New Colombian Goja Navás (Psocodea: ‘ Psocoptera’: Epipsocidae) with peculiar genitalia, and the first Goja with brachypterous male, from Oaxaca, Mexico Author Carrejo, Nancy Departamento de Biología, Facultad de Ciencias Naturales y Exactas, Universidad del Valle, Santiago de Cali, COLOMBIA. Instituto de Estudios Ambientales, Universidad de la Sierra Juárez, Ixtlán de Juárez, Oaxaca, MÉXICO. & nancy. carrejo @ correounivalle. edu. co; htps: // orcid. org / 0000 - 0002 - 2395 - 6887 nancy.carrejo@correounivalle.edu.co Author Obando, Ranulfo González Departamento de Biología, Facultad de Ciencias Naturales y Exactas, Universidad del Valle, Santiago de Cali, COLOMBIA. Instituto de Estudios Ambientales, Universidad de la Sierra Juárez, Ixtlán de Juárez, Oaxaca, MÉXICO. & ranulfo. gonzález @ correounivalle. edu. co; htps: // orcid. org / 0000 - 0003 - 4047 - 8677 ranulfo.gonzález@correounivalle.edu.co Author Casasola-González, José Arturo 0000-0002-4765-6707 casasola @ unsij. edu. mx; https: // orcid. org / 0000 - 0002 - 4765 - 6707 casasola@unsij.edu.mx Author García Aldrete, Alfonso N. Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma de México, CdMx, MÉXICO. & anga @ ib. unam. mx; htps: // orcid. org / 0000 - 0001 - 7214 - 7966 anga@ib.unam.mx text Zootaxa 2021 2021-09-23 5040 4 451 481 journal article 10.11646/zootaxa.5040.4.1 1175-5326 5531583 FFFA7323-04AC-4B70-9DBE-6EAD909A9D2A Goja risaraldensis n. sp. Male ( Figs 19–24 ) Diagnosis. Similar to P. andina n. sp. , from which it differs in the wing venation, in having larger external parameres, in the head pattern, and in the endophallic sclerites (Compare Figs 1–2 with 19–20, 3 with 21, and 6 with 24 in this paper). Apex of the aedeagal arch robust, with the posterior border concave and a pair of the left endophallic sclerites more widened distally than in P. andina ( Fig. 24 ). Color (in 80% ethanol). Body brown with light spots. Head pattern ( Fig. 21 ); forehead brown, with broad ochre band between the compound eyes, enclosing the ocellar group. Compound eyes dark brown; ocelli hyaline, with ochre centripetal crescents. Anteclypeus brown, with ochre striae converging mesally as illustrated; labrum dark brown; genae cream to light brown, with ochre anterior areas; pre- and postgenitalia cream. Base of maxillae cream, with brown spots, maxillary palps brown, Mx4 darker.Antennae light brown. Thorax light brown, pronotum cream, tergal lobes of meso- and metathorax brown; pleura cream, with ochre spots. Legs light brown, coxae with basal ochre spots. Wings hyaline, veins light brown. Forewings ( Fig. 19 ) with abundant brown setae. Abdomen cream, with ochre brown subcuticular spots. Clunium, epiproct and paraprocts light brown, with brown spots. Hypandrium light brown. Phallosome proximally creamy, distally brown; aedeagal arch and endophallic sclerites dark brown, external parameres brown. FIGURES 13‒18. Goja meremberg n. sp. Male. 13 . Forewing. 14 . Hindwing. 15 . Front view of head. 16 . Left paraproct and epiproct. 17 . Hypandrium. 18 . Phallosome. Scales in mm. FIGURES 19‒24. Goja risaraldensis n. sp. Male. 19 . Forewing. 20 . Hindwing. 21 . Front view of head. 22 . Left paraproct and epiproct. 23 . Hypandrium. 24 . Phallosome. Scales in mm. Morphology. Head: H/MxW: 1.48; compound eyes small, H/d: 3.75; IO/MxW: 0.61. Outer cusp of lacinial tips broad, with seven denticles. Mx4/Mx2: 1.15. Labrum elongate, lower margin with a pair of projections with rounded apex, with five distal sensillae, one central placoid, flanked by a pair of trichoid-placoid. Forewings ( Fig. 19 ): FW/W: 2.81; pterostigma elongate, wider in the middle: lp/wp: 6.04; Rs with seven branches; areola postica elongate: al/ah: 4.32; M with six branches, M5 and M6 forked. Hindwings ( Fig. 20 ): HW/w: 3.12. Rs and M with four branches. Legs: Fore- tarsi without ctenidiobothria, mid- and hind- tarsi with ctenidiobothria. Hypandrium ( Fig. 23 ) broad, setose, posterior border slightly concave in the middle. Phallosome ( Fig. 24 ): deeply obtusely concave anteriorly, side struts slender, straight; aedeagal arch robust, apically with two short projections, directed outward; external parameres well developed, inner border almost straight, outer border rounded, right one distally acuminate, left one blunt distally. Three pairs of asymmetric endophallic sclerites, as illustrated ( Fig. 24 ). Clunium projecting posteriorly over the area of the epiproct. Paraprocts large, broadly elliptic ( Fig. 22 ), setae on posterior border as illustrated with setae and macrosetae; large elliptic sensory fields with 32-33 trichobothria in basal rosettes. Epiproct convex anteriorly, sides converging to almost straight posterior border; four macrosetae mesally, other setae as illustrated ( Fig. 22 ) broad. Measurements. FW: 4500, HW: 3275, F: 1080, T: 1980, t1: 995, t2: 280, ctt1: 39, f1: 1020, f2: 960, f3: 780, Mx4: 390, IO: 295, D: 252, d: 315, IO/d: 1.55, PO: 0.85. Specimen studied . Holotype male. COLOMBIA . Risaralda , Santuario , Planes de San Rafael , 5º07’32.4”N : 76º00’32.4”W , 2099 m . 18.VII.2015 , on tree trunk. R . González, N . Calderón and O. Saenz. Etymology. The specific epithet refers to the department of Risaralda , Colombia , where the holotype was found.