Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)
Author
Wesener, Thomas
urn:lsid:zoobank.org:author:
Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. & Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. & B 54 D 3623309
Author
Enghoff, Henrik
urn:lsid:zoobank.org:author:
Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark
Author
Sierwald, Petra
urn:lsid:zoobank.org:author:
Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.
text
ZooKeys
2009
2009-09-04
19
19
1
128
journal article
10.3897/zookeys.19.221
e6ea8eea-7156-4785-8313-f4bd88dadad1
1313–2970
576503
C473F9F6-1AE7-4B3F-B17F-CA1C2709010C
Hylekobolus goodmani
Wesener
,
sp. n.
urn:lsid:zoobank.org:act:
C097B4BE-0F42-4C7D-B087-4C7B4396C826
Material
examined:
6 ♂,
3 ♀
Holotype
:
1 ♂ (hind end missing),
FMMC 7936
,
Province
Toliara
, RNI
Andohahela
parcel I,
440 m
, rainforest,
24°37.6’ S
,
46°45.9’ E
, leg.
S. Goodman
,
19–28.X.1995
.
Paratypes
:
2 ♂,
1 imm.
,
FMMC 8190
,
Province
Toliara
, RNI
Andohahela
parcel I,
1200 m
, rainforest,
24°35.0’ S
,
46°44.1’ E
, leg.
S. Goodman
,
7–17.XI.1995
;
Figure 58.
Hylekobolus albicollaris
sp. n.
, male holotype.
A
head, lateral view
B
male legs 1–7, ventral view
C
telson
D
anterior gonopod, anterior view
E
left posterior gonopod, anterior view
F
anterior gonopod, posterior view
G
left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm.
Other
material examined
:
2 ♂,
1 ♀
,
FMMC 8191
, same data as previous
;
1 ♂,
2 ♀
,
FMMC 8202
, RNI
d’ Andohahela
parcel 1, 1875 m, montane rainforest,
24°33.7’ S
,
46°43.3’ E
,
pitfall traps
, leg.
S. Goodman
,
5.XII.1995
.
Differential
diagnosis:
the gonopods clearly identify this species (
Figs 59
D–G).
H. goodmani
is quite similar in colour to the sympatric
H. montanus
sp. n.
The latter, however, is much larger and has completely different posterior gonopods.
Description
.
Measurements
: male
holotype
with 49 rings,
55 mm
long,
4.7–4.9 mm
wide.
Coloration:
head, median part of collum, anterior 3/4 of body rings and anterior half of anal segment dark grey (
Fig. 59A
). Posterior margin of body rings, telson and margins of collum light brown (
Fig. 59C
). Legs light brown, antennae darker grey (
Fig. 59B
).
Eyes
with 40–44 ocelli (
Fig. 59A
).
Anterior gonopod
sternite basally wide, apically elevated into a rectangular lobe with a well-rounded tip (
Fig. 59D
). Mesal process of coxite slender and protruding (
Fig. 59D
). Apical process of telopodite large and well-rounded. Retrorse margin slender, widely projecting above telopodite margin (
Fig. 59F
), basally of apical process not projecting (
Fig. 59F
).
Figure 59.
Hylekobolus goodmani
sp. n.
, male holotype.
A
head, lateral view
B
legs 1–7, ventral view
C
telson
D
anterior gonopod, anterior view
E
left posterior gonopod, anterior view
F
anterior gonopod, posterior view
G
left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm.
Posterior gonopod
telopodite bent 90° (
Fig. 59G
). Projection absent from lateral margin (
w
,
Fig. 59E
). Apical process (
x
) basally extraordinary slender, only twice as wide as sperm canal (
Fig. 59E
). Projection of sperm canal (
z
) long and slender, running parallel to apical process (
Fig. 59G
). Tip sharp-edged. Membranous lobe (
y
) large and well-rounded (
Fig. 59E
).
Distribution
and ecology
: this species is only known from the Anosy Mountains, north of the Isaka-Ivondro corridor (
Fig. 54
).
Etymology
:
goodmani
, adjective, named after Dr. Steven M. Goodman who collected this and numerous other remarkable Malagasy millipede species.