New Colombian Goja Navás (Psocodea: ‘ Psocoptera’: Epipsocidae) with peculiar genitalia, and the first Goja with brachypterous male, from Oaxaca, Mexico
Author
Carrejo, Nancy
Departamento de Biología, Facultad de Ciencias Naturales y Exactas, Universidad del Valle, Santiago de Cali, COLOMBIA. Instituto de Estudios Ambientales, Universidad de la Sierra Juárez, Ixtlán de Juárez, Oaxaca, MÉXICO. & nancy. carrejo @ correounivalle. edu. co; htps: // orcid. org / 0000 - 0002 - 2395 - 6887
nancy.carrejo@correounivalle.edu.co
Author
Obando, Ranulfo González
Departamento de Biología, Facultad de Ciencias Naturales y Exactas, Universidad del Valle, Santiago de Cali, COLOMBIA. Instituto de Estudios Ambientales, Universidad de la Sierra Juárez, Ixtlán de Juárez, Oaxaca, MÉXICO. & ranulfo. gonzález @ correounivalle. edu. co; htps: // orcid. org / 0000 - 0003 - 4047 - 8677
ranulfo.gonzález@correounivalle.edu.co
Author
Casasola-González, José Arturo
0000-0002-4765-6707
casasola @ unsij. edu. mx; https: // orcid. org / 0000 - 0002 - 4765 - 6707
casasola@unsij.edu.mx
Author
García Aldrete, Alfonso N.
Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma de México, CdMx, MÉXICO. & anga @ ib. unam. mx; htps: // orcid. org / 0000 - 0001 - 7214 - 7966
anga@ib.unam.mx
text
Zootaxa
2021
2021-09-23
5040
4
451
481
journal article
10.11646/zootaxa.5040.4.1
1175-5326
5531583
FFFA7323-04AC-4B70-9DBE-6EAD909A9D2A
Goja risaraldensis
n. sp.
Male
(
Figs 19–24
)
Diagnosis.
Similar to
P.
andina
n. sp.
, from which it differs in the wing venation, in having larger external parameres, in the head pattern, and in the endophallic sclerites (Compare
Figs 1–2
with 19–20, 3 with 21, and 6 with
24 in
this paper). Apex of the aedeagal arch robust, with the posterior border concave and a pair of the left endophallic sclerites more widened distally than in
P.
andina
(
Fig. 24
).
Color
(in 80% ethanol). Body brown with light spots. Head pattern (
Fig. 21
); forehead brown, with broad ochre band between the compound eyes, enclosing the ocellar group. Compound eyes dark brown; ocelli hyaline, with ochre centripetal crescents. Anteclypeus brown, with ochre striae converging mesally as illustrated; labrum dark brown; genae cream to light brown, with ochre anterior areas; pre- and postgenitalia cream. Base of maxillae cream, with brown spots, maxillary palps brown, Mx4 darker.Antennae light brown. Thorax light brown, pronotum cream, tergal lobes of meso- and metathorax brown; pleura cream, with ochre spots. Legs light brown, coxae with basal ochre spots. Wings hyaline, veins light brown. Forewings (
Fig. 19
) with abundant brown setae. Abdomen cream, with ochre brown subcuticular spots. Clunium, epiproct and paraprocts light brown, with brown spots. Hypandrium light brown. Phallosome proximally creamy, distally brown; aedeagal arch and endophallic sclerites dark brown, external parameres brown.
FIGURES 13‒18.
Goja meremberg
n. sp.
Male.
13
. Forewing.
14
. Hindwing.
15
. Front view of head.
16
. Left paraproct and epiproct.
17
. Hypandrium.
18
. Phallosome. Scales in mm.
FIGURES 19‒24.
Goja risaraldensis
n. sp.
Male.
19
. Forewing.
20
. Hindwing.
21
. Front view of head.
22
. Left paraproct and epiproct.
23
. Hypandrium.
24
. Phallosome. Scales in mm.
Morphology.
Head: H/MxW: 1.48; compound eyes small, H/d: 3.75; IO/MxW: 0.61. Outer cusp of lacinial tips broad, with seven denticles. Mx4/Mx2: 1.15. Labrum elongate, lower margin with a pair of projections with rounded apex, with five distal sensillae, one central placoid, flanked by a pair of trichoid-placoid. Forewings (
Fig. 19
): FW/W: 2.81; pterostigma elongate, wider in the middle: lp/wp: 6.04; Rs with seven branches; areola postica elongate: al/ah: 4.32; M with six branches, M5 and M6 forked. Hindwings (
Fig. 20
): HW/w: 3.12. Rs and M with four branches. Legs: Fore- tarsi without ctenidiobothria, mid- and hind- tarsi with ctenidiobothria. Hypandrium (
Fig. 23
) broad, setose, posterior border slightly concave in the middle. Phallosome (
Fig. 24
): deeply obtusely concave anteriorly, side struts slender, straight; aedeagal arch robust, apically with two short projections, directed outward; external parameres well developed, inner border almost straight, outer border rounded, right one distally acuminate, left one blunt distally. Three pairs of asymmetric endophallic sclerites, as illustrated (
Fig. 24
). Clunium projecting posteriorly over the area of the epiproct. Paraprocts large, broadly elliptic (
Fig. 22
), setae on posterior border as illustrated with setae and macrosetae; large elliptic sensory fields with 32-33 trichobothria in basal rosettes. Epiproct convex anteriorly, sides converging to almost straight posterior border; four macrosetae mesally, other setae as illustrated (
Fig. 22
) broad.
Measurements.
FW: 4500, HW: 3275, F: 1080, T: 1980, t1: 995, t2: 280, ctt1: 39, f1: 1020, f2: 960, f3: 780, Mx4: 390, IO: 295, D: 252, d: 315, IO/d: 1.55, PO: 0.85.
Specimen studied
.
Holotype
male.
COLOMBIA
.
Risaralda
,
Santuario
,
Planes de San Rafael
,
5º07’32.4”N
:
76º00’32.4”W
,
2099 m
.
18.VII.2015
, on tree trunk.
R
.
González, N
. Calderón and
O. Saenz.
Etymology.
The specific epithet refers to the department of
Risaralda
,
Colombia
, where the
holotype
was found.