The Isoperla Of California (Plecoptera: Perlodidae); Updated Male Descriptions And Adult Keys For 18 Western Nearctic Species Author Sandberg, John B. , & Boris C. Kondratieff & California Dept. of Fish & Wildlife - A. B. L. and CSUC Research Foundation, California State University, Chico, CA 95929 - 0555 U. S. A. E-mail: jsandberg @ csuchico. edu jsandberg@csuchico.edu Author Kondratieff, Boris C. Department of Bioagricultural Sciences and Pest Management, Colorado State University, Fort Collins, Colorado 80523, U. S. A. E-mail: Boris. Kondratieff @ Colostate. edu Boris.Kondratieff@Colostate.edu text Illiesia 2013 9 5 34 64 journal article 6837 10.5281/zenodo.4760870 53754ae5-8913-4e45-b546-b9edae8f726b 1854-0392 4760870 150B2555-91E3-4110-8BBC-25F1AAC0E6B2 Isoperla roguensis Szczytko & Stewart ( Figs. 14 a-d, 20 e) Isoperla roguensis Szczytko & Stewart 1984 . Holotype , Oregon , Curry County , mouth of Rogue River. Isoperla karuk Baumann & Lee 2009 . Holotype , California , Humboldt County , Klamath River @ junction of Aikens Creek . Syn. Sandberg, 2011b . Male . Aedeagus: sclerotized posterior process present; body of aedeagus with one posterior lobe, one large dorsal lobe, and one anterior lobe ( Fig. 14a ); one bulbous and usually bi-hemispherical shaped patch of spinulae above sclerotized process and a small pair of spinule patches placed laterally ( Figs. 14 a-c); sclerotized process length approaching 0.5 mm , blade-like in lateral view ( Fig. 14a ), distal margin expanded and slightly membranous in posterior view ( Fig. 14b ), shape of distal expanded margin variable, and narrow proximal portion of sclerotized process usually inverted inside posterior aedeagal membrane. Abdominal terga 8-9, 9, 9-10: without stout spinulae or long stout setae. Posterolateral margins of at least abdominal segment 8 with scale-like setae clustered in brushes of several setae. Paraprocts: curved dorsally, length if straightened subequal to combined first and second cercal segments, tapering abruptly to tiny sharp apices ( Fig. 14c ). Vesicle: rounded lobe, widest at base with broadly rounded apical margin ( Fig. 14d ). Isoperla roguensis shares similar characters with the species of the I. marmorata complex ( Table 1 ). See comments under I. fulva . Three distinct spinule patches distinguishes this species. Recently, Sandberg (2011b) placed I. karuk Baumann & Lee 2009 as a junior subjective synonym. The previously described everted anterior tubelike lobes ( Szczytko & Stewart 1984 , fig. 55) remained inverted after live eversion, their positions indicated by a pair of small inverted areas of membrane (openings). After clearing in KOH, the internal tubelike processes were visible with openings along anteroapical margin ( Fig. 20e , ventral aspect provided clearest view). Additional Material Examined. CALIFORNIA , Plumas Co. , MF Feather River, FR 12N61X access off Hwy 70, 10.5 mi ( 16.9 km ) SE Quincy , 39.86453°N , 120.76138°W , 18/ VI /2013, J.L. York , 1♂ , 1♀ ( JBSC ); Jamison Creek , 1.6 mi ( 2.6 km ) E Two Rivers exit off Hwy 70, 39.813°N , 120.6821°W , 19/ VI /2013, J.L. York , 1♀ ( JBSC ).