<document id="981FF11D76AA322F4757CF92E278C246" ID-CLB-Dataset="312558" ID-DOI="10.3897/zookeys.1255.158380" ID-GBIF-Dataset="fc490eb6-835d-440a-99a5-298ced25981a" ID-ZooBank="584332C1-8B4B-41B9-A17D-327F47B5616A" ID-publisher-id="158380" URI-arpha="C1B74D6C-6F55-56D8-B801-6574AC43F552" URI-zoobank="http://zoobank.org/584332C1-8B4B-41B9-A17D-327F47B5616A" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1760020140826" checkinUser="pensoft" docAuthor="Benjamin, Suresh P., Dhiya’ulhaq, Naufal Urfi, Deeleman-Reinhold, Christa, Buchori, Damayanti, Hidayat, Purnama, Scheu, Stefan &amp; Drescher, Jochen" docDate="2025" docId="6487E2EB928A57BE84E8F71297687052" docLanguage="en" docName="ZooKeys 1255: 95-125" docOrigin="ZooKeys 1255" docSource="https://zookeys.pensoft.net/article/158380/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Zametopina calceata Simon 1909" docType="treatment" docVersion="6" dtd-version="3.0" lastPageNumber="125" masterDocId="C1B74D6C6F5556D8B8016574AC43F552" masterDocTitle="Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia" masterLastPageNumber="125" masterPageNumber="95" pageNumber="95" updateTime="1760033433488" updateUser="ExternalLinkService">
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<mods:title id="A5FFCABBD79B3649AED7BEB464535F71">Taxonomic notes on the crab-spider genera Nyctimus Thorell, 1877 and Zametopina Simon, 1909 (Araneae, Thomisidae) with descriptions of six new species from Southeast Asia</mods:title>
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<mods:namePart id="749F139CF2345A564B185E05BC7D124E">Benjamin, Suresh P.</mods:namePart>
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<mods:namePart id="A286DD7048BFBD251229C5992A09E6B3">Buchori, Damayanti</mods:namePart>
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<mods:affiliation id="C85471DC5ADC48ECD8388DD8FD75B4D8">Department of Plant Protection, Faculty of Agriculture, IPB University, Kampus IPB Dramaga Bogor, Jl. Raya Dramaga, West Java 16680, Indonesia</mods:affiliation>
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<mods:affiliation id="C5D9ABE6A749A7A070E47242DCD27819">Department of Plant Protection, Faculty of Agriculture, IPB University, Kampus IPB Dramaga Bogor, Jl. Raya Dramaga, West Java 16680, Indonesia</mods:affiliation>
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<mods:namePart id="3EA236783EFFAB6950BABDA5ED7D0BA8">Scheu, Stefan</mods:namePart>
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<mods:affiliation id="8DD51D4B2A61F89840DC0BB512970FEA">Animal Ecology, J. F. Blumenbach Institute of Zoology and Anthropology, University of Göttingen, Untere Karspüle 2, Göttingen 37073, Germany</mods:affiliation>
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<treatment id="6487E2EB928A57BE84E8F71297687052" ID-DOI="http://doi.org/10.5281/zenodo.17306149" ID-GBIF-Taxon="302826298" ID-Zenodo-Dep="17306149" ID-arpha="6487E2EB-928A-57BE-84E8-F71297687052" LSID="urn:lsid:plazi:treatment:6487E2EB928A57BE84E8F71297687052" httpUri="http://treatment.plazi.org/id/6487E2EB928A57BE84E8F71297687052" scope_family="Thomisidae" scope_order="Araneae">
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<paragraph id="0E9527BE1DF71BAF03142FB306196879">
<taxonomicName id="1575D12434C3B3CE0ED7C66C38CC8F97" ID-CoL="5CVNT" authority="Simon, 1909" authorityName="Simon" authorityYear="1909" class="Arachnida" family="Thomisidae" genus="Zametopina" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="calceata">
<emphasis id="06E2536A6563B8A924E0BFCCE30A307F" italics="true">Zametopina calceata</emphasis>
<bibRefCitation id="8B1DA16A022442BACFFD76FA9E0B6845" DOI="10.5962/bhl.part.24151" author="Simon E" journalOrPublisher="Bulletin Biologique de la France et de la Belgique" pagination="69-147" refId="B15" refString="Simon E (1909) Etude sur les arachnides du Tonkin (1 re partie). Bulletin Biologique de la France et de la Belgique 42: 69-147. https://doi.org/10.5962/bhl.part.24151" title="Etude sur les arachnides du Tonkin (1 re partie)." volume="42" year="1909">Simon, 1909</bibRefCitation>
</taxonomicName>
</paragraph>
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<taxonomicName id="BA5506AD592342924BDBD2E9D548AFE6" ID-CoL="5CVNT" authority="Simon, 1909" authorityName="Simon" authorityYear="1909" class="Arachnida" family="Thomisidae" genus="Zametopina" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="calceata">
<emphasis id="FF6D93E8EF370F1E3A3DA40C929793F4" italics="true">Zametopina calceata</emphasis>
<comment id="5CC134246FCA9FF3B1BA75939CF8A968">
<bibRefCitation id="3B650CDC8763F1200941DAAE9FD076F4" DOI="10.5962/bhl.part.24151" author="Simon E" journalOrPublisher="Bulletin Biologique de la France et de la Belgique" pagination="69-147" refId="B15" refString="Simon E (1909) Etude sur les arachnides du Tonkin (1 re partie). Bulletin Biologique de la France et de la Belgique 42: 69-147. https://doi.org/10.5962/bhl.part.24151" title="Etude sur les arachnides du Tonkin (1 re partie)." volume="42" year="1909">Simon, 1909</bibRefCitation>
: 
<treatmentCitation id="420FEE1E6C172D1AC0E13363940A8695" author="Simon E" page="123" year="1909">
<bibRefCitation id="CCF184CA0A963B684763E61918CAD209" DOI="10.5962/bhl.part.24151" author="Simon E" firstAuthor="Simon" journalOrPublisher="Bulletin Biologique de la France et de la Belgique" page="123" pagination="69-147" refId="B15" refString="Simon E (1909) Etude sur les arachnides du Tonkin (1 re partie). Bulletin Biologique de la France et de la Belgique 42: 69–147. https://doi.org/10.5962/bhl.part.24151" title="Etude sur les arachnides du Tonkin (1 re partie)." volume="42" year="1909">Simon 1909: 123</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="7DE11E118A04277F0BA65F30940509FA" ID-DOI="http://doi.org/10.5281/zenodo.5295462" ID-Zenodo-Dep="5295462" author="Tang G &amp; Blick T &amp; Ono H" httpUri="http://treatment.plazi.org/id/1605DA212003FFBEFF7BFD47FADB2830" page="66" year="2010">
<bibRefCitation id="F34B1439B530CE728EDF2A37186876C0" author="Tang G &amp; Blick T &amp; Ono H" etAl="et al." firstAuthor="Tang" journalOrPublisher="Bulletin of the National Museum of Nature and Science Tokyo (A)" pagination="65-70" refId="B16" refString="Tang G, Blick T, Ono H (2010) Rediscovery of an obscure spider genus Zametopina Simon, 1909 (Araneae, Thomisidae) from Yunnan, China. Bulletin of the National Museum of Nature and Science Tokyo (A) 36: 65–70." title="Rediscovery of an obscure spider genus Zametopina Simon, 1909 (Araneae, Thomisidae) from Yunnan, China." volume="36" year="2010">Tang et al. (2010)</bibRefCitation>
: 66
</treatmentCitation>
, figs 1-18.
</comment>
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</paragraph>
</subSubSection>
<subSubSection id="SECID0EQOAI" type="type material">
<paragraph id="5E4867C2557288F7F6C904723DAFAF06">
<heading id="203ADCC8F2CC0C4603B6C088BA6E40AF" reason="title">Type material.</heading>
</paragraph>
<paragraph id="6795C2443EC829996CD6EE0EBD9D5CF6">
<materialsCitation id="4728526C7191B67C8C9FE414274BAB86" ID-GBIF-Occurrence="5838966302" collectionCode="MNHN" collectorName="de Pelacot" country="Vietnam" county="Boa Luc" location="a l'Ouest de Cao-Bang et pres la frontiere du Yun-Nan et du Kuang-Si" municipality="Haut-Tonkin" specimenCode="MNHN 22937" specimenCount="1" specimenCount-male="1" stateProvince="Cao B ang Province" typeStatus="holotype">
<emphasis id="3C40984793B5C34ADCA640231449B493" bold="true">
<emphasis id="9417EDD91F0C1726F3A7F602109A2BE7" bold="true" italics="true">
<typeStatus id="5CA8A2B7FD30F6F7C57E46868E4DC0C8" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="72E58BB5C0FDE58B5D3535AD50687B1A" name="Vietnam">Vietnam</collectingCountry>
– 
<collectingRegion id="CB14AB9313678CE79B40DBBEE9AC8D62">Cao B ằng Province</collectingRegion>
• 
<specimenCount id="7F982179D82B7A68291E0DE5D0876CFC" count="1" type="male">♂</specimenCount>
; “ 
<collectingCounty id="8EAA43423EC7FD41BC2F5E70A0401010">Boa Luc</collectingCounty>
, dans 
<collectingMunicipality id="C2FA17FEB5BB51BDB8D942F8E9F09B7C">Haut-Tonkin</collectingMunicipality>
, 
<location id="4BB5E5C78D8FB87ADF8D50203D64C1E9" LSID="urn:lsid:plazi:treatment:6487E2EB928A57BE84E8F71297687052:4BB5E5C78D8FB87ADF8D50203D64C1E9" country="Vietnam" county="Boa Luc" municipality="Haut-Tonkin" name="a l'Ouest de Cao-Bang et pres la frontiere du Yun-Nan et du Kuang-Si" stateProvince="Cao B ang Province">à l’Ouest de Cao-Bang et prés la frontière du Yun-Nan et du Kuang-Si</location>
” (probably B ảo L ạc District); 1906 or 1907; 
<collectorName id="FA824855661E0C672B1C2BFDBA9C6471">de Pelacot</collectorName>
leg.; 
<specimenCode id="870CEFF2C513613D9301DC9EDAB42FA3" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">
<named-content id="7F6312070A30C36AD7907E314673FFC8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum National d’Histoire Naturelle, Paris">MNHN</named-content>
22937
</specimenCode>
. Examined.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6OAI" type="diagnosis">
<paragraph id="8723657F3CA1409CE8F7B955EA9E46B2">
<heading id="9A5B15A6CAA4215E48B9770DC8DE8FDF" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="8603E63DF538D651AF2ADD223A738092">
See 
<bibRefCitation id="7349CAF703166BB0490E3343C8386F76" DOI="10.11865/zs.2023101" author="Lin YJ &amp; Li SQ &amp; Pham DS" etAl="et al." firstAuthor="Lin" issue="1" journalOrPublisher="Zoological Systematics" pagination="1-99" refId="B8" refString="Lin YJ, Li SQ, Pham DS (2023) Taxonomic notes on some spider species (Arachnida: Araneae) from China and Vietnam. Zoological Systematics 48 (1): 1–99. https://doi.org/10.11865/zs.2023101" title="Taxonomic notes on some spider species (Arachnida: Araneae) from China and Vietnam." volume="48" year="2023">Lin et al. (2023)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJPAI" type="description">
<paragraph id="FF10808F0E5BAB23796AEADBAFB5B9E3">
<heading id="A6B29DA19A90D0222B213F417E659AA3" reason="title">Description.</heading>
</paragraph>
<paragraph id="A8B83417FD40A19B6FE34EF6005463B6">
See 
<bibRefCitation id="8DFE31980DE39EADA607682ACE3FC139" author="Tang G &amp; Blick T &amp; Ono H" etAl="et al." firstAuthor="Tang" journalOrPublisher="Bulletin of the National Museum of Nature and Science Tokyo (A)" pagination="65-70" refId="B16" refString="Tang G, Blick T, Ono H (2010) Rediscovery of an obscure spider genus Zametopina Simon, 1909 (Araneae, Thomisidae) from Yunnan, China. Bulletin of the National Museum of Nature and Science Tokyo (A) 36: 65–70." title="Rediscovery of an obscure spider genus Zametopina Simon, 1909 (Araneae, Thomisidae) from Yunnan, China." volume="36" year="2010">Tang et al. (2010)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETPAI" type="distribution">
<paragraph id="F0BBAA47BD87637BBF47EB8904BB272A">
<heading id="5C1C00B1FBB360FBB3DE6F128635BD63" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="15BAAB253691A90628B6D3D82C5517F9">
<collectingCountry id="84A3DC93502E135F5BF39D7B96DC850A" name="China">China</collectingCountry>
(
<collectingRegion id="9CB694BF44222804F889099389ED3E4C" country="China" name="Yunnan">Yunnan</collectingRegion>
), 
<collectingCountry id="8F6942C915C4DFA0DD3F65CDAF1F07A8" name="Vietnam">Vietnam</collectingCountry>
(Cao B ằng) (Fig. 
<figureCitation id="7C3AE983821E0C80463FDEF6068D6343" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of all described Nyctimus and Zametopina (Araneae: Thomisidae) species. Grey circles surrounding white dots indicate species collected from the same or very closely located sampling points in Brunei, Sabah (Malaysia), and Jambi (Indonesia)." figureDoi="10.3897/zookeys.1255.158380.figure19" httpUri="https://binary.pensoft.net/fig/1434042">19</figureCitation>
).
</paragraph>
<caption id="CE304BA90A491906306248BAA647A3EC" ID-DOI="10.3897/zookeys.1255.158380.figure19" ID-arpha="25EAD12D-97A1-5D0F-B2EE-A5F83C6F9CB9" httpUri="https://binary.pensoft.net/fig/1434042" startId="F19">
<paragraph id="3409806FCD019EA6D684648BDE9E5CC6">
<label id="621AA6C922EAC1BEE9E25AE27FA0208B">Figure 19.</label>
</paragraph>
<paragraph id="A2AD8EF4D1F0CFB27783638505125646">
Distribution of all described 
<taxonomicName id="FC24A543A8E7E4D5AC2446FBAD03BC93" authorityName="Thorell" authorityYear="1877" class="Arachnida" family="Thomisidae" genus="Nyctimus" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="0F2639308BD46D339A50FBF58BA28F41" italics="true">Nyctimus</emphasis>
</taxonomicName>
and 
<taxonomicName id="A2CF8A27A4050751780CD302BD67C518" authorityName="Simon" authorityYear="1909" class="Arachnida" family="Thomisidae" genus="Zametopina" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="A3BB60FAB1D035D6C9A9B41D6832EDB4" italics="true">Zametopina</emphasis>
</taxonomicName>
(
<taxonomicName id="6B6825659EE664B3326935B1536B53F1" class="Arachnida" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="order">Araneae</taxonomicName>
: 
<taxonomicName id="4D19D043E572998CD894690ABB6F0A55" authorityName="Sundevall" authorityYear="1833" class="Arachnida" family="Thomisidae" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="family">Thomisidae</taxonomicName>
) species. Grey circles surrounding white dots indicate species collected from the same or very closely located sampling points in Brunei, Sabah (Malaysia), and Jambi (Indonesia).
</paragraph>
</caption>
</subSubSection>
</treatment>
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