<document id="FED4598847FE6DEEA50DBC56282395D8" ID-CLB-Dataset="79793" ID-DOI="10.5281/zenodo.6709103" ID-GBIF-Dataset="715ac68b-c7fd-4eac-9128-b57bfd6ecddf" ID-ISBN="978-84-96553-89-7" ID-Zenodo-Dep="6709103" IM.metadata_requiresApprovalFor="plazi" IM.taxonomicNames_requiresApprovalFor="plazi" checkinTime="1656002980909" checkinUser="jonas" docAuthor="Russell A. Mittermeier, Anthony B. Rylands &amp; Don E. Wilson" docDate="2013" docId="5D3287905C43FFFCAED7F6CC8DEBF3ED" docLanguage="en" docName="hbmw_3_Indriidae_0142.pdf.imf" docOrigin="Handbook of the Mammals of the World – Volume 3 Primates, Barcelona: Lynx Edicions" docTitle="Avahi laniger" docType="treatment" docVersion="11" lastPageNumber="164" masterDocId="A10BFFE85C42FFFDAE67FFDF8512FFEF" masterDocTitle="Indriidae" masterLastPageNumber="175" masterPageNumber="142" pageNumber="164" updateTime="1732884232467" updateUser="ExternalLinkService">
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<mods:title id="5A7CF52D92938143014F2D4B9E69079E">Indriidae</mods:title>
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<mods:namePart id="9FA05CE7441A38D594DC23E1873C7553">Russell A. Mittermeier</mods:namePart>
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<mods:namePart id="6C2A621E82586C4AD60A112887FEA939">Anthony B. Rylands</mods:namePart>
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<mods:namePart id="0F946E83638BC014BDF9E6B5A7DB19B0">Don E. Wilson</mods:namePart>
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<mods:dateIssued id="DB784575B16AE176F391E335BB3142FF">2013</mods:dateIssued>
<mods:dateOther id="5E160CF49F12C71A261091BD510CF011" type="pubDate">2013-03-31</mods:dateOther>
<mods:publisher id="6230C96CC1B8563720DD3141CF836968">Lynx Edicions</mods:publisher>
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<mods:placeTerm id="BF9E70444D388197DE3740CEAF4B0185">Barcelona</mods:placeTerm>
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<mods:title id="3FCD5F71BABEEDB3C391C6423635AF7D">Handbook of the Mammals of the World – Volume 3 Primates</mods:title>
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<treatment id="5D3287905C43FFFCAED7F6CC8DEBF3ED" ID-DOI="http://doi.org/10.5281/zenodo.6708793" ID-GBIF-Taxon="196159553" ID-Zenodo-Dep="6708793" LSID="urn:lsid:plazi:treatment:5D3287905C43FFFCAED7F6CC8DEBF3ED" httpUri="http://treatment.plazi.org/id/5D3287905C43FFFCAED7F6CC8DEBF3ED" lastPageNumber="164" pageId="1" pageNumber="164" scope_class="Mammalia" scope_infraOrder="Llemuriformes" scope_order="Primates" scope_subOrder="Strepsirrhini">
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<paragraph id="D52436865C43FFFCAED7F6CC85DFF6AE" blockId="1.[173,1036,2323,2471]" box="[176,205,2323,2369]" pageId="1" pageNumber="164">
<heading id="8E6C81EA5C43FFFCAED7F6CC85DFF6AE" box="[176,205,2323,2369]" pageId="1" pageNumber="164">
<figureCitation id="4DA02A035C43FFFCAED7F6CC85DFF6AE" box="[176,205,2323,2369]" captionStart="On" captionStartId="6.[80,110,3397,3422]" captionTargetBox="[16,2704,15,3633]" captionTargetPageId="5" captionText="On following pages: 12. Crowned Sifaka (Propithecus coronatus); 13. Coquerel’s Sifaka (Propithecus coquerell); 14. Tattersall’s Sifaka (Propithecus tattersall); 15. Diademed Sifaka (Propithecus diadema); 16. Milne-Edwards's Sifaka (Propithecus edwardsi); 17. Silky Sifaka (Propithecus candidus); 18. Perrier’s Sifaka (Propithecus perrieri)." figureDoi="http://doi.org/10.5281/zenodo.6709199" httpUri="https://zenodo.org/record/6709199/files/figure.png" pageId="1" pageNumber="164">1.</figureCitation>
</heading>
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<subSubSection id="9D81650D5C43FFFCAEB9F6CC87A2F6AE" box="[222,688,2323,2369]" pageId="1" pageNumber="164" type="vernacular_names">
<paragraph id="D52436865C43FFFCAEB9F6CC87A2F6AE" blockId="1.[173,1036,2323,2471]" box="[222,688,2323,2369]" pageId="1" pageNumber="164">
<heading id="8E6C81EA5C43FFFCAEB9F6CC87A2F6AE" box="[222,688,2323,2369]" pageId="1" pageNumber="164">
<vernacularName id="5B9846A85C43FFFCAEB9F6CC87A2F6AE" ID-CoL="K29Q" baseAuthorityName="Gmelin" baseAuthorityYear="1788" box="[222,688,2323,2369]" class="Mammalia" family="Indriidae" genus="Avahi" kingdom="Animalia" language="eng" order="Primates" pageId="1" pageNumber="164" phylum="Chordata" rank="species" species="laniger">Eastern Woolly Lemur</vernacularName>
</heading>
</paragraph>
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<subSubSection id="9D81650D5C43FFFCAC93F6CC86F4F6AE" box="[756,998,2323,2369]" pageId="1" pageNumber="164" type="nomenclature">
<paragraph id="D52436865C43FFFCAC93F6CC86F4F6AE" blockId="1.[173,1036,2323,2471]" box="[756,998,2323,2369]" pageId="1" pageNumber="164">
<heading id="8E6C81EA5C43FFFCAC93F6CC86F4F6AE" box="[756,998,2323,2369]" pageId="1" pageNumber="164">
<taxonomicName id="129B4D055C43FFFCAC93F6CC86F4F6AE" ID-CoL="K29Q" baseAuthorityName="Gmelin" baseAuthorityYear="1788" box="[756,998,2323,2369]" class="Mammalia" family="Indriidae" genus="Avahi" kingdom="Animalia" order="Primates" pageId="1" pageNumber="164" phylum="Chordata" rank="species" species="laniger">
<emphasis id="E7EFEA945C43FFFCAC93F6CC86F4F6AE" box="[756,998,2323,2369]" italics="true" pageId="1" pageNumber="164">Avahi laniger</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9D81650D5C43FFFCAECAF68D85C6F648" pageId="1" pageNumber="164" type="vernacular_names">
<paragraph id="D52436865C43FFFCAECAF68D811EF688" blockId="1.[173,1036,2323,2471]" box="[173,1036,2386,2407]" pageId="1" pageNumber="164">
<heading id="8E6C81EA5C43FFFCAECAF68D811EF688" box="[173,1036,2386,2407]" pageId="1" pageNumber="164">
<emphasis id="E7EFEA945C43FFFCAECAF68D85EBF688" bold="true" box="[173,249,2386,2407]" pageId="1" pageNumber="164">French:</emphasis>
<vernacularName id="5B9846A85C43FFFCAF65F68D8496F688" ID-CoL="K29Q" baseAuthorityName="Gmelin" baseAuthorityYear="1788" box="[258,388,2386,2407]" class="Mammalia" family="Indriidae" genus="Avahi" kingdom="Animalia" language="fra" order="Primates" pageId="1" pageNumber="164" phylum="Chordata" rank="species" species="laniger">Avahi laineux</vernacularName>
/ 
<emphasis id="E7EFEA945C43FFFCAFFEF68D84E1F688" bold="true" box="[409,499,2386,2407]" pageId="1" pageNumber="164">German:</emphasis>
<vernacularName id="5B9846A85C43FFFCAF9AF68D87ABF688" ID-CoL="K29Q" baseAuthorityName="Gmelin" baseAuthorityYear="1788" box="[509,697,2386,2407]" class="Mammalia" family="Indriidae" genus="Avahi" kingdom="Animalia" language="deu" order="Primates" pageId="1" pageNumber="164" phylum="Chordata" rank="species" species="laniger">Ostlicher Wollmaki</vernacularName>
/ 
<emphasis id="E7EFEA945C43FFFCACA9F68D863BF688" bold="true" box="[718,809,2386,2407]" pageId="1" pageNumber="164">Spanish:</emphasis>
<vernacularName id="5B9846A85C43FFFCAD54F68D811EF688" ID-CoL="K29Q" baseAuthorityName="Gmelin" baseAuthorityYear="1788" box="[819,1036,2386,2407]" class="Mammalia" family="Indriidae" genus="Avahi" kingdom="Animalia" language="esp" order="Primates" pageId="1" pageNumber="164" phylum="Chordata" rank="species" species="laniger">Lémur lanudo oriental</vernacularName>
</heading>
</paragraph>
<paragraph id="D52436865C43FFFCAECAF6A585C6F648" blockId="1.[173,1036,2323,2471]" pageId="1" pageNumber="164">
<heading id="8E6C81EA5C43FFFCAECAF6A585C6F648" pageId="1" pageNumber="164">
<emphasis id="E7EFEA945C43FFFCAECAF6A584B0F660" bold="true" box="[173,418,2426,2447]" pageId="1" pageNumber="164">Other common names:</emphasis>
<vernacularName id="5B9846A85C43FFFCAFCAF6A58727F660" ID-CoL="K29Q" baseAuthorityName="Gmelin" baseAuthorityYear="1788" box="[429,565,2426,2447]" class="Mammalia" family="Indriidae" genus="Avahi" kingdom="Animalia" language="eng" order="Primates" pageId="1" pageNumber="164" phylum="Chordata" rank="species" species="laniger">Eastern Avahi</vernacularName>
, 
<vernacularName id="5B9846A85C43FFFCAC24F6A585C6F648" ID-CoL="K29Q" baseAuthorityName="Gmelin" baseAuthorityYear="1788" class="Mammalia" family="Indriidae" genus="Avahi" kingdom="Animalia" language="eng" order="Primates" pageId="1" pageNumber="164" phylum="Chordata" rank="species" species="laniger">Gmelin’s Woolly Lemur my</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9D81650D5C43FFFCAD73F61F804EF632" box="[788,1372,2496,2525]" pageId="1" pageNumber="164" type="reference_group">
<paragraph id="D52436865C43FFFCAD73F61F804EF632" blockId="1.[788,1374,2496,2918]" box="[788,1372,2496,2525]" pageId="1" pageNumber="164">
<emphasis id="E7EFEA945C43FFFCAD73F61F86BDF632" bold="true" box="[788,943,2496,2525]" pageId="1" pageNumber="164">Taxonomy.</emphasis>
<taxonomicName id="129B4D055C43FFFCADA4F61F8045F632" authority="Gmelin, 1788" authorityName="Gmelin" authorityYear="1788" box="[963,1367,2496,2525]" class="Mammalia" family="Indriidae" genus="Lemur" kingdom="Animalia" order="Primates" pageId="1" pageNumber="164" phylum="Chordata" rank="species" species="laniger">Lemur laniger Gmelin, 1788</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection id="9D81650D5C43FFFCAD71F63886B7F5C3" pageId="1" pageNumber="164" type="materials_examined">
<paragraph id="D52436865C43FFFCAD71F63886B7F5C3" blockId="1.[788,1374,2496,2918]" pageId="1" pageNumber="164">
<materialsCitation id="65F33CDB5C43FFFCAD71F63886B7F5C3" ID-GBIF-Occurrence="3818561309" pageId="1" pageNumber="164">Madagascar, Betanimena country, Antongil Bay.</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="9D81650D5C43FFFCAD73F5E9816FF5BC" box="[788,1149,2614,2643]" pageId="1" pageNumber="164" type="discussion">
<paragraph id="D52436865C43FFFCAD73F5E9816FF5BC" blockId="1.[788,1374,2496,2918]" box="[788,1149,2614,2643]" pageId="1" pageNumber="164">This species is monotypic.</paragraph>
</subSubSection>
<subSubSection id="9D81650D5C43FFFCAD73F58686DEF462" pageId="1" pageNumber="164" type="distribution">
<caption id="81E4660E5C43FFFCAD73F58686DEF462" ID-DOI="http://doi.org/10.5281/zenodo.6709111" ID-Zenodo-Dep="6709111" httpUri="https://zenodo.org/record/6709111/files/figure.png" inLine="true" pageId="1" pageNumber="164" targetBox="[176,756,2503,2910]" targetPageId="1">
<paragraph id="D52436865C43FFFCAD73F58686DEF462" blockId="1.[788,1374,2496,2918]" lastBlockId="1.[173,1375,2924,3468]" pageId="1" pageNumber="164">
<emphasis id="E7EFEA945C43FFFCAD73F58686D6F595" bold="true" box="[788,964,2649,2682]" pageId="1" pageNumber="164">Distribution.</emphasis>
NE &amp; E Madagascar from the Bemarivo River in the N to the Mangoro/Nesivolo rivers in the S; prior to the recent taxonomic splitting of this species, it was thought to range almost the entire length of E Madagascar’s rainforests from the Ankarana Massif in the extreme N to the Tolagnaro (= Fort-Dauphin) region in the extreme S.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="9D81650D5C43FFFCAECAF44C804EF3BD" pageId="1" pageNumber="164" type="description">
<paragraph id="D52436865C43FFFCAECAF44C804EF3BD" blockId="1.[173,1375,2924,3468]" pageId="1" pageNumber="164">
<emphasis id="E7EFEA945C43FFFCAECAF44C84B0F45B" bold="true" box="[173,418,2963,2996]" pageId="1" pageNumber="164">Descriptive notes.</emphasis>
Head—body 27.7-32.2 cm,tail 30.4-36.6 cm; weight 1.1-1.3 kg. The dorsal coat of the Eastern Woolly Lemur is gray-brown to reddish, becoming paler toward the rusty-red tail. The chest and abdomen are gray. The face is brownish, with a lighter band or distinct patches above the eyes and lighter fur on the cheeks and throat. Small ears are largely hidden by thick fur. Dense, short fur is tightly curled on the back.
</paragraph>
</subSubSection>
<subSubSection id="9D81650D5C43FFFCAEC9F3BF8120F396" box="[174,1074,3168,3193]" pageId="1" pageNumber="164" type="biology_ecology">
<paragraph id="D52436865C43FFFCAEC9F3BF8120F396" blockId="1.[173,1375,2924,3468]" box="[174,1074,3168,3193]" pageId="1" pageNumber="164">
<emphasis id="E7EFEA945C43FFFCAEC9F3BF840EF396" bold="true" box="[174,284,3168,3193]" pageId="1" pageNumber="164">Habitat.</emphasis>
Tropical moist lowland, montane, and secondary forest.
</paragraph>
</subSubSection>
<subSubSection id="9D81650D5C43FFFCAEC9F3588607F327" pageId="1" pageNumber="164" type="food_feeding">
<paragraph id="D52436865C43FFFCAEC9F3588607F327" blockId="1.[173,1375,2924,3468]" pageId="1" pageNumber="164">
<emphasis id="E7EFEA945C43FFFCAEC9F35884A0F34F" bold="true" box="[174,434,3207,3232]" pageId="1" pageNumber="164">Food and Feeding.</emphasis>
The Eastern Woolly Lemur eats mainly immature leaves and buds, supplemented with fruits, flowers, and bark.
</paragraph>
</subSubSection>
<subSubSection id="9D81650D5C43FFFCAEC9F31186E2F300" box="[174,1008,3278,3311]" pageId="1" pageNumber="164" type="breeding">
<paragraph id="D52436865C43FFFCAEC9F31186E2F300" blockId="1.[173,1375,2924,3468]" box="[174,1008,3278,3311]" pageId="1" pageNumber="164">
<emphasis id="E7EFEA945C43FFFCAEC9F3118426F300" bold="true" box="[174,308,3278,3311]" pageId="1" pageNumber="164">Breeding.</emphasis>
There is no information available for this species.
</paragraph>
</subSubSection>
<subSubSection id="9D81650D5C43FFFCAECAF3268648F2D2" pageId="1" pageNumber="164" type="activity">
<paragraph id="D52436865C43FFFCAECAF3268648F2D2" blockId="1.[173,1375,2924,3468]" pageId="1" pageNumber="164">
<emphasis id="E7EFEA945C43FFFCAECAF3268488F2F9" bold="true" box="[173,410,3321,3350]" pageId="1" pageNumber="164">Activity patterns.</emphasis>
There is no specific information available for this species, but the Eastern Woolly Lemur is nocturnal and arboreal.
</paragraph>
</subSubSection>
<subSubSection id="9D81650D5C43FFFCAEC8F29783FFF6D0" pageId="1" pageNumber="164" type="biology_ecology">
<paragraph id="D52436865C43FFFCAEC8F29783FFF6D0" blockId="1.[173,1375,2924,3468]" lastBlockId="1.[1443,2650,2180,3077]" pageId="1" pageNumber="164">
<emphasis id="E7EFEA945C43FFFCAEC8F297868AF28A" bold="true" box="[175,920,3400,3429]" pageId="1" pageNumber="164">Movements, Home range and Social organization.</emphasis>
Groups of up to five Eastern Woolly Lemurs have been reported, and home ranges of 1-2 ha are defended aggressively. Small group size indicates monogamous groupings, but longitudinal and genetic data are needed to better understand this social behavior. Research at the Analamazaotra Special Reserve suggests a social organization based on monogamous pairs and their offspring. In Analamazaotra, densities have been estimated to reach 72-100 ind/km?.
</paragraph>
</subSubSection>
<subSubSection id="9D81650D5C43FFFCABC2F6968C53F4D2" pageId="1" pageNumber="164" type="conservation">
<paragraph id="D52436865C43FFFCABC2F6968C53F4D2" blockId="1.[1443,2650,2180,3077]" pageId="1" pageNumber="164">
<emphasis id="E7EFEA945C43FFFCABC2F6968212F689" bold="true" box="[1445,1792,2377,2406]" pageId="1" pageNumber="164">Status and Conservation.</emphasis>
CITES Appendix I. Classified as Least Concern on The [UCN Red List. However, at the IUCN/SSC Lemur Red-Listing Workshop held in July 2012, A. lanigerwas re-assessed and elevated to endangered. It suffers mainly from habitat destruction due to logging and slash-and-burn agriculture. It is sometimes captured opportunistically at its daytime sleeping sites or in traps baited with fruit. In some places, it is actively pursued by hunters with slingshots (Mananara-Nord) or spears (Makira). Indeed, recent data indicate that there has been a considerable upsurge in hunting of the Eastern Woolly Lemur. It occurs in four national parks (Mananara-Nord, Mantadia, Marojejy, and Zahamena), two strict nature reserves (Betampona and Zahamena), five special reserves (Ambatovaky, Ambohitantely, Analamazaotra, Anjanaharibe-Sud, and Mangerivola), and the soon-to-be-protected forests of Makira. It is still unclear which species of woolly lemur occurs in Ankarana National Park; for now,it is tentatively considered to be the Western Woolly Lemur (A. 
<taxonomicName id="129B4D055C43FFFCA6F8F4FF8C27F4D2" authorityName="Lorenz" authorityYear="1898" box="[2207,2357,2848,2877]" class="Mammalia" family="Indriidae" genus="Avahi" kingdom="Animalia" order="Primates" pageId="1" pageNumber="164" phylum="Chordata" rank="species" species="occidentalis">occidentalis</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="9D81650D5C43FFFCABC1F4948DEBF3ED" pageId="1" pageNumber="164" type="bibRefCitation_list">
<paragraph id="D52436865C43FFFCABC1F4948DEBF3ED" blockId="1.[1443,2650,2180,3077]" pageId="1" pageNumber="164">
<emphasis id="E7EFEA945C43FFFCABC1F4948352F48B" bold="true" box="[1446,1600,2891,2916]" pageId="1" pageNumber="164">Bibliography.</emphasis>
Albignac (1981b), Andriantompohavana et al. (2007), Britt et al. (1999), Fowler et al. (1989), Ganzhorn (1988), Ganzhorn et al. (1985), Glander et al. (1992), Goodman, S.M. et al. (1998), Harcourt (1991), Hawkins et al. (1990), Lei et al. (2008), Mittermeier et al. (2010), Napier &amp; Napier (1967), Nicoll &amp; Langrand (1989), Petter et al. (1977), Razanahoera-Rakotomalala (1981), Rumpler et al. (2011), Schmid &amp; Smolker (1998), Sterling &amp; Ramaroson (1996), Tattersall (1982, 1986b), Zaramody et al. (2006).
</paragraph>
</subSubSection>
</treatment>
</document>