<document ID-CLB-Dataset="24666" ID-DOI="http://dx.doi.org/10.3897/evolsyst.3.37464" ID-GBIF-Dataset="8ac0c1d8-2ebc-40bd-bcf6-7216ce5d8d66" ID-Pensoft-Pub="2535-0730-2-109" ID-Pensoft-UUID="48BB2ADCDFB750ACA7D9EC306EC80801" ID-ZooBank="87602625AF8D4A3FBAE5F35C09FB6C00" ModsDocID="2535-0730-2-109" checkinTime="1575997079287" checkinUser="pensoft" docAuthor="Monod, Lionel, Duperre, Nadine &amp; Harms, Danilo" docDate="2019" docId="6C60DECD0BEA5DB88B92297E47889DD9" docLanguage="en" docName="EvolutSyst 3(2): 109-200" docOrigin="Evolutionary Systematics 3 (2)" docSource="http://dx.doi.org/10.3897/evolsyst.3.37464" docTitle="Hormurus boholiensis" docType="treatment" docVersion="6" id="48BB2ADCDFB750ACA7D9EC306EC80801" lastPageNumber="109" masterDocId="48BB2ADCDFB750ACA7D9EC306EC80801" masterDocTitle="An annotated catalogue of the scorpion types (Arachnida, Scorpiones) held in the Zoological Museum Hamburg. Part I: Parvorder Iurida Soleglad &amp; Fet, 2003" masterLastPageNumber="200" masterPageNumber="109" pageNumber="109" updateTime="1732626668489" updateUser="ExternalLinkService">
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<mods:title id="843F689A0C2B724F9F586E8494D52495">An annotated catalogue of the scorpion types (Arachnida, Scorpiones) held in the Zoological Museum Hamburg. Part I: Parvorder Iurida Soleglad &amp; Fet, 2003</mods:title>
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<mods:namePart id="198F3B4BA6B550EA64EE550D61FC8DC2">Monod, Lionel</mods:namePart>
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<mods:namePart id="02ACE325DECDBD33FDC4B3F3191B37C0">Duperre, Nadine</mods:namePart>
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<mods:namePart id="04D2209343CF01D2F2E7DE3A20B322FF">Harms, Danilo</mods:namePart>
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<paragraph id="9C01DA84CBAB89F0C01A07DD81558D72" pageId="0" pageNumber="109">
<taxonomicName id="0E474BE836A25E3DAAF09DD3977FF502" ID-CoL="6MDBJ" LSID="6C60DECD-0BEA-5DB8-8B92-297E47889DD9" class="Arachnida" family="Hormuridae" genus="Hormurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hormurus boholiensis" order="Scorpiones" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="boholiensis">Hormurus boholiensis</taxonomicName>
<figureCitation id="42B70749D6A9DAA5D390D3ADA1FA4F55" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Hormurus boholiensis Kraepelin, 1914, female lectotype (A-B). Ischnurus caudicula L. Koch, 1867 [= Hormurus waigiensis (Gervais, 1843)], subadult female holotype (C-D): A, C dorsal aspect of habitus B, D ventral aspect of habitus. Scale bars: 10 mm." httpUri="https://binary.pensoft.net/fig/355041" pageId="0" pageNumber="109">
Fig. 28 
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</paragraph>
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<paragraph id="E3F72012A67761FC00751B831A3F41B3" pageId="0" pageNumber="109">
<taxonomicName id="FDEEA346A6AAEA0E2A0EAC7D96CBCA8C" class="Arachnida" family="Hormuridae" genus="Hormurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hormurus boholiensis" order="Scorpiones" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="boholiensis">Hormurus boholiensis</taxonomicName>
Kraepelin, 1914d: 333
</paragraph>
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<subSubSection id="31BB450B7B24D33241FE7CA423A639A2" pageId="0" pageNumber="109" type="current combination">
<paragraph id="0D4AE1E24E2461BD63CAD2D69405EF1D" pageId="0" pageNumber="109">Current combination.</paragraph>
<paragraph id="FA0CE478FAFBB9C395FE7963AC5386F6" pageId="0" pageNumber="109">
<taxonomicName id="7C647EEEAE4F70E7422049AE1F464569" class="Arachnida" family="Hormuridae" genus="Hormurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hormurus boholiensis" order="Scorpiones" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="boholiensis">
<emphasis id="C20EDB1FDC911E47CC570BF0085CCE39" italics="true" pageId="0" pageNumber="109">Hormurus boholiensis</emphasis>
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Kraepelin, 1914
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<subSubSection id="F09EDEA643C54B96D66438A43E1C914C" pageId="0" pageNumber="109" type="lectotype">
<paragraph id="CC7440413A9FDE7AB26DA0E68E7F8EAD" pageId="0" pageNumber="109">Lectotype.</paragraph>
<paragraph id="80F6E7221F3EE18E3A87942F56DE226E" pageId="0" pageNumber="109">
(
<figureCitation id="097E20638D68D1CDCBB283405E8A467D" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Hormurus boholiensis Kraepelin, 1914, female lectotype (A-B). Ischnurus caudicula L. Koch, 1867 [= Hormurus waigiensis (Gervais, 1843)], subadult female holotype (C-D): A, C dorsal aspect of habitus B, D ventral aspect of habitus. Scale bars: 10 mm." httpUri="https://binary.pensoft.net/fig/355041" pageId="0" pageNumber="109">
Fig. 28 
<normalizedToken id="402D5482F131AB95DA812002AE03783E" originalValue="A–B">A-B</normalizedToken>
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) ♀, Philippines, [Central Visayas], Bohol [
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9°50 
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N
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, 
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124°10 
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E
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], 10.1863, ZMK 3938 [Carl Gottfried Semper leg.].
</paragraph>
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<subSubSection id="F743B26231F7CAAAB333A3CE3BD4B3BA" pageId="0" pageNumber="109" type="paralectotype">
<paragraph id="1745D6090D972AD4CE586D59ADEDE9E7" pageId="0" pageNumber="109">Paralectotype.</paragraph>
<paragraph id="E38B7B1D145FFCDBC707F30DB5E9C031" pageId="0" pageNumber="109">♀, same data as lectotype.</paragraph>
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<subSubSection id="648C393A912043D2C46EB482416A491D" pageId="0" pageNumber="109" type="remarks">
<paragraph id="F342AE3F8D43E49DF1590903FD54CA9F" pageId="0" pageNumber="109">Remarks.</paragraph>
<paragraph id="371CB975F94830AF072AAE3A8663CEED" pageId="0" pageNumber="109">
<bibRefCitation id="C065FA6B62742971A9F6A9A6D72BC5F9" author="Kraepelin, K" editor="Sarasin, F" journalOrPublisher="Memoires de la Societe Neuchateloise des Sciences Naturelles" pageId="0" pageNumber="109" refId="B89" refString="Kraepelin, K, Sarasin, F, Roux, J, 1914d. Die Skorpione und Pedipalpen von Neu-Caledonien und den benachbarten Inselgruppen In: SarasinFRouxJ (Eds) Nova Caledonia, Zoologie.[Vol. 1.] C.W. Kreidals Verlag, Wiesbaden, 327-337." title="Die Skorpione und Pedipalpen von Neu-Caledonien und den benachbarten Inselgruppen In: SarasinFRouxJ (Eds) Nova Caledonia, Zoologie. [Vol. 1.] C. W. Kreidals Verlag, Wiesbaden, 327 - 337." year="1914 d">Kraepelin (1914d)</bibRefCitation>
mentioned that the two type specimens were poorly preserved: &quot;Von dieser Arte liegen mir nur zwei schlecht erhaltene ♀ des Hamburger Museum vor.&quot;. 
<taxonomicName id="D130F602A94179DDB934FF04F4009D57" lsidName="H. boholiensis" pageId="0" pageNumber="109" rank="species" species="boholiensis">
<emphasis id="1ED0F1B63CFE12D877284C1EE56F7430" italics="true" pageId="0" pageNumber="109">H. boholiensis</emphasis>
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was considered a junior synonym of 
<taxonomicName id="DA552FC7F0493D5D7E64E00DC83A9EE5" class="Arachnida" family="Hormuridae" genus="Liocheles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liocheles australasiae" order="Scorpiones" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="australasiae">
<emphasis id="B016729B150404B71360770CEA7C5408" italics="true" pageId="0" pageNumber="109">Liocheles australasiae</emphasis>
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(Fabricius, 1775) by L. E. 
<bibRefCitation id="BA94DB48F42D4AE43F28A2BEB3407EC9" author="Koch, LE" journalOrPublisher="Records of the Western Australian Museum" pageId="0" pageNumber="109" pagination="83 - 367" refId="B40" refString="Koch, LE, 1977. The taxonomy, geographic distribution and evolutionary radiation of Australo-Papuan scorpions. Records of the Western Australian Museum 5: 83 - 367" title="The taxonomy, geographic distribution and evolutionary radiation of Australo-Papuan scorpions." volume="5" year="1977">Koch (1977</bibRefCitation>
: 161). 
<bibRefCitation id="53BE214C3118B77C58AF36E2632B197C" author="Monod, L" journalOrPublisher="Revue Suisse de Zoologie" pageId="0" pageNumber="109" pagination="723 - 758" refId="B123" refString="Monod, L, 2011b. Taxonomic emendations in the genus Liocheles Sundevall, 1833 (Scorpiones, Liochelidae). Revue Suisse de Zoologie 118: 723 - 758" title="Taxonomic emendations in the genus Liocheles Sundevall, 1833 (Scorpiones, Liochelidae)." volume="118" year="2011 b">Monod (2011b)</bibRefCitation>
revalidated the species as 
<taxonomicName id="1C0A725412ADFFBF289CA362FE2090F6" class="Arachnida" family="Hormuridae" genus="Liocheles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liocheles boholiensis" order="Scorpiones" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="boholiensis">
<emphasis id="FD0DE9A326E3CF0EB74686E4FE4862F3" italics="true" pageId="0" pageNumber="109">Liocheles boholiensis</emphasis>
</taxonomicName>
(Kraepelin, 1914). 
<bibRefCitation id="938889924203B96B99D6D552BF228F87" DOI="https://doi.org/10.1111/cla.12067" author="Monod, L" journalOrPublisher="Cladistics" pageId="0" pageNumber="109" pagination="71 - 111" refId="B125" refString="Monod, L, Prendini, L, 2015. Evidence for Eurogondwana: the roles of dispersal, extinction and vicariance in the evolution and biogeography of Indo-Pacific Hormuridae (Scorpiones: Scorpionoidea). Cladistics 31: 71 - 111" title="Evidence for Eurogondwana: the roles of dispersal, extinction and vicariance in the evolution and biogeography of Indo-Pacific Hormuridae (Scorpiones: Scorpionoidea)." url="https://doi.org/10.1111/cla.12067" volume="31" year="2015">Monod and Prendini (2015)</bibRefCitation>
subsequently removed the genus 
<taxonomicName id="1D657F3C4D7970613D171850E536E060" class="Arachnida" family="Hormuridae" genus="Hormurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hormurus" order="Scorpiones" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis id="080335B1BA32AFD614898ACED794431A" italics="true" pageId="0" pageNumber="109">Hormurus</emphasis>
</taxonomicName>
from synonymy with 
<taxonomicName id="0DDD335981B190A9B8B2235BA64C55B9" class="Arachnida" family="Hormuridae" genus="Liocheles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liocheles" order="Scorpiones" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis id="5A3FA62D49BE8BB89D378303A94A10DE" italics="true" pageId="0" pageNumber="109">Liocheles</emphasis>
</taxonomicName>
and confirmed the species 
<taxonomicName id="745F0290D64C3430A8646A7023A6CB2A" class="Magnoliopsida" family="Asteraceae" genus="Oligochaeta" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Oligochaeta boholiensis" order="Asterales" pageId="0" pageNumber="109" phylum="Tracheophyta" rank="species" species="boholiensis">
<emphasis id="E434D9F6CE2AEE9DFBE6833FA55EE5BF" italics="true" pageId="0" pageNumber="109">boholiensis</emphasis>
</taxonomicName>
as belonging to 
<taxonomicName id="45503DFE97128B7394F7488C37F9C081" class="Arachnida" family="Hormuridae" genus="Hormurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hormurus" order="Scorpiones" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis id="D0DBCAD5E7D94C29309623EB18469699" italics="true" pageId="0" pageNumber="109">Hormurus</emphasis>
</taxonomicName>
. The larger female of this series was designated as lectotype by 
<bibRefCitation id="C6A096384992C6A396387CBC08772EB0" author="Monod, L" journalOrPublisher="Revue Suisse de Zoologie" pageId="0" pageNumber="109" pagination="723 - 758" refId="B123" refString="Monod, L, 2011b. Taxonomic emendations in the genus Liocheles Sundevall, 1833 (Scorpiones, Liochelidae). Revue Suisse de Zoologie 118: 723 - 758" title="Taxonomic emendations in the genus Liocheles Sundevall, 1833 (Scorpiones, Liochelidae)." volume="118" year="2011 b">Monod (2011b</bibRefCitation>
: 726)
</paragraph>
</subSubSection>
<subSubSection id="7422BED08EFC80B5F6F3C4FFCB4C793F" pageId="0" pageNumber="109" type="remarks">
<paragraph id="FF4A005E7109903F67EDE05B725D0783" pageId="0" pageNumber="109">Remarks on collector.</paragraph>
<paragraph id="0B539B882135F117A4610CE1B5E7478A" pageId="0" pageNumber="109">
Although not mentioned on the original labels, the specimens were probably collected by Carl Gottfried Semper (1832-1893), a German malacologist who obtained a Ph.D. degree at the University of 
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in 1856 and continued his studies in Kiel. An inheritance from his father allowed him to pay for a seven years trip to the Philippines and Palau (1858-1865) (
<bibRefCitation id="065CC6905C46B0683CCF57DA9E5CEE2C" author="Schuberg, A" editor="Bergh, R" journalOrPublisher="Mitteilungen aus dem Museum fuer Naturkunde in Berlin" pageId="0" pageNumber="109" refId="B153" refString="Schuberg, A, Bergh, R, 1880. Carl Semper. In: BerghR (Ed.) Reisen im Archipel der Philippinen von Dr.C. Semper, Zweiter Theil, Wissenschaftliche Resultate, Band 2, Supplemente, Malacologische Untersuchungen von Dr. Rud Bergh, Nudibranchien. CW Kreisel's Verlag. Wiesbaden, 7-21." title="Carl Semper. In: BerghR (Ed.) Reisen im Archipel der Philippinen von Dr. C. Semper, Zweiter Theil, Wissenschaftliche Resultate, Band 2, Supplemente, Malacologische Untersuchungen von Dr. Rud Bergh, Nudibranchien. CW Kreisel's Verlag. Wiesbaden, 7 - 21." year="1880">Schuberg 1880</bibRefCitation>
). After returning to Germany, he was appointed lecturer (1866) and professor (1869) at the University of 
<normalizedToken id="E1B70280909B18ECFFB04E6C7EDF1B21" originalValue="Würzburg">Wuerzburg</normalizedToken>
. He visited Bohol in 1863, which corresponds to the collecting date of the two present specimens.
</paragraph>
</subSubSection>
</treatment>
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