Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda) Author Wesener, Thomas urn:lsid:zoobank.org:author: Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. & Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. & B 54 D 3623309 Author Enghoff, Henrik urn:lsid:zoobank.org:author: Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark Author Sierwald, Petra urn:lsid:zoobank.org:author: Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. text ZooKeys 2009 2009-09-04 19 19 1 128 journal article 10.3897/zookeys.19.221 e6ea8eea-7156-4785-8313-f4bd88dadad1 1313–2970 576503 C473F9F6-1AE7-4B3F-B17F-CA1C2709010C Hylekobolus goodmani Wesener , sp. n. urn:lsid:zoobank.org:act: C097B4BE-0F42-4C7D-B087-4C7B4396C826 Material examined: 6 ♂, 3 ♀ Holotype : 1 ♂ (hind end missing), FMMC 7936 , Province Toliara , RNI Andohahela parcel I, 440 m , rainforest, 24°37.6’ S , 46°45.9’ E , leg. S. Goodman , 19–28.X.1995 . Paratypes : 2 ♂, 1 imm. , FMMC 8190 , Province Toliara , RNI Andohahela parcel I, 1200 m , rainforest, 24°35.0’ S , 46°44.1’ E , leg. S. Goodman , 7–17.XI.1995 ; Figure 58. Hylekobolus albicollaris sp. n. , male holotype. A head, lateral view B male legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm. Other material examined : 2 ♂, 1 ♀ , FMMC 8191 , same data as previous ; 1 ♂, 2 ♀ , FMMC 8202 , RNI d’ Andohahela parcel 1, 1875 m, montane rainforest, 24°33.7’ S , 46°43.3’ E , pitfall traps , leg. S. Goodman , 5.XII.1995 . Differential diagnosis: the gonopods clearly identify this species ( Figs 59 D–G). H. goodmani is quite similar in colour to the sympatric H. montanus sp. n. The latter, however, is much larger and has completely different posterior gonopods. Description . Measurements : male holotype with 49 rings, 55 mm long, 4.7–4.9 mm wide. Coloration: head, median part of collum, anterior 3/4 of body rings and anterior half of anal segment dark grey ( Fig. 59A ). Posterior margin of body rings, telson and margins of collum light brown ( Fig. 59C ). Legs light brown, antennae darker grey ( Fig. 59B ). Eyes with 40–44 ocelli ( Fig. 59A ). Anterior gonopod sternite basally wide, apically elevated into a rectangular lobe with a well-rounded tip ( Fig. 59D ). Mesal process of coxite slender and protruding ( Fig. 59D ). Apical process of telopodite large and well-rounded. Retrorse margin slender, widely projecting above telopodite margin ( Fig. 59F ), basally of apical process not projecting ( Fig. 59F ). Figure 59. Hylekobolus goodmani sp. n. , male holotype. A head, lateral view B legs 1–7, ventral view C telson D anterior gonopod, anterior view E left posterior gonopod, anterior view F anterior gonopod, posterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; Gn = gnathochilarium; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm. Posterior gonopod telopodite bent 90° ( Fig. 59G ). Projection absent from lateral margin ( w , Fig. 59E ). Apical process ( x ) basally extraordinary slender, only twice as wide as sperm canal ( Fig. 59E ). Projection of sperm canal ( z ) long and slender, running parallel to apical process ( Fig. 59G ). Tip sharp-edged. Membranous lobe ( y ) large and well-rounded ( Fig. 59E ). Distribution and ecology : this species is only known from the Anosy Mountains, north of the Isaka-Ivondro corridor ( Fig. 54 ). Etymology : goodmani , adjective, named after Dr. Steven M. Goodman who collected this and numerous other remarkable Malagasy millipede species.