 4L</figureCitation>
). Dorsal margin of parameres slightly curved inward from the basal to medial portions and slightly curved outward at the apical portion. Ventral margin of parameres substraight. SRP circular, with curved handle-shaped extension (
<figureCitation id="8D8048C43A4FFF88972FFF12FD54C27A" box="[585,677,1873,1900]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="22" pageNumber="323">Fig. 5K</figureCitation>
). FLP I-shaped (
<figureCitation id="8D8048C43A4FFF889606FF12FC4AC27A" box="[864,955,1873,1900]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="22" pageNumber="323">Fig. 5K</figureCitation>
). A+SA with two superposed and elongate sclerites (
<figureCitation id="8D8048C43A4FFF889464FF36FEACC286" box="[258,349,1909,1936]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="22" pageNumber="323">Fig. 5K</figureCitation>
). 
<collectingCountry id="6DAC14D13A4FFF889409FF36FE62C286" box="[367,403,1910,1936]" name="American Samoa" pageId="22" pageNumber="323">AS</collectingCountry>
I-shaped (
<figureCitation id="8D8048C43A4FFF88976FFF36FD92C286" box="[521,611,1909,1936]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="22" pageNumber="323">Fig. 5K</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5DA107CA3A4FFF8995A1FFDAFB31C455" lastPageId="23" lastPageNumber="324" pageId="22" pageNumber="323" type="materials_examined">
<paragraph id="150454413A4FFF8895A1FFDAFEA0C2EA" blockId="22.[151,1437,1405,2044]" pageId="22" pageNumber="323">
<emphasis id="27CF88533A4FFF8895A1FFDAFC86C2A2" bold="true" box="[199,887,1945,1972]" pageId="22" pageNumber="323">
Type material. 
<taxonomicName id="D2BB2FC23A4FFF88941FFFD9FC83C2A2" authority="Pereira &amp; Martinez" authorityName="Pereira &amp; Martinez" authorityYear="1956" box="[377,882,1945,1972]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="kelleri">
<emphasis id="27CF88533A4FFF88941FFFD9FD70C2A2" bold="true" box="[377,641,1945,1972]" italics="true" pageId="22" pageNumber="323">Scybalocanthon kelleri</emphasis>
Pereira &amp; Martínez
</taxonomicName>
,
</emphasis>
<materialsCitation id="A5D35E1C3A4FFF88961BFFDAFF56C2CE" ID-GBIF-Occurrence="2847261459" collectionCode="MZSP" pageId="22" pageNumber="323" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="27CF88533A4FFF88961BFFDAFC0FC2A2" bold="true" box="[893,1022,1945,1972]" pageId="22" pageNumber="323">
<typeStatus id="CA00EAE33A4FFF88961BFFDAFC15C2A2" box="[893,996,1946,1972]" pageId="22" pageNumber="323" type="holotype">holotype</typeStatus>
<specimenCount id="03BD9FC83A4FFF88968FFFD9FC0FC2A2" box="[1001,1022,1945,1972]" pageId="22" pageNumber="323" type="male">♂</specimenCount>
</emphasis>
(
<collectionCode id="73AACC843A4FFF88916AFFDAFBADC2A2" box="[1036,1116,1946,1972]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="22" pageNumber="323" type="Museum">MZSP</collectionCode>
): “Sosomoco / Ost Columb.” //
</materialsCitation>
<materialsCitation id="A5D35E1C3A4FFF8895CBFFFEFEBFC2EA" ID-GBIF-Occurrence="2847261483" collectorName="F. S. Pereira" determinerName="P. Pereira" pageId="22" pageNumber="323" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
“ 
<typeStatus id="CA00EAE33A4FFF8895D0FFFEFE93C2CE" box="[182,354,1982,2008]" pageId="22" pageNumber="323" type="holotype">HOLOTYPUS</typeStatus>
” / 
<emphasis id="27CF88533A4FFF88941CFFFDFE73C2CE" bold="true" box="[378,386,1981,2008]" pageId="22" pageNumber="323">/</emphasis>
“ 
<taxonomicName id="D2BB2FC23A4FFF8894F2FFFEFDB6C2CE" authorityName="Martinez" authorityYear="1948" box="[404,583,1982,2008]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="genus">
<emphasis id="27CF88533A4FFF8894F2FFFEFDB6C2CE" box="[404,583,1982,2008]" italics="true" pageId="22" pageNumber="323">Scybalocanthon</emphasis>
</taxonomicName>
/ 
<emphasis id="27CF88533A4FFF88973CFFFEFD12C2CE" box="[602,739,1982,2008]" italics="true" pageId="22" pageNumber="323">
<taxonomicName id="D2BB2FC23A4FFF88973CFFFEFD53C2CE" authority="Silva &amp; Valois, 2019" authorityName="Silva &amp; Valois" authorityYear="2019" box="[602,674,1982,2008]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="kelleri" status="sp. nov.">kelleri</taxonomicName>
<taxonomicNameLabel id="3CFC35283A4FFF8897CEFFFFFD12C2CE" box="[680,739,1983,2008]" pageId="22" pageNumber="323" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
<specimenCount id="03BD9FC83A4FFF88978FFFFFFD0FC2CF" box="[745,766,1983,2009]" pageId="22" pageNumber="323" type="male">♂</specimenCount>
/ 
<emphasis id="27CF88533A4FFF889677FFFFFC6AC2CE" box="[785,923,1983,2009]" italics="true" pageId="22" pageNumber="323">
<collectorName id="B84E31973A4FFF889677FFFFFC6AC2CE" box="[785,923,1983,2009]" pageId="22" pageNumber="323">F. S. Pereira</collectorName>
</emphasis>
” // “ 
<taxonomicName id="D2BB2FC23A4FFF8896B6FFFEFBC1C2CE" authorityName="Hoffmannsegg" authorityYear="1817" box="[976,1072,1982,2008]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="genus">
<emphasis id="27CF88533A4FFF8896B6FFFEFBC1C2CE" box="[976,1072,1982,2008]" italics="true" pageId="22" pageNumber="323">Canthon</emphasis>
</taxonomicName>
/ 
<taxonomicName id="D2BB2FC23A4FFF889122FFFFFB5DC2CE" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[1092,1196,1983,2008]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="27CF88533A4FFF889122FFFFFB5DC2CE" box="[1092,1196,1983,2008]" italics="true" pageId="22" pageNumber="323">nigriceps</emphasis>
</taxonomicName>
/ 
<emphasis id="27CF88533A4FFF8891A6FFFFFB00C2CE" box="[1216,1265,1983,2008]" italics="true" pageId="22" pageNumber="323">Har.</emphasis>
/ 
<determinerName id="E0D8B8853A4FFF889062FFFFFA9CC2CE" box="[1284,1389,1982,2009]" pageId="22" pageNumber="323">P. Pereira</determinerName>
det. 951” // “10196”
</materialsCitation>
.
</paragraph>
<paragraph id="150454413A4EFF8995A1F8D9FB31C455" blockId="23.[151,1437,152,503]" pageId="23" pageNumber="324">
<emphasis id="27CF88533A4EFF8995A1F8D9FE45C5A5" bold="true" box="[199,436,152,179]" pageId="23" pageNumber="324">Non-type material [</emphasis>
<materialsCitation id="A5D35E1C3A4EFF8994D2F8D9FE06C5ED" ID-GBIF-Occurrence="2847261665" collectingDate="1897-01" collectingDateMax="1997-04" collectingDateMin="1897-01" collectionCode="CMNC, CEMT" collectorName="S. &amp; J. Peck &amp; Finca El Esfuerzo &amp; Buena Vista" country="Colombia" elevation="1000" latitude="3.0455556" location="Quebrada Susamuco" longLatPrecision="21" longitude="-73.595" municipality="Sosomoco" pageId="23" pageNumber="324" specimenCount="22" specimenCount-female="9" specimenCount-male="13" stateProvince="Meta">
<emphasis id="27CF88533A4EFF8994D2F8D9FCBBC5A5" bold="true" box="[436,842,152,179]" pageId="23" pageNumber="324">
<specimenCount id="03BD9FC83A4EFF8994D2F8D9FDE3C5A5" box="[436,530,153,179]" pageId="23" pageNumber="324" type="male">7 males</specimenCount>
, 
<specimenCount id="03BD9FC83A4EFF899746F8D9FD66C5A5" box="[544,663,152,179]" pageId="23" pageNumber="324" type="female">5 females</specimenCount>
]: 
<collectingCountry id="6DAC14D13A4EFF8997CDF8D8FCBBC5A5" box="[683,842,152,179]" name="Colombia" pageId="23" pageNumber="324">COLOMBIA</collectingCountry>
</emphasis>
: 
<collectingMunicipality id="F560CE3B3A4EFF89963CF8D9FC23C5A5" box="[858,978,153,179]" pageId="23" pageNumber="324">Sosomoco</collectingMunicipality>
, [no date], [anonymous]— 
<specimenCount id="03BD9FC83A4EFF89906DF8D8FAD7C5A5" box="[1291,1318,152,179]" pageId="23" pageNumber="324" type="male">1♂</specimenCount>
(
<collectionCode id="73AACC843A4EFF899051F8D9FA61C5A5" box="[1335,1424,153,179]" pageId="23" pageNumber="324">CMNC</collectionCode>
); 
<collectingRegion id="D77F9AA33A4EFF8995F1F8FDFF16C5C1" box="[151,231,189,215]" country="Colombia" name="Meta" pageId="23" pageNumber="324">META</collectingRegion>
, 
<location id="1064029A3A4EFF899590F8FDFE12C5C1" LSID="urn:lsid:plazi:treatment:9D12E5573A4FFF8995F1FC3EFBFEC4E1:1064029A3A4EFF899590F8FDFE12C5C1" box="[246,483,189,215]" country="Colombia" latitude="3.0455556" longLatPrecision="21" longitude="-73.595" municipality="Sosomoco" name="Quebrada Susamuco" pageId="23" pageNumber="324" stateProvince="Meta">Quebrada Susamuco</location>
, 
<locationDeviation id="04B64D633A4EFF899494F8FDFCF2C5C1" box="[498,771,189,215]" location="Villavicencio" pageId="23" pageNumber="324">
<quantity id="D243F9A43A4EFF899494F8FDFDCCC5C1" box="[498,573,189,215]" metricMagnitude="4" metricUnit="m" metricValue="2.3" pageId="23" pageNumber="324" unit="km" value="23.0">23 km</quantity>
W Villavicencio
</locationDeviation>
, 
<quantity id="D243F9A43A4EFF899674F8FDFC9BC5C1" box="[786,874,189,215]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="324" unit="m" value="1000.0">
<elevation id="9E96B3723A4EFF899674F8FDFC9BC5C1" box="[786,874,189,215]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="324" unit="m" value="1000.0">1000 m</elevation>
</quantity>
(
<geoCoordinate id="708F32863A4EFF899618F8FDFC03C5C1" box="[894,1010,189,215]" degrees="4" direction="north" minutes="12" orientation="latitude" pageId="23" pageNumber="324" precision="15" seconds="4" value="4.201111">4°12’4’’N</geoCoordinate>
, 
<geoCoordinate id="708F32863A4EFF899167F8FDFB69C5C1" box="[1025,1176,189,215]" degrees="73" direction="west" minutes="47" orientation="longitude" pageId="23" pageNumber="324" precision="15" seconds="40" value="-73.79444">73°47’40’’W</geoCoordinate>
), 
<date id="610572813A4EFF8991D6F8FCFACBC5C1" box="[1200,1338,188,215]" pageId="23" pageNumber="324" value="1972-03-03" valueMax="1972-03-05" valueMin="1972-03-03">
<collectingDate id="71418B693A4EFF8991D6F8FCFACBC5C1" box="[1200,1338,188,215]" pageId="23" pageNumber="324" value="1972-03-03" valueMax="1972-03-05" valueMin="1972-03-03">3–5.iii.1972</collectingDate>
</date>
, 
<collectorName id="B84E31973A4EFF89902FF8FDFF21C5ED" pageId="23" pageNumber="324">S. &amp; J. Peck</collectorName>
, forest— 
<specimenCount id="03BD9FC83A4EFF89945AF8A1FEABC5ED" box="[316,346,225,251]" pageId="23" pageNumber="324" type="male">3♂</specimenCount>
<specimenCount id="03BD9FC83A4EFF899407F8A1FE75C5ED" box="[353,388,225,251]" pageId="23" pageNumber="324" type="female">4♀</specimenCount>
(
<collectionCode id="73AACC843A4EFF8994F4F8A1FE1AC5ED" box="[402,491,225,251]" pageId="23" pageNumber="324">CMNC</collectionCode>
);
</materialsCitation>
<materialsCitation id="A5D35E1C3A4EFF89949BF8A1FBACC409" ID-GBIF-Occurrence="2847261371" collectingDate="1897-01" collectingDateMax="1997-04" collectingDateMin="1897-01" collectionCode="CMNC, CEMT" collectorName="S. &amp; J. Peck &amp; Finca El Esfuerzo &amp; Buena Vista" country="Colombia" elevation="1000" latitude="3.0455556" location="Quebrada Susamuco" longLatPrecision="21" longitude="-73.595" municipality="Sosomoco" pageId="23" pageNumber="324" specimenCount="22" specimenCount-female="9" specimenCount-male="13" stateProvince="Meta">
Vista Hermosa, 
<collectorName id="B84E31973A4EFF8997D6F8A1FC89C5ED" box="[688,888,225,251]" pageId="23" pageNumber="324">Finca El Esfuerzo</collectorName>
(
<geoCoordinate id="708F32863A4EFF8996E0F8A0FBE6C5ED" box="[902,1047,224,251]" degrees="73" direction="west" minutes="35" orientation="longitude" pageId="23" pageNumber="324" precision="15" seconds="42" value="-73.595">73°35’42”W</geoCoordinate>
, 
<geoCoordinate id="708F32863A4EFF899144F8A1FB63C5ED" box="[1058,1170,225,251]" degrees="3" direction="north" minutes="2" orientation="latitude" pageId="23" pageNumber="324" precision="15" seconds="44" value="3.0455556">3°2’44”N</geoCoordinate>
), 
<date id="610572813A4EFF8991C0F8A1FB0DC5ED" box="[1190,1276,225,251]" pageId="23" pageNumber="324" value="1997-04">
<collectingDate id="71418B693A4EFF8991C0F8A1FB0DC5ED" box="[1190,1276,225,251]" pageId="23" pageNumber="324" value="1997-04">iv.1997</collectingDate>
</date>
, 
<collectingMethod id="CCFA2C563A4EFF899060F8A1FAB8C5ED" box="[1286,1353,225,251]" pageId="23" pageNumber="324">pitfall</collectingMethod>
human feces, 
<quantity id="D243F9A43A4EFF8995B9F945FED8C409" box="[223,297,261,287]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="324" unit="m" value="200.0">
<elevation id="9E96B3723A4EFF8995B9F945FED8C409" box="[223,297,261,287]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="324" unit="m" value="200.0">200 m</elevation>
</quantity>
, A. Lopera— 
<specimenCount id="03BD9FC83A4EFF8994AFF945FE14C409" box="[457,485,261,287]" pageId="23" pageNumber="324" type="male">1♂</specimenCount>
(
<collectionCode id="73AACC843A4EFF899492F945FDB9C409" box="[500,584,261,287]" pageId="23" pageNumber="324">CEMT</collectionCode>
); 
<collectorName id="B84E31973A4EFF89973AF945FD17C409" box="[604,742,261,287]" pageId="23" pageNumber="324">Buena Vista</collectorName>
, 
<date id="610572813A4EFF899794F945FCCAC409" box="[754,827,261,287]" pageId="23" pageNumber="324" value="1897-01">
<collectingDate id="71418B693A4EFF899794F945FCCAC409" box="[754,827,261,287]" pageId="23" pageNumber="324" value="1897-01">i.1897</collectingDate>
</date>
, S. Burger— 
<specimenCount id="03BD9FC83A4EFF8996B0F945FC03C409" box="[982,1010,261,287]" pageId="23" pageNumber="324" type="male">1♂</specimenCount>
(
<collectionCode id="73AACC843A4EFF899167F945FBA4C409" box="[1025,1109,261,287]" pageId="23" pageNumber="324">CEMT</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="A5D35E1C3A4EFF89910EF944FB4CC455" ID-GBIF-Occurrence="2847261407" collectingDate="1986-02-15" collectingDateMax="1986-02-21" collectingDateMin="1986-02-15" collectionCode="CMNC" collectorName="F. Genier" country="Ecuador" elevation="450" latitude="-0.9891667" location="Tena" longLatPrecision="21" longitude="-77.81528" pageId="23" pageNumber="324" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Napo">
<emphasis id="27CF88533A4EFF89910EF944FB07C409" bold="true" box="[1128,1270,260,287]" pageId="23" pageNumber="324">
<collectingCountry id="6DAC14D13A4EFF89910EF944FB07C409" box="[1128,1270,260,287]" name="Ecuador" pageId="23" pageNumber="324">ECUADOR</collectingCountry>
</emphasis>
: 
<collectingRegion id="D77F9AA33A4EFF899063F945FAA5C409" box="[1285,1364,261,287]" country="Ecuador" name="Napo" pageId="23" pageNumber="324">NAPO</collectingRegion>
, 
<location id="1064029A3A4EFF899039F945FA69C409" LSID="urn:lsid:plazi:treatment:9D12E5573A4FFF8995F1FC3EFBFEC4E1:1064029A3A4EFF899039F945FA69C409" box="[1375,1432,261,287]" country="Ecuador" latitude="-0.9891667" longLatPrecision="21" longitude="-77.81528" name="Tena" pageId="23" pageNumber="324" stateProvince="Napo">Tena</location>
, 
<quantity id="D243F9A43A4EFF8995F1F968FEE7C455" box="[151,278,296,323]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="23" pageNumber="324" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">
<elevation id="9E96B3723A4EFF8995F1F968FEE7C455" box="[151,278,296,323]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="23" pageNumber="324" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">400–500 m</elevation>
</quantity>
(
<geoCoordinate id="708F32863A4EFF899440F968FE52C455" box="[294,419,296,323]" degrees="0" direction="south" minutes="59" orientation="latitude" pageId="23" pageNumber="324" precision="15" seconds="21" value="-0.9891667">0°59’21’’S</geoCoordinate>
, 
<geoCoordinate id="708F32863A4EFF8994C9F968FDB4C455" box="[431,581,296,323]" degrees="77" direction="west" minutes="48" orientation="longitude" pageId="23" pageNumber="324" precision="15" seconds="55" value="-77.81528">77°48’55’’W</geoCoordinate>
), 
<date id="610572813A4EFF89973CF968FD06C455" box="[602,759,296,323]" pageId="23" pageNumber="324" value="1986-02-15" valueMax="1986-02-21" valueMin="1986-02-15">
<collectingDate id="71418B693A4EFF89973CF968FD06C455" box="[602,759,296,323]" pageId="23" pageNumber="324" value="1986-02-15" valueMax="1986-02-21" valueMin="1986-02-15">15–21.ii.1986</collectingDate>
</date>
, 
<collectorName id="B84E31973A4EFF899665F969FC9CC455" box="[771,877,297,323]" pageId="23" pageNumber="324">F. Génier</collectorName>
, feces trap— 
<specimenCount id="03BD9FC83A4EFF899161F969FBD2C455" box="[1031,1059,297,323]" pageId="23" pageNumber="324" type="male">1♂</specimenCount>
<specimenCount id="03BD9FC83A4EFF89914CF969FBBCC455" box="[1066,1101,297,323]" pageId="23" pageNumber="324" type="female">1♀</specimenCount>
(
<collectionCode id="73AACC843A4EFF89913AF969FB44C455" box="[1116,1205,297,323]" pageId="23" pageNumber="324">CMNC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5DA107CA3A4EFF8995A1F90CFF35C4B9" pageId="23" pageNumber="324" type="distribution">
<paragraph id="150454413A4EFF8995A1F90CFF35C4B9" blockId="23.[151,1437,152,503]" pageId="23" pageNumber="324">
<emphasis id="27CF88533A4EFF8995A1F90CFE90C471" bold="true" box="[199,353,332,359]" pageId="23" pageNumber="324">Distribution.</emphasis>
Known from 
<collectingCountry id="6DAC14D13A4EFF89949BF90DFD9AC471" box="[509,619,333,359]" name="Colombia" pageId="23" pageNumber="324">Colombia</collectingCountry>
and 
<collectingCountry id="6DAC14D13A4EFF8997C6F90DFD0CC471" box="[672,765,333,359]" name="Ecuador" pageId="23" pageNumber="324">Ecuador</collectingCountry>
(
<figureCitation id="8D8048C43A4EFF89966AF90DFC97C47E" box="[780,870,333,360]" captionStart="FIGURE 6" captionStartId="17.[151,250,1784,1809]" captionTargetBox="[179,1376,186,1734]" captionTargetId="figure@17.[174,1414,181,1759]" captionTargetPageId="17" captionText="FIGURE 6. Known distribution of species of Scybalocanthon." figureDoi="http://doi.org/10.5281/zenodo.3995522" httpUri="https://zenodo.org/record/3995522/files/figure.png" pageId="23" pageNumber="324">Fig. 6A</figureCitation>
). 
<emphasis id="27CF88533A4EFF89961FF90CFBCFC471" bold="true" box="[889,1086,332,359]" pageId="23" pageNumber="324">Endemism areas</emphasis>
: 
<emphasis id="27CF88533A4EFF89912AF90CFAB1C471" bold="true" box="[1100,1344,332,359]" pageId="23" pageNumber="324">Brazilian sub-region</emphasis>
: Pacific dominion: Sabana and Magdalena provinces; Boreal Brazilian dominion: 
<collectingRegion id="D77F9AA33A4EFF8996B8F931FB74C49D" box="[990,1157,369,395]" country="Ecuador" name="Napo" pageId="23" pageNumber="324">Napo province</collectingRegion>
(see 
<bibRefCitation id="712A29B03A4EFF8991A5F931FA9BC49D" author="Morrone, J. J." box="[1219,1386,369,395]" pageId="23" pageNumber="324" pagination="1 - 110" refId="ref47051" refString="Morrone, J. J. (2014) Biogeographical regionalisation of the Neotropical region. Zootaxa, 3782 (1), 1 - 110. https: // doi. org / 10.11646 / zootaxa. 3782.1.1" type="journal article" year="2014">Morrone 2014</bibRefCitation>
; fig. 12).
</paragraph>
</subSubSection>
<subSubSection id="5DA107CA3A4EFF8995A1F9F9FBFEC4E1" pageId="23" pageNumber="324" type="discussion">
<paragraph id="150454413A4EFF8995A1F9F9FBFEC4E1" blockId="23.[151,1437,152,503]" pageId="23" pageNumber="324">
<emphasis id="27CF88533A4EFF8995A1F9F9FEC5C4C5" bold="true" box="[199,308,441,467]" pageId="23" pageNumber="324">Remarks</emphasis>
. “Sosomoco” is apparently an incorrect citation for quebrada Susumuco in the eastern portion of the 
<collectingRegion id="D77F9AA33A4EFF8995F1F99DFEA7C4E1" box="[151,342,477,503]" country="Colombia" name="Meta" pageId="23" pageNumber="324">Meta department</collectingRegion>
in 
<collectingCountry id="6DAC14D13A4EFF89941FF99DFE16C4E1" box="[377,487,477,503]" name="Colombia" pageId="23" pageNumber="324">Colombia</collectingCountry>
(
<bibRefCitation id="712A29B03A4EFF899490F99DFC90C4E1" author="Maldonado-Ocampo, J. A. &amp; Ortega-Lara, A. &amp; Usma Oviedo, J. S. &amp; Galvis, G. &amp; Villa-Navarro, F. &amp; Vasquez, L. &amp; Prada-Pedreros, S. &amp; Rodriguez, C. A." box="[502,865,476,503]" pageId="23" pageNumber="324" refId="ref46464" refString="Maldonado-Ocampo, J. A., Ortega-Lara, A., Usma Oviedo, J. S., Galvis, G., Villa-Navarro, F., Vasquez, L., Prada-Pedreros, S. &amp; Rodriguez, C. A. (2005) Peces de los Andes de Colombia. Guia de campo. Instituto de Investigacion de Recursos Biologicos &quot; Alexander von Humboldt, &quot; Bogota, 346 pp." type="book" year="2005">
Maldonado-Ocampo 
<emphasis id="27CF88533A4EFF899781F99EFCE9C4E1" box="[743,792,477,503]" italics="true" pageId="23" pageNumber="324">et al</emphasis>
. 2005
</bibRefCitation>
; 
<bibRefCitation id="712A29B03A4EFF89960BF99DFBF2C4E1" author="Salazar, J. A." box="[877,1027,477,503]" pageId="23" pageNumber="324" pagination="143 - 156" refId="ref47445" refString="Salazar, J. A. (2008) Mariposas Colombianas X. Apuntes sobre la variacion cromatica en hembras de Pterourus cacicus Lucas, 1852 (Lepidoptera: Papilionidae). Boletin Cientifico Museo de Historia Natural, 12, 143 - 156." type="journal article" year="2008">Salazar 2008</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>