<document id="497C3EA6D9858A3AA4FBFFCB91C572CB" ID-CLB-Dataset="102847" ID-DOI="10.5281/zenodo.6870843" ID-GBIF-Dataset="8a6c6dcf-1911-46fc-8ea0-48fe55c5ed11" ID-ISBN="978-84-16728-08-4" ID-Zenodo-Dep="6870843" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1658335596803" checkinUser="felipe" docAuthor="Russell A. Mittermeier &amp; Don E. Wilson" docDate="2018" docId="3D474A54A061870DFF26AD8F1350F328" docLanguage="en" docName="hbmw_8_Soricidae_0332.pdf.imf" docOrigin="Handbook of the Mammals of the World – Volume 8 Insectivores, Sloths and Colugos, Barcelona: Lynx Edicions" docTitle="Sylvisorex isabellae" docType="treatment" docVersion="10" lastPageNumber="468" masterDocId="C17E322CA0288744FF8DAB47125EFFF9" masterDocTitle="Soricidae" masterLastPageNumber="551" masterPageNumber="332" pageNumber="468" updateTime="1768228605763" updateUser="ExternalLinkService">
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<mods:title id="45A59CEAC19B49F55C621403FC7362F4">Soricidae</mods:title>
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<mods:namePart id="04EEBDE0B0B5D0DC55375A4E032C8692">Russell A. Mittermeier</mods:namePart>
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<mods:namePart id="4275B7B24B233E1051CBC896CD246C66">Don E. Wilson</mods:namePart>
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<mods:title id="4F755089AE06F2148E733669DA36C3D3">Handbook of the Mammals of the World – Volume 8 Insectivores, Sloths and Colugos</mods:title>
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<treatment id="3D474A54A061870DFF26AD8F1350F328" ID-DOI="http://doi.org/10.5281/zenodo.6870038" ID-GBIF-Taxon="197841062" ID-Zenodo-Dep="6870038" LSID="urn:lsid:plazi:treatment:3D474A54A061870DFF26AD8F1350F328" httpUri="http://treatment.plazi.org/id/3D474A54A061870DFF26AD8F1350F328" lastPageNumber="468" pageId="73" pageNumber="468" scope_class="Mammalia" scope_family="Soricidae" scope_order="Eulipotyphla">
<subSubSection id="FDF4A8C9A061870DFF26AD8F12A3F903" box="[171,253,1736,1786]" pageId="73" pageNumber="468" type="multiple">
<paragraph id="B551FB42A061870DFF26AD8F12A3F903" blockId="73.[168,1359,1736,1863]" box="[171,253,1736,1786]" pageId="73" pageNumber="468">
<heading id="EE194C2EA061870DFF26AD8F12A3F903" box="[171,253,1736,1786]" pageId="73" pageNumber="468">
<figureCitation id="2DD5E7C7A061870DFF26AD8F12A3F903" box="[171,253,1736,1786]" captionStart="Plate 19: Soricidae" captionStartId="64.[118,148,3136,3161]" captionTargetBox="[11,2741,14,3634]" captionTargetPageId="63" captionText="182. Palawan Moss Shrew (Palawanosorex muscorum), 183. Pearson’s Long-clawed Shrew (Solisorex pearsoni), 184. Kelaart’s Long-clawed Shrew (Feroculus feroculus), 185. Etruscan Shrew (Suncus etruscus), 186. Sri Lankan Shrew (Suncus fellowesgordoni), 187. Malayan Pygmy Shrew (Suncus malayanus), 188. Bornean Pygmy Shrew (Suncus hosei), 189. Jungle Shrew (Suncus zeylanicus), 190. Asian House Shrew (Suncus murinus), 191. Sr 1 Lankan Highland Shrew (Suncus montanus), 192. Nilgir 1 Highland Shrew (Suncus niger), 193. Anderson’s Shrew (Suncus stoliczkanus), 194. Day’s Shrew (Suncus dayi), 195. Flores Shrew (Suncus mertensi), 196. Black Shrew (Suncus ater), 197. Taita Shrew (Suncus aequatorius), 198. Greater Dwarf Shrew (Suncus lixa), 199. Hutu-Tutsi Dwarf Shrew (Suncus hututsi), 200. Least Dwarf Shrew (Suncus infinitestmus), 201. Remy’s Pygmy Shrew (Suncus remyi), 202. Lesser Dwarf Shrew (Suncus varilla), 203. Climbing Shrew (Suncus megalurus), 204. Rwenzori Shrew (Ruwenzorisorex suncoides), 205. Johnston's Forest Shrew (Sylvisorex johnstoni), 206. Akaibe’s Forest Shrew (Sylvisorex akaiber), 207. Moon Forest Shrew (Sylvisorex lunaris), 208. Greater Forest Shrew (Sylvisorex ollula), 209. Lesser Forest Shrew (Sylvisorex oriundus), 210. Bioko Forest Shrew (Sylvisorex isabellae), 211. Corbet’s Forest Shrew (Sylvisorex corbeti), 212. Bamenda Forest Shrew (Sylvisorex silvanorum), 213. Rainforest Shrew (Sylvisorex pluvialis), 214. Cameroonian Forest Shrew (Sylvisorex camerunensis), 215. Mount Cameroon Forest Shrew (Sylvisorex moro), 216. Kongana Forest Shrew (Sylvisorex konganensis), 217. Volcano Forest Shrew (Sylvisorex vulcanorum), 218. Grant's Forest Shrew (Sylvisorex granti), 219. Howell's Forest Shrew (Sylvisorex howelli), 220. Armored Hero Shrew (Scutisorex somereni), 221. Thor’s Hero Shrew (Scutisorex thor), 222. Greater Large-headed Shrew (Paracrocidura maxima), 223. Lesser Large-headed Shrew (Paracrocidura schoutedeni), 224. Grauer’s Large-headed Shrew (Paracrocidura graven), 225. Piebald Shrew (Diplomesodon pulchellus)" figureDoi="http://doi.org/10.5281/zenodo.6871917" httpUri="https://zenodo.org/record/6871917/files/figure.png" pageId="73" pageNumber="468">210.</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FDF4A8C9A061870DFE80AD8F10FDF903" box="[269,675,1736,1786]" pageId="73" pageNumber="468" type="vernacular_names">
<paragraph id="B551FB42A061870DFE80AD8F10FDF903" blockId="73.[168,1359,1736,1863]" box="[269,675,1736,1786]" pageId="73" pageNumber="468">
<heading id="EE194C2EA061870DFE80AD8F10FDF903" box="[269,675,1736,1786]" pageId="73" pageNumber="468">
<vernacularName id="3BED8B6CA061870DFE80AD8F10FDF903" ID-CoL="53NBP" baseAuthorityName="Heim de Balsac" baseAuthorityYear="1968" box="[269,675,1736,1786]" class="Mammalia" family="Soricidae" genus="Sylvisorex" kingdom="Animalia" language="eng" order="Soricomorpha" pageId="73" pageNumber="468" phylum="Chordata" rank="species" species="isabellae">Bioko Forest Shrew</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FDF4A8C9A061870DFD6BAD8F166CF903" box="[742,1074,1736,1786]" pageId="73" pageNumber="468" type="nomenclature">
<paragraph id="B551FB42A061870DFD6BAD8F166CF903" blockId="73.[168,1359,1736,1863]" box="[742,1074,1736,1786]" pageId="73" pageNumber="468">
<heading id="EE194C2EA061870DFD6BAD8F166CF903" box="[742,1074,1736,1786]" pageId="73" pageNumber="468">
<taxonomicName id="72EE80C1A061870DFD6BAD8F166CF903" ID-CoL="53NBP" baseAuthorityName="Heim de Balsac" baseAuthorityYear="1968" box="[742,1074,1736,1786]" class="Mammalia" family="Soricidae" genus="Sylvisorex" kingdom="Animalia" order="Soricomorpha" pageId="73" pageNumber="468" phylum="Chordata" rank="species" species="isabellae">
<emphasis id="879A2750A061870DFD6BAD8F166CF903" box="[742,1074,1736,1786]" italics="true" pageId="73" pageNumber="468">Sylvisorex isabellae</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FDF4A8C9A061870DFF27AC4C117EF8BE" pageId="73" pageNumber="468" type="vernacular_names">
<paragraph id="B551FB42A061870DFF27AC4C1710F8D9" blockId="73.[168,1359,1736,1863]" box="[170,1358,1803,1824]" pageId="73" pageNumber="468">
<heading id="EE194C2EA061870DFF27AC4C1710F8D9" box="[170,1358,1803,1824]" pageId="73" pageNumber="468">
<emphasis id="879A2750A061870DFF27AC4C12ABF8D9" bold="true" box="[170,245,1803,1824]" pageId="73" pageNumber="468">French:</emphasis>
<vernacularName id="3BED8B6CA061870DFF72AC4C13A6F8D9" ID-CoL="53NBP" baseAuthorityName="Heim de Balsac" baseAuthorityYear="1968" box="[255,504,1803,1824]" class="Mammalia" family="Soricidae" genus="Sylvisorex" kingdom="Animalia" language="fra" order="Soricomorpha" pageId="73" pageNumber="468" phylum="Chordata" rank="species" species="isabellae">Pachyure de Santa Isabel</vernacularName>
/ 
<emphasis id="879A2750A061870DFD83AC4C1036F8D9" bold="true" box="[526,616,1803,1824]" pageId="73" pageNumber="468">German:</emphasis>
<vernacularName id="3BED8B6CA061870DFDFFAC4C11FDF8D9" ID-CoL="53NBP" baseAuthorityName="Heim de Balsac" baseAuthorityYear="1968" box="[626,931,1803,1824]" class="Mammalia" family="Soricidae" genus="Sylvisorex" kingdom="Animalia" language="deu" order="Soricomorpha" pageId="73" pageNumber="468" phylum="Chordata" rank="species" species="isabellae">Bioko-Waldmoschusspitzmaus</vernacularName>
/ 
<emphasis id="879A2750A061870DFC35AC4C164DF8D9" bold="true" box="[952,1043,1803,1824]" pageId="73" pageNumber="468">Spanish:</emphasis>
<vernacularName id="3BED8B6CA061870DFB93AC4C1710F8D9" ID-CoL="53NBP" baseAuthorityName="Heim de Balsac" baseAuthorityYear="1968" box="[1054,1358,1803,1824]" class="Mammalia" family="Soricidae" genus="Sylvisorex" kingdom="Animalia" language="esp" order="Soricomorpha" pageId="73" pageNumber="468" phylum="Chordata" rank="species" species="isabellae">Musarana de bosque de Bioko</vernacularName>
</heading>
</paragraph>
<paragraph id="B551FB42A061870DFF24AC75117EF8BE" blockId="73.[168,1359,1736,1863]" box="[169,800,1842,1863]" pageId="73" pageNumber="468">
<heading id="EE194C2EA061870DFF24AC75117EF8BE" box="[169,800,1842,1863]" pageId="73" pageNumber="468">
<emphasis id="879A2750A061870DFF24AC7513FEF8BE" bold="true" box="[169,416,1842,1863]" pageId="73" pageNumber="468">Other common names:</emphasis>
<vernacularName id="3BED8B6CA061870DFE26AC7510DFF8BE" ID-CoL="53NBP" baseAuthorityName="Heim de Balsac" baseAuthorityYear="1968" box="[427,641,1842,1863]" class="Mammalia" family="Soricidae" genus="Sylvisorex" kingdom="Animalia" language="eng" order="Soricomorpha" pageId="73" pageNumber="468" phylum="Chordata" rank="species" species="isabellae">Isabella Forest Shrew</vernacularName>
, 
<vernacularName id="3BED8B6CA061870DFD1DAC75117EF8BE" ID-CoL="53NBP" baseAuthorityName="Heim de Balsac" baseAuthorityYear="1968" box="[656,800,1842,1863]" class="Mammalia" family="Soricidae" genus="Sylvisorex" kingdom="Animalia" language="eng" order="Soricomorpha" pageId="73" pageNumber="468" phylum="Chordata" rank="species" species="isabellae">Isabella Shrew</vernacularName>
</heading>
</paragraph>
</subSubSection>
<paragraph id="B551FB42A061870DFC83AC3D162BF7F4" blockId="73.[782,1374,1914,2337]" pageId="73" pageNumber="468">
<subSubSection id="FDF4A8C9A061870DFC83AC3D11A2F846" pageId="73" pageNumber="468" type="reference_group">
<emphasis id="879A2750A061870DFC83AC3D11F7F86E" bold="true" box="[782,937,1914,1943]" pageId="73" pageNumber="468">Taxonomy.</emphasis>
<taxonomicName id="72EE80C1A061870DFC31AC3D11A9F846" ID-CoL="53NBP" authority="Heim de Balsac, 1968" authorityName="Heim de Balsac" authorityYear="1968" class="Mammalia" family="Soricidae" genus="Sylvisorex" kingdom="Animalia" order="Soricomorpha" pageId="73" pageNumber="468" phylum="Chordata" rank="subSpecies" species="morio" subSpecies="isabellae">Sylvisorex morio isabellae Heim de Balsac, 1968</taxonomicName>
,
</subSubSection>
<subSubSection id="FDF4A8C9A061870DFB82ACE1162BF7F4" pageId="73" pageNumber="468" type="materials_examined">
<materialsCitation id="0586F11FA061870DFB82ACE1162BF7F4" ID-GBIF-Occurrence="3864799545" country="Equatorial Guinea" elevation="2000" location="Pic Santa Isabel" pageId="73" pageNumber="468" specimenCount="1">
<location id="B031AD99A061870DFB82ACE116A5F846" LSID="urn:lsid:plazi:treatment:3D474A54A061870DFF26AD8F1350F328:B031AD99A061870DFB82ACE116A5F846" box="[1039,1275,1958,1983]" country="Equatorial Guinea" name="Pic Santa Isabel" pageId="73" pageNumber="468">Pic Santa Isabel</location>
, 
<location id="B031AD99A061870DFA99ACE11106F81F" LSID="urn:lsid:plazi:treatment:3D474A54A061870DFF26AD8F1350F328:B031AD99A061870DFA99ACE11106F81F" country="Equatorial Guinea" name="Refugium" pageId="73" pageNumber="468">Refugium</location>
, 
<quantity id="721656A7A061870DFCE0AC8A1186F81F" box="[877,984,1997,2022]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="73" pageNumber="468" unit="m" value="2000.0">
<elevation id="3EC31C71A061870DFCE0AC8A1186F81F" box="[877,984,1997,2022]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="73" pageNumber="468" unit="m" value="2000.0">2000 m</elevation>
</quantity>
, 
<location id="B031AD99A061870DFC60AC8A1105F7F4" LSID="urn:lsid:plazi:treatment:3D474A54A061870DFF26AD8F1350F328:B031AD99A061870DFC60AC8A1105F7F4" country="Equatorial Guinea" name="Fernando Po (= Bioko Island)" pageId="73" pageNumber="468">Fernando Po (= Bioko Island)</location>
, 
<collectingCountry id="CDF9BBD2A061870DFCE0ACAB162FF7F4" box="[877,1137,2028,2061]" name="Equatorial Guinea" pageId="73" pageNumber="468">Equatorial Guinea</collectingCountry>
.
</materialsCitation>
</subSubSection>
</paragraph>
<subSubSection id="FDF4A8C9A061870DFC83A3541616F72B" pageId="73" pageNumber="468" type="discussion">
<paragraph id="B551FB42A061870DFC83A3541616F72B" blockId="73.[782,1374,1914,2337]" pageId="73" pageNumber="468">
There is a population in the Bamenda Highlands of south-western Cameroon that has been attributed to S. 
<taxonomicName id="72EE80C1A061870DFB4EA3211774F77A" authorityName="Heim de Balsac" authorityYear="1968" box="[1219,1322,2150,2179]" class="Mammalia" family="Soricidae" genus="Sylvisorex" kingdom="Animalia" order="Soricomorpha" pageId="73" pageNumber="468" phylum="Chordata" rank="species" species="isabellae">isabellae</taxonomicName>
, although it might represent a distinct cryptic species. Monotypic.
</paragraph>
</subSubSection>
<subSubSection id="FDF4A8C9A061870DFC83A39F1704F700" box="[782,1370,2264,2297]" pageId="73" pageNumber="468" type="distribution">
<caption id="E191ABCAA061870DFC83A39F1704F700" ID-DOI="http://doi.org/10.5281/zenodo.6871353" ID-Zenodo-Dep="6871353" box="[782,1370,2264,2297]" httpUri="https://zenodo.org/record/6871353/files/figure.png" inLine="true" pageId="73" pageNumber="468" targetBox="[167,758,1918,2332]" targetPageId="73">
<paragraph id="B551FB42A061870DFC83A39F1704F700" blockId="73.[782,1374,1914,2337]" box="[782,1370,2264,2297]" pageId="73" pageNumber="468">
<emphasis id="879A2750A061870DFC83A39F11E4F700" bold="true" box="[782,954,2264,2297]" pageId="73" pageNumber="468">Distribution.</emphasis>
N Bioko I and SW Cameroon.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="FDF4A8C9A061870DFC83A24F1686F5A2" pageId="73" pageNumber="468" type="description">
<paragraph id="B551FB42A061870DFC83A24F1686F5A2" blockId="73.[782,1374,1914,2337]" lastBlockId="73.[166,1372,2351,3281]" pageId="73" pageNumber="468">
<emphasis id="879A2750A061870DFC83A24F165BF6D8" bold="true" box="[782,1029,2312,2337]" pageId="73" pageNumber="468">Descriptive notes.</emphasis>
Head-body 54-63 mm, tail 54-59 mm, ear 8-9 mm, hindfoot 10-13 mm; weight 5-5-9 g. The Bioko Forest Shrew is small, with soft and dense pelage. Dorsum is dark reddish brown, and venter is generally slightly paler reddish brown, depending on the angle. Feet are brown, with inconspicuous dark brown scales and sparse short dark brown hairs. Tail is 90% of headbody length, blackish brown, and slightly lighter below. Braincase is high-domed, first incisors are long and hooked,first unicuspid is large while second to fourth are smaller and subequal, and third molar is medium in size. There are four unicuspids. Chromosomal complement has 2n = 36 and FN = 50 for the Cameroonian population.
</paragraph>
</subSubSection>
<subSubSection id="FDF4A8C9A061870DFF25A125107FF550" pageId="73" pageNumber="468" type="biology_ecology">
<paragraph id="B551FB42A061870DFF25A125107FF550" blockId="73.[166,1372,2351,3281]" pageId="73" pageNumber="468">
<emphasis id="879A2750A061870DFF25A1251348F57A" bold="true" box="[168,278,2658,2691]" pageId="73" pageNumber="468">Habitat.</emphasis>
Montane tropical moist forests and grasslands found above the tree line at elevations of 2000-2400 m.
</paragraph>
</subSubSection>
<subSubSection id="FDF4A8C9A061870DFF2AA1F710C5F528" box="[167,667,2736,2769]" pageId="73" pageNumber="468" type="food_feeding">
<paragraph id="B551FB42A061870DFF2AA1F710C5F528" blockId="73.[166,1372,2351,3281]" box="[167,667,2736,2769]" pageId="73" pageNumber="468">
<emphasis id="879A2750A061870DFF2AA1F713F3F528" bold="true" box="[167,429,2736,2769]" pageId="73" pageNumber="468">Food and Feeding.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection id="FDF4A8C9A061870DFF2AA1901042F501" box="[167,540,2775,2808]" pageId="73" pageNumber="468" type="breeding">
<paragraph id="B551FB42A061870DFF2AA1901042F501" blockId="73.[166,1372,2351,3281]" box="[167,540,2775,2808]" pageId="73" pageNumber="468">
<emphasis id="879A2750A061870DFF2AA1901373F501" bold="true" box="[167,301,2775,2808]" pageId="73" pageNumber="468">Breeding.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection id="FDF4A8C9A061870DFF2BA0441021F4D9" box="[166,639,2819,2848]" pageId="73" pageNumber="468" type="activity">
<paragraph id="B551FB42A061870DFF2BA0441021F4D9" blockId="73.[166,1372,2351,3281]" box="[166,639,2819,2848]" pageId="73" pageNumber="468">
<emphasis id="879A2750A061870DFF2BA04413CFF4D9" bold="true" box="[166,401,2819,2848]" pageId="73" pageNumber="468">Activity patterns.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection id="FDF4A8C9A061870DFF2AA061160CF4BE" box="[167,1106,2854,2887]" pageId="73" pageNumber="468" type="biology_ecology">
<paragraph id="B551FB42A061870DFF2AA061160CF4BE" blockId="73.[166,1372,2351,3281]" box="[167,1106,2854,2887]" pageId="73" pageNumber="468">
<emphasis id="879A2750A061870DFF2AA061113DF4BE" bold="true" box="[167,867,2854,2887]" pageId="73" pageNumber="468">Movements, Home range and Social organization.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection id="FDF4A8C9A061870DFF25A0091706F37A" pageId="73" pageNumber="468" type="conservation">
<paragraph id="B551FB42A061870DFF25A0091706F37A" blockId="73.[166,1372,2351,3281]" pageId="73" pageNumber="468">
<emphasis id="879A2750A061870DFF25A00913A5F496" bold="true" box="[168,507,2894,2927]" pageId="73" pageNumber="468">Status and Conservation.</emphasis>
Classified as Endangered on The IUCN Red List. The Bioko Forest Shrew has a very small extent of occurrence (less than 5000 km?), all individuals are in fewer than five locations, and there is continuing decline in extent and quality ofits habitat. The Bioko Forest Shrew is threatened by habitat loss mainly from logging operations and conversion of land to agricultural uses. Although it is present in Mount Oku Faunal Reserve in the Bamenda Highlands of Cameroon (although its taxonomyis uncertain), there are no conservation plans in place for the Bioko Forest Shrew, and there is urgent need to conserve suitable montane forest habitat, particularly on Bioko Island.
</paragraph>
</subSubSection>
<subSubSection id="FDF4A8C9A061870DFF2AA7D51350F328" pageId="73" pageNumber="468" type="bibRefCitation_list">
<paragraph id="B551FB42A061870DFF2AA7D51350F328" blockId="73.[166,1372,2351,3281]" pageId="73" pageNumber="468">
<emphasis id="879A2750A061870DFF2AA7D5131EF352" bold="true" box="[167,320,3218,3243]" pageId="73" pageNumber="468">Bibliography.</emphasis>
Heim de Balsac &amp; Meester (1977), Hutterer (2013ah), Hutterer &amp; Howell (2008e), Schlitter et al. (1999).
</paragraph>
</subSubSection>
</treatment>
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