Complex diversity in a mainly tropical group of ant parasitoids: Revision of the Orasema stramineipes species group (Hymenoptera: Chalcidoidea: Eucharitidae)
Author
Burks, Roger A.
Author
Heraty, John M.
Author
Dominguez, Chrysalyn
Author
Mottern, Jason L.
text
Zootaxa
2018
2018-03-26
4401
1
1
107
journal article
30421
10.11646/zootaxa.4401.1.1
d89af63d-c093-4908-bd61-2b26133bb7d6
1175-5326
1207463
CE7364CD-1A9E-45F2-ABA1-FB574F03BCBE
Orasema arimbome
Dominguez, Burks & Heraty
n. sp.
urn:lsid:zoobank.org:act:A3AA1AB0-9854-4E68-BCEE-16A374F318C0 (
Figs 11–18
)
Diagnosis.
F2 longer than F3; funiculars dark brown. Mesoscutal midlobe and mesoscutellar disc areolate; mesoscutal midlobe with a forked or paired reddish area anteriorly. Axilla with distinct longitudinal and oblique carinae anteriorly (dorsally). Axillula areolate but with some longitudinal carinae. Stigmal vein narrow, at nearly a right angle with wing margin. Propodeum medially without reticulate sculpture.
This species is similar to
O. stramineipes
and
O. yaax
in having a usually large body size, but more strongly resembles
O. pirca
and others with a completely areolate axillula. It is especially similar to
O. pirca
, from which it differs in having some longitudinal axillular carinae amidst its areolate sculpture (
Fig. 15
) instead of having a regularly areolate axillula, and in having a differently shaped stigmal vein (
Fig. 16
versus
Fig. 160
). These differences are slight, but are upheld by distinctive molecular differences.
Description. Female.
Length 4.0–
5.1 mm
.
Color:
Head and mesosoma green with reddish to yellowish luster on the mesoscutum medially, axilla laterally, and at frenal groove. Pedicel pale brown; anellus pale brown; flagellum brown. Coxae brown to greenish; fore femur mostly brown, tips pale; mid and hind femora yellow. Fore wing slightly infuscate (
Fig. 11
); venation dark brown. Petiole green; gaster brown to green.
Head:
Head width:head height 1.1; face reticulate; longitudinal groove between eye and torulus present; eyes bare, inter-ocular distance:eye height 1.5–1.6; malar space:eye height 0.6–0.8; supraclypeal area longer than broad, smooth; clypeus smooth. Flagellum length:head height 11.6; F2 3.0–3.3× as long as broad, 1.3× as long as F3; following funiculars subequal in width, progressively shorter.
Mesosoma:
Mesosoma length:height 1.4–1.5. Mesoscutal midlobe coarsely areolate, with minute setae; lateral lobe areolate to nearly smooth. Axilla areolate with some longitudinal and oblique carinae anteriorly (dorsally); mesoscutellar disc areolate, with broader sculpture posteriorly; frenal line irregularly foveate; frenum areolatereticulate; axillular groove anteriorly absent, broad and foveate posteriorly; axillula areolate-reticulate with longitudinal carinae. Propodeum areolate, with irregular or no median carina; propodeal disc areolate; callus areolate-reticulate, with 3–5 setae. Mesepisternum areolate-reticulate laterally, smooth ventrally, broadly rounded anterior to mid coxa. Upper mesepimeron smooth; lower mesepimeron areolate-reticulate; transepimeral sulcus distinct. Metepisternum laterally nearly smooth dorsally, areolate-reticulate ventrally. Hind coxa 2.1–2.3× as long as broad; hind femur 6.4–7.8× as long as broad. Fore wing 2.6–2.7× as long as broad, 2.9–3.0× as long as mesothorax; stigma with 5 sensilla, 2 basal and 3 apical. Hind wing costal cell with a broad bare area.
FIGURES 11–16.
Orasema arimbome
. ♀:
11.
Habitus (holotype).
12.
Head, cly = clypeus, scly = supraclypeal area.
13.
Antenna (holotype), with F2 and F6–F7 inset, F1 = anellus, F2–F8 = funiculars, ped = pedicel, scp = scape.
14.
Mesosoma (holotype), dorsal view, ax = axilla, llm = mesoscutal lateral lobe, mlm = mesoscutal midlobe.
15.
Oblique view of mesoscutellum, axl = axillula, frn = frenum, scd = mesoscutellar disc.
16.
Stigma and postmarginal vein, pmv = postmarginal vein, stv = stigma vein, cc = costal cell.
Metasoma:
Petiole 2.7–2.8× as long as broad, 1.0–1.2× as long as hind coxa, areolate but sometimes nearly smooth anteriorly, anterior carina moderate. Antecostal sulcus crossed by carinae; acrosternite posteriorly angulate.
Male.
Length
3.5–3.7 mm
. Head width:head height 1.2; scape pale brown; flagellum length:head height 2.0– 2.1, F2 2.2–2.6× as long as broad. Fore femur mostly brown, tips pale; fore tibia yellow; mid femur mostly dark brown basally, becoming yellow apically; mid tibia yellow; hind femur mostly brown, tips pale. hind tibia yellow. Petiole 7.0–7.6× as long as broad, petiole length:hind coxa length 2.2–2.3.
Distribution.
Costa
Rica
,
Guatemala
,
Honduras
,
Panama
.
Material
examined.
Holotype
:
COSTA RICA
:
Cartago
:
Monumento Nacional Guayabo
,
1100m
,
9°58'21"N
,
83°41'34"W
,
xi.1994
,
G. Fonseca
[
1 ♀
, INBIOCRI02093415], deposited in
MUCR
.
Paratypes
:
COSTA RICA
:
Cartago
:
Campo Escuela Nacional Iztaru
,
1800m
,
9°53'20"N
,
83°58'20"W
,
28.ix.1991
, J
.A. Ugalde [2 ♀, INBIO: INBIOCRI00876618, INBIOCRI00876636]. Dulce Nombre, Vivero Linda Vista,
1300m
,
vi– viii.1993
, Hanson [3 ♀, MUCR: UCRCENT00479323, UCRCENT00479325, UCRCENT00479330]. Dulce Nombre, Vivero Linda Vista,
1300m
,
viii–x.1993
, P. Hanson [3 ♀, MUCR: UCRCENT00479326–28]. Dulce Nombre, Vivero Linda Vista,
300m
,
viii–ix.1999
, P. Hanson [1 ♀, MUCR: UCRCENT00479320]. Monumento Nacional Guayabo,
1100m
, 9°58'21"N, 83°41'34"W,
xi.1994
, G. Fonseca [6 ♀, INBIO: INBIOCRI02093376, INBIOCRI02093401, INBIOCRI02093444, INBIOCRI02093453, INBIOCRI02093478, INBIOCRI02093515]. Monumento Nacional Guayabo, A.C.A.C. Amistad,
1100m
, 9°58'21"N, 83°41'34"W,
vi.1994
, G. Fonseca [7 ♀, INBIO: INBIOCRI01881684, INBIOCRI01881711, INBIOCRI01881770, INBIOCRI01881771, INBIOCRI01881900, INBIOCRI01882037, INBIOCRI01882053]. Monumento Nacional Guayabo, A.C.A.C. Amistad,
1100m
, 9°58'21"N, 83°41'34"W,
vii.1994
, G. Fonseca [3 ♀, INBIO: INBIOCRI01888930, INBIOCRI01889002, INBIOCRI01889162]. Monumento Nacional Guayabo, Turrialba,
1100m
, 9°58'14"N, 83°41'24"W,
ix.1994
,
G. Fonseca
[
21 ♀
,
INBIO
:
INBIOCRI02039320
, INBIOCRI02039326, INBIOCRI 0 2039330, INBIOCRI02039332, INBIOCRI02039348–49, INBIOCRI02039379–80, INBIOCRI02039492,
INBI
OCRI02039515
, INBIOCRI02039550, INBIOCRI02039557–58, INBIOCRI02039560–61, INBIOCRI02039591, INBIOCRI02039608, INBIOCRI02039611, INBIOCRI02039625, INBIOCRI02039645, INBIOCRI02039757].
La Unión
,
Café
, Finca André, Tres Ríos de
Cartago
,
9°54'00"N
,
83°59'23"W
,
2.vii.1973
,
D.J. Calvert
& I.
Jolis
[
2 ♀
,
USNM
:
UCRCENT00237538–39
].
Guanacaste
:
Est. Cacao
,
Lado SO Vol. Cacao, P.N.G.
,
1000–1400m
,
10°55'42"N
,
85°28'06"W
,
v–vi.1991
, Tp malaise [1 ♀, INBIO: INBIOCRI00634862]. Est. Cacao, Lado SO Vol. Cacao, P.N.G.,
1000–1400m
, 10°55'42"N, 85°28'06"W,
vii.1991
, C. Chaves [1 ♀, INBIO: INBIOCRI00323190]. Est. Cacao, Lado suroeste del Volcan Cacao,
1000–1400m
, 10°55'42"N, 85°28'06"W,
ii–iii.1989
,
PNG
Inventario de Biod.
[
1 ♀
,
INBIO
:
INBIOCRI00290512
]. Est. Cacao, Lado suroeste del
Volcan Cacao
,
1000–1400m
,
10°55'42"N
,
85°28'06"W
,
vi.1990
, I curso Parataxon. [10 ♀, INBIO: INBIOCRI00224643, INBIOCRI00224745, INBIOCRI00224747–48, INBIOCRI00224752–53, INBIOCRI00224755, INBIOCRI00224761, INBIOCRI00224763, INBIOCRI00252641]. Est. Cacao, Send. a la Cima.
2 km
SW del Cerro Cacao,
1500m
, 10°55'59"N, 85°28'03"W,
iii.1996
, C. Moraga, Sombrereta [1 ♀, INBIO: INBIOCRI02473632]. Est. Cacao, SW side Volcan Cacao,
1000–1400m
, 10°55'42"N, 85°28'06"W,
iii.1990
, GNP Biod. Sur. [5 ♀, INBIO: INBIOCRI00165685, INBIOCRI00165693, INBIOCRI00165714, INBIOCRI00165716–17]. Est. Cacao, SW side Volcan Cacao,
1000–1400m
, 10°55'42"N, 85°28'06"W,
xi–xii.1989
, URCG R.
Blanco
&
C. Chaves
[
16 ♀
,
INBIO
:
INBIOCRI00144443
, INBIOCRI00148517, INBIOCRI00148561–65,
INBIO
:
INBIOCRI00148568
, INBIOCRI00148570–71, INBIOCRI00148573, INBIOCRI00148599, INBIOCRI00148603, INBIOCRI00148637, INBIOCRI00148639, INBIOCRI00148842].
Est. Pitilla
,
9 km
S
Santa Cecilia
, P.N.
Guanacaste
,
700m
,
10°59'28"N
,
85°25'39"W
,
31.iii–15.iv.1992
, C. Moraga [1 ♀, INBIO: INBIOCRI00416160]. Estac. Mengo, SW Volcan Cacao,
10°55'36"N
,
85°28'06"W
,
ix–x.1989
[1 ♀, UCRC: UCRCENT00413248]. Estac. Mengo, SW Volcan Cacao,
1100m
,
10°55'36"N
,
85°28'06"W
,
ix–x.1989
[2 ♀, UCRC: UCRCENT00479156–57].
Guanacaste
N.P.,
Pitilla Biological Station
,
10°59'31"N
,
85°28'20"W
,
1.ii.1995
, L. Masner, screen sweep [8 ♀, UCRC: UCRCENT00312149–55, UCRCENT00312157]. Macizo Miravalles, Estación Cabro Muco,
1100m
, 10°43'00"N, 85°08'34"W,
15.iii–2.iv.2003
,
J. Azofeifa
, sweep [
1 ♀
,
INBIO
:
INB03710916
].
P.N.
Santa Rosa
,
Rio Calera
,
Est.
Santa Rosa
,
0–100m
,
10°48'11"N
,
85°40'27"W
,
21.vii.2000
,
D. Briceno
[
1 ♀
,
UCRC
:
UCRCENT00479158
]
.
Sector Las Pailas
,
Sendero Volcan, P.N
.
Rincon
de la Vieja,
800m
,
10°46'31"N
,
85°20'59"W
,
28.xi.1993
–
5.i.1994
, E. Araya,
malaise trap
[1 ♀, INBIO: INBIOCRI01981909]. Vn Cacao (ACG),
1659m
,
13.ii.1995
, J. Whitfield [1 ♀, BMNH: UCRCENT00309943]. Volca Cacao, Arenales, W side,
900m
,
10°56'03"N
,
85°27'22"W
,
1988–1989
[1 ♀, UCRC: UCRCENT00413504].
Heredia
:
10 km
N
San Jose
,
10°02'28"N
,
84°07'52"W
,
vii.1971
, J
.
Maldonado
C. [
2 ♀
,
USNM
:
UCRCENT00416817–18
]
.
San Jose
,
1100–1200m
,
9°56'00"N
,
84°05'00"W
,
viii.1980
,
N.L.H. Krauss
[
3 ♀
,
AMNH
:
UCRCENT00424174–76
]
.
Santo Domingo
,
1200m
,
9°58'48"N
,
84°05'24"W
,
26.iv.2016
, P. Hanson, YPT [3 ♀, UCRC: UCRCENT00479159 (D4700), UCRCENT00479160 (D4702), UCRCENT00479161 (D4701)].
Santo Domingo
,
1200m
,
i–ii.2013
,
P. Hanson
[
1 ♀
,
MUCR
:
UCRCENT00479329
]
.
Santo Domingo
,
INBio
Cafetal
,
1100m
,
9°58'48"N
,
84°05'24"W
,
25–28.vi.1997
,
S. & J. Peck
, FIT [
1 ♀
,
CNC
:
UCRCENT00425818
(D4644)].
Limón
:
Esc Agricultura
,
10°12'56"N
,
83°35'30"W
,
10.viii.1972
, J. Maldonado C. [1 ♀, USNM: UCRCENT00416819].
Puntarenas
:
Est. Altamira
,
Buenos Aires
,
1430m
,
9°01'53"N
,
83°00'32"W
,
15.ix–14.x.1993
, R. Delgado [3 ♀, INBIO: INBIOCRI01620784, INBIOCRI01620807, INBIOCRI01620860].
San Jose
:
San Antonio de Escazu, Eberhard property,
1300m
,
9°53'49"N
,
84°08'15"W
,
4–5.iii.1996
,
L. Masner
, creekbed,
Yellow Pan Trap
[
1 ♀
,
CNC
:
UCRCENT00425961
]
.
San Gerardo
,
11.ix.1986
,
L. Masner
[
1 ♀
,
CNC
:
UCRCENT00425960
].
GUATEMALA
:
Huehuetenango
:
Todos Santos
,
2500m
,
15°30'14"N
,
91°36'04"W
,
30.viii.1987
,
M. Sharkey
[
1♂
,
CNC
:
UCRCENT00425820
].
HONDURAS
:
Yoro
:
Atlantida
,
100m
,
15°40'11"N
,
87°08'32"W
,
12.ix.1972
,
J. Helava
[
1 ♀
,
CNC
:
UCRCENT00247569
]
.
PANAMA
:
Chiriqui
:
5 km
.
NE Santa Clara
,
1500m
,
8°51'52"N
,
82°43'44"W
,
6.vii.1996
, Gillogly and Schaffner [
3 ♀
,
TAMU
:
UCRCENT00426525– 27
].
Reserva La Fortuna Estacion Biologica
,
1150m
,
8°44'03"N
,
82°14'28"W
,
11–13.vii.1996
,
A.R. Gillogly
[
4 ♀
,
TAMU
:
UCRCENT00426529–32
].
Lagunas
del
Volcan
,
5 km
SW Volcan
,
1286m
,
8°45'52"N
,
82°40'33"W
,
30.vii.1999
,
J.B. Woolley
[
1 ♀
,
TAMU
:
UCRCENT00426536
].
Coclé
:
El
Valle
,
8°34'48"N
,
80°10'12"W
,
xi.1946
,
N.L.H. Krauss
[
2 ♀
,
USNM
:
UCRCENT00248644–45
].
Ngäbe-Buglé Comarca
:
3 km
W Fortuna Hwy
,
Oleoducto Rd
,
1170m
,
8°47'07"N
,
82°12'05"W
,
6–9.i.2001
, M. Yoder & J.B. Woolley [1 ♀, TAMU: UCRCENT00400702].
Etymology.
The Mangue word for “red,” referring to the large reddish area on the mesoscutal midlobe (see
Fig. 14
and color section in description above).