Complex diversity in a mainly tropical group of ant parasitoids: Revision of the Orasema stramineipes species group (Hymenoptera: Chalcidoidea: Eucharitidae) Author Burks, Roger A. Author Heraty, John M. Author Dominguez, Chrysalyn Author Mottern, Jason L. text Zootaxa 2018 2018-03-26 4401 1 1 107 journal article 30421 10.11646/zootaxa.4401.1.1 d89af63d-c093-4908-bd61-2b26133bb7d6 1175-5326 1207463 CE7364CD-1A9E-45F2-ABA1-FB574F03BCBE Orasema arimbome Dominguez, Burks & Heraty n. sp. urn:lsid:zoobank.org:act:A3AA1AB0-9854-4E68-BCEE-16A374F318C0 ( Figs 11–18 ) Diagnosis. F2 longer than F3; funiculars dark brown. Mesoscutal midlobe and mesoscutellar disc areolate; mesoscutal midlobe with a forked or paired reddish area anteriorly. Axilla with distinct longitudinal and oblique carinae anteriorly (dorsally). Axillula areolate but with some longitudinal carinae. Stigmal vein narrow, at nearly a right angle with wing margin. Propodeum medially without reticulate sculpture. This species is similar to O. stramineipes and O. yaax in having a usually large body size, but more strongly resembles O. pirca and others with a completely areolate axillula. It is especially similar to O. pirca , from which it differs in having some longitudinal axillular carinae amidst its areolate sculpture ( Fig. 15 ) instead of having a regularly areolate axillula, and in having a differently shaped stigmal vein ( Fig. 16 versus Fig. 160 ). These differences are slight, but are upheld by distinctive molecular differences. Description. Female. Length 4.0– 5.1 mm . Color: Head and mesosoma green with reddish to yellowish luster on the mesoscutum medially, axilla laterally, and at frenal groove. Pedicel pale brown; anellus pale brown; flagellum brown. Coxae brown to greenish; fore femur mostly brown, tips pale; mid and hind femora yellow. Fore wing slightly infuscate ( Fig. 11 ); venation dark brown. Petiole green; gaster brown to green. Head: Head width:head height 1.1; face reticulate; longitudinal groove between eye and torulus present; eyes bare, inter-ocular distance:eye height 1.5–1.6; malar space:eye height 0.6–0.8; supraclypeal area longer than broad, smooth; clypeus smooth. Flagellum length:head height 11.6; F2 3.0–3.3× as long as broad, 1.3× as long as F3; following funiculars subequal in width, progressively shorter. Mesosoma: Mesosoma length:height 1.4–1.5. Mesoscutal midlobe coarsely areolate, with minute setae; lateral lobe areolate to nearly smooth. Axilla areolate with some longitudinal and oblique carinae anteriorly (dorsally); mesoscutellar disc areolate, with broader sculpture posteriorly; frenal line irregularly foveate; frenum areolatereticulate; axillular groove anteriorly absent, broad and foveate posteriorly; axillula areolate-reticulate with longitudinal carinae. Propodeum areolate, with irregular or no median carina; propodeal disc areolate; callus areolate-reticulate, with 3–5 setae. Mesepisternum areolate-reticulate laterally, smooth ventrally, broadly rounded anterior to mid coxa. Upper mesepimeron smooth; lower mesepimeron areolate-reticulate; transepimeral sulcus distinct. Metepisternum laterally nearly smooth dorsally, areolate-reticulate ventrally. Hind coxa 2.1–2.3× as long as broad; hind femur 6.4–7.8× as long as broad. Fore wing 2.6–2.7× as long as broad, 2.9–3.0× as long as mesothorax; stigma with 5 sensilla, 2 basal and 3 apical. Hind wing costal cell with a broad bare area. FIGURES 11–16. Orasema arimbome . ♀: 11. Habitus (holotype). 12. Head, cly = clypeus, scly = supraclypeal area. 13. Antenna (holotype), with F2 and F6–F7 inset, F1 = anellus, F2–F8 = funiculars, ped = pedicel, scp = scape. 14. Mesosoma (holotype), dorsal view, ax = axilla, llm = mesoscutal lateral lobe, mlm = mesoscutal midlobe. 15. Oblique view of mesoscutellum, axl = axillula, frn = frenum, scd = mesoscutellar disc. 16. Stigma and postmarginal vein, pmv = postmarginal vein, stv = stigma vein, cc = costal cell. Metasoma: Petiole 2.7–2.8× as long as broad, 1.0–1.2× as long as hind coxa, areolate but sometimes nearly smooth anteriorly, anterior carina moderate. Antecostal sulcus crossed by carinae; acrosternite posteriorly angulate. Male. Length 3.5–3.7 mm . Head width:head height 1.2; scape pale brown; flagellum length:head height 2.0– 2.1, F2 2.2–2.6× as long as broad. Fore femur mostly brown, tips pale; fore tibia yellow; mid femur mostly dark brown basally, becoming yellow apically; mid tibia yellow; hind femur mostly brown, tips pale. hind tibia yellow. Petiole 7.0–7.6× as long as broad, petiole length:hind coxa length 2.2–2.3. Distribution. Costa Rica , Guatemala , Honduras , Panama . Material examined. Holotype : COSTA RICA : Cartago : Monumento Nacional Guayabo , 1100m , 9°58'21"N , 83°41'34"W , xi.1994 , G. Fonseca [ 1 ♀ , INBIOCRI02093415], deposited in MUCR . Paratypes : COSTA RICA : Cartago : Campo Escuela Nacional Iztaru , 1800m , 9°53'20"N , 83°58'20"W , 28.ix.1991 , J .A. Ugalde [2 ♀, INBIO: INBIOCRI00876618, INBIOCRI00876636]. Dulce Nombre, Vivero Linda Vista, 1300m , vi– viii.1993 , Hanson [3 ♀, MUCR: UCRCENT00479323, UCRCENT00479325, UCRCENT00479330]. Dulce Nombre, Vivero Linda Vista, 1300m , viii–x.1993 , P. Hanson [3 ♀, MUCR: UCRCENT00479326–28]. Dulce Nombre, Vivero Linda Vista, 300m , viii–ix.1999 , P. Hanson [1 ♀, MUCR: UCRCENT00479320]. Monumento Nacional Guayabo, 1100m , 9°58'21"N, 83°41'34"W, xi.1994 , G. Fonseca [6 ♀, INBIO: INBIOCRI02093376, INBIOCRI02093401, INBIOCRI02093444, INBIOCRI02093453, INBIOCRI02093478, INBIOCRI02093515]. Monumento Nacional Guayabo, A.C.A.C. Amistad, 1100m , 9°58'21"N, 83°41'34"W, vi.1994 , G. Fonseca [7 ♀, INBIO: INBIOCRI01881684, INBIOCRI01881711, INBIOCRI01881770, INBIOCRI01881771, INBIOCRI01881900, INBIOCRI01882037, INBIOCRI01882053]. Monumento Nacional Guayabo, A.C.A.C. Amistad, 1100m , 9°58'21"N, 83°41'34"W, vii.1994 , G. Fonseca [3 ♀, INBIO: INBIOCRI01888930, INBIOCRI01889002, INBIOCRI01889162]. Monumento Nacional Guayabo, Turrialba, 1100m , 9°58'14"N, 83°41'24"W, ix.1994 , G. Fonseca [ 21 ♀ , INBIO : INBIOCRI02039320 , INBIOCRI02039326, INBIOCRI 0 2039330, INBIOCRI02039332, INBIOCRI02039348–49, INBIOCRI02039379–80, INBIOCRI02039492, INBI OCRI02039515 , INBIOCRI02039550, INBIOCRI02039557–58, INBIOCRI02039560–61, INBIOCRI02039591, INBIOCRI02039608, INBIOCRI02039611, INBIOCRI02039625, INBIOCRI02039645, INBIOCRI02039757]. La Unión , Café , Finca André, Tres Ríos de Cartago , 9°54'00"N , 83°59'23"W , 2.vii.1973 , D.J. Calvert & I. Jolis [ 2 ♀ , USNM : UCRCENT00237538–39 ]. Guanacaste : Est. Cacao , Lado SO Vol. Cacao, P.N.G. , 1000–1400m , 10°55'42"N , 85°28'06"W , v–vi.1991 , Tp malaise [1 ♀, INBIO: INBIOCRI00634862]. Est. Cacao, Lado SO Vol. Cacao, P.N.G., 1000–1400m , 10°55'42"N, 85°28'06"W, vii.1991 , C. Chaves [1 ♀, INBIO: INBIOCRI00323190]. Est. Cacao, Lado suroeste del Volcan Cacao, 1000–1400m , 10°55'42"N, 85°28'06"W, ii–iii.1989 , PNG Inventario de Biod. [ 1 ♀ , INBIO : INBIOCRI00290512 ]. Est. Cacao, Lado suroeste del Volcan Cacao , 1000–1400m , 10°55'42"N , 85°28'06"W , vi.1990 , I curso Parataxon. [10 ♀, INBIO: INBIOCRI00224643, INBIOCRI00224745, INBIOCRI00224747–48, INBIOCRI00224752–53, INBIOCRI00224755, INBIOCRI00224761, INBIOCRI00224763, INBIOCRI00252641]. Est. Cacao, Send. a la Cima. 2 km SW del Cerro Cacao, 1500m , 10°55'59"N, 85°28'03"W, iii.1996 , C. Moraga, Sombrereta [1 ♀, INBIO: INBIOCRI02473632]. Est. Cacao, SW side Volcan Cacao, 1000–1400m , 10°55'42"N, 85°28'06"W, iii.1990 , GNP Biod. Sur. [5 ♀, INBIO: INBIOCRI00165685, INBIOCRI00165693, INBIOCRI00165714, INBIOCRI00165716–17]. Est. Cacao, SW side Volcan Cacao, 1000–1400m , 10°55'42"N, 85°28'06"W, xi–xii.1989 , URCG R. Blanco & C. Chaves [ 16 ♀ , INBIO : INBIOCRI00144443 , INBIOCRI00148517, INBIOCRI00148561–65, INBIO : INBIOCRI00148568 , INBIOCRI00148570–71, INBIOCRI00148573, INBIOCRI00148599, INBIOCRI00148603, INBIOCRI00148637, INBIOCRI00148639, INBIOCRI00148842]. Est. Pitilla , 9 km S Santa Cecilia , P.N. Guanacaste , 700m , 10°59'28"N , 85°25'39"W , 31.iii–15.iv.1992 , C. Moraga [1 ♀, INBIO: INBIOCRI00416160]. Estac. Mengo, SW Volcan Cacao, 10°55'36"N , 85°28'06"W , ix–x.1989 [1 ♀, UCRC: UCRCENT00413248]. Estac. Mengo, SW Volcan Cacao, 1100m , 10°55'36"N , 85°28'06"W , ix–x.1989 [2 ♀, UCRC: UCRCENT00479156–57]. Guanacaste N.P., Pitilla Biological Station , 10°59'31"N , 85°28'20"W , 1.ii.1995 , L. Masner, screen sweep [8 ♀, UCRC: UCRCENT00312149–55, UCRCENT00312157]. Macizo Miravalles, Estación Cabro Muco, 1100m , 10°43'00"N, 85°08'34"W, 15.iii–2.iv.2003 , J. Azofeifa , sweep [ 1 ♀ , INBIO : INB03710916 ]. P.N. Santa Rosa , Rio Calera , Est. Santa Rosa , 0–100m , 10°48'11"N , 85°40'27"W , 21.vii.2000 , D. Briceno [ 1 ♀ , UCRC : UCRCENT00479158 ] . Sector Las Pailas , Sendero Volcan, P.N . Rincon de la Vieja, 800m , 10°46'31"N , 85°20'59"W , 28.xi.1993 5.i.1994 , E. Araya, malaise trap [1 ♀, INBIO: INBIOCRI01981909]. Vn Cacao (ACG), 1659m , 13.ii.1995 , J. Whitfield [1 ♀, BMNH: UCRCENT00309943]. Volca Cacao, Arenales, W side, 900m , 10°56'03"N , 85°27'22"W , 1988–1989 [1 ♀, UCRC: UCRCENT00413504]. Heredia : 10 km N San Jose , 10°02'28"N , 84°07'52"W , vii.1971 , J . Maldonado C. [ 2 ♀ , USNM : UCRCENT00416817–18 ] . San Jose , 1100–1200m , 9°56'00"N , 84°05'00"W , viii.1980 , N.L.H. Krauss [ 3 ♀ , AMNH : UCRCENT00424174–76 ] . Santo Domingo , 1200m , 9°58'48"N , 84°05'24"W , 26.iv.2016 , P. Hanson, YPT [3 ♀, UCRC: UCRCENT00479159 (D4700), UCRCENT00479160 (D4702), UCRCENT00479161 (D4701)]. Santo Domingo , 1200m , i–ii.2013 , P. Hanson [ 1 ♀ , MUCR : UCRCENT00479329 ] . Santo Domingo , INBio Cafetal , 1100m , 9°58'48"N , 84°05'24"W , 25–28.vi.1997 , S. & J. Peck , FIT [ 1 ♀ , CNC : UCRCENT00425818 (D4644)]. Limón : Esc Agricultura , 10°12'56"N , 83°35'30"W , 10.viii.1972 , J. Maldonado C. [1 ♀, USNM: UCRCENT00416819]. Puntarenas : Est. Altamira , Buenos Aires , 1430m , 9°01'53"N , 83°00'32"W , 15.ix–14.x.1993 , R. Delgado [3 ♀, INBIO: INBIOCRI01620784, INBIOCRI01620807, INBIOCRI01620860]. San Jose : San Antonio de Escazu, Eberhard property, 1300m , 9°53'49"N , 84°08'15"W , 4–5.iii.1996 , L. Masner , creekbed, Yellow Pan Trap [ 1 ♀ , CNC : UCRCENT00425961 ] . San Gerardo , 11.ix.1986 , L. Masner [ 1 ♀ , CNC : UCRCENT00425960 ]. GUATEMALA : Huehuetenango : Todos Santos , 2500m , 15°30'14"N , 91°36'04"W , 30.viii.1987 , M. Sharkey [ 1♂ , CNC : UCRCENT00425820 ]. HONDURAS : Yoro : Atlantida , 100m , 15°40'11"N , 87°08'32"W , 12.ix.1972 , J. Helava [ 1 ♀ , CNC : UCRCENT00247569 ] . PANAMA : Chiriqui : 5 km . NE Santa Clara , 1500m , 8°51'52"N , 82°43'44"W , 6.vii.1996 , Gillogly and Schaffner [ 3 ♀ , TAMU : UCRCENT00426525– 27 ]. Reserva La Fortuna Estacion Biologica , 1150m , 8°44'03"N , 82°14'28"W , 11–13.vii.1996 , A.R. Gillogly [ 4 ♀ , TAMU : UCRCENT00426529–32 ]. Lagunas del Volcan , 5 km SW Volcan , 1286m , 8°45'52"N , 82°40'33"W , 30.vii.1999 , J.B. Woolley [ 1 ♀ , TAMU : UCRCENT00426536 ]. Coclé : El Valle , 8°34'48"N , 80°10'12"W , xi.1946 , N.L.H. Krauss [ 2 ♀ , USNM : UCRCENT00248644–45 ]. Ngäbe-Buglé Comarca : 3 km W Fortuna Hwy , Oleoducto Rd , 1170m , 8°47'07"N , 82°12'05"W , 6–9.i.2001 , M. Yoder & J.B. Woolley [1 ♀, TAMU: UCRCENT00400702]. Etymology. The Mangue word for “red,” referring to the large reddish area on the mesoscutal midlobe (see Fig. 14 and color section in description above).