Three new species of Rhipidocladum (Poaceae: Bambusoideae: Arthrostylidiinae) from South America
Author
Tyrrell, Christopher D.
Iowa State University, Dept. of Ecology, Evolution, and Organismal Biology, 251 Bessey Hall, Ames, IA 50011 USA. & University of New Brunswick, Biology Dept., P. O. Box 4400, Fredericton, NB E 3 B 5 A 3 Canada. E-mail: christopher. d. tyrrell @ unb. ca
Author
Clark, Lynn G.
Iowa State University, Dept. of Ecology, Evolution, and Organismal Biology, 251 Bessey Hall, Ames, IA 50011 USA.
text
Phytotaxa
2013
2013-05-08
98
2
55
64
http://dx.doi.org/10.11646/phytotaxa.98.2.3
journal article
10.11646/phytotaxa.98.2.3
1179-3163
5072461
Rhipidocladum arenicolum
C.D. Tyrrell & L.G. Clark
,
sp. nov.
,
Fig 2
.
Type:—
PERU
.
San Martín
.
Rioja
:
Low forest
over white sands, ca.
1 km
above Aquas Verdes
along trail to
San Pablo
,
5°41'14"S
77°37'58"W
,
1200 m
,
7 July 2002
(fl),
J
.
L
. Luteyn,
I
. Sánchez-Vega, &
M
. Zapata Cruz 15520
(
holotype
:
USM
!; isotypes:
ISC
!,
MO
!,
NY
!,
US
!)
.
Diagnosis:—
Rhipidocladum arenicolum
differs from
R. harmonicum
in its smaller culm diameter (
5–8 mm
), longer branches (
57–65 cm
), longer (
16–23 cm
) synflorescences and larger florets (lemmas
18.5–19 mm
).
Description:—
Culm height unknown; internodes ca.
26 cm
long,
5–8 mm
diameter, hollow. Culm leaves unknown. Branch complements with 18–25 branchlets, branchlets
57–65 cm
long, occasionally rebranching from the base. Foliage leaves 4–6 per flowering branchlet; sheaths
32–54 mm
long, abaxially glabrous, margins glabrous; fimbriae
1–3 mm
long, stramineous to brown, mostly deciduous; pseudopetioles
4–5 mm
long, abaxially glabrous, adaxially puberulent, dark brown; inner ligules ca.
0.5 mm
, truncate; blades
7–10 cm
long,
13–17 mm
wide, L:W ratio 5–6, broadly lanceolate, abaxially and adaxially glabrous, base rounded, apex acuminate, margins smooth. Synflorescences
16–23 cm
long, spicate, bearing 6–7(–8) spikelets spaced
20–40 mm
apart; rachis at least apically geniculate. Spikelets 4.4–5.0 cm long, comprising 2–3 glumes and 6– 7 fertile florets; rachilla internodes (2–)
4 mm
long. Lowest glume ca.
4 mm
long, 7-nerved, ovate-lanceolate, apex obtuse, abaxially glabrous, mucronate; upper glumes ovate-lanceolate, apex obtuse, abaxially glabrous, adaxially puberulent: second glume ca.
9 mm
long, 7-nerved, mucronate; third glume ca.
11 mm
long, 9- nerved, mucronate. Lemma 18.5–19.0 mm long, 13-nerved, ovate-lanceolate, apex obtuse, abaxially glabrous, mucronate. Palea ca.
13 mm
long, abaxially glabrous, adaxially scabrous, stramineous. Lodicules 3, the anterior pair ca.
6 mm
long, posterior one ca.
4 mm
long, lanceolate. Anthers
7–8 mm
long. Ovary ca.
1.7 mm
long, white. Caryopsis ca.
6.5 mm
long,
2 mm
wide, fusiform, subterete, glabrous, style base persistent, dark brown.
Distribution and Habitat:—
Known only from the
type
locality, growing in (and possibly endemic to) a white sand forest at
1200 m
elevation in northern
Peru
.
Etymology:—
Named for the habitats with white sandy soils where it occurs:
aren-
= sand,
cola
= dwelling.