A revision of the genus Bromeloecia Spuler (Diptera: Sphaeroceridae: Limosininae)
Author
Yau, Tiffany
Author
Marshall, Stephen A.
text
Zootaxa
2018
2018-07-13
4445
1
1
115
journal article
29191
10.11646/zootaxa.4445.1.1
0ce95227-0383-471e-9cac-4cab7c0e34f0
1175-5326
1454109
95850B7B-4E31-4C9D-89BA-EE65BDB79BDD
Bromeloecia ponsa
Yau and Marshall
sp. n.
(Figs. 10.1–10.14)
Description:
Body length
1.1–1.5 mm
(males),
1. 1–1.6 mm
(females).
Head: Length of anterior fronto-orbital seta 0.6x posterior seta. Flagellomere 1 with 1 long pit surrounding 5/6 of basal outer edge. Eye ovoid to almost circular; genal height 0.6x eye height.
Thorax: Katepisternum with a row of 3–4 smaller anterior setae with dorsal most seta 0.5x and almost level with strong dorsal seta.
Wing:
Costa
ends at 3/4 of wing length. Angle between costa and R4+5 90°. Distance between radial fork and rm
2x
length of r-m; dm-cu equal in length to r-m and distance between crossveins no more than length of dm-cu. M-pseudovein slightly bent towards costa, bent posteriorly to meet level of basal portion in distal one third and ending close to wing margin.
WIPs: Ripple pattern of cell r2+3 distinct, main length of pattern generally parallel to R2+3 and costa, distally dipped, narrowest cross section at mid length of CS3. Basal angle of distal spot wide, 125°.
Male abdomen and terminalia: Syntergite 1+2 and T3 with long lateral setae, a distinctly long pair of divergent medial setae and multiple short marginal setae in between. Anteroventral arm of synsternite 6+7 similar to a laterally stretched out Ω-shape; posteroventral lobe long and curled inwards to abdominal cavity, with anteroventral edge sclerotized. Right side of epandrium without anteroventral process. Surstylus ventrally covered with mollisetae.
Hypandrium Y-shaped, lateral extension plate starts at distal half of posteromedial hypandrial processes; hypandrial arms round. Ejaculatory apodeme finger-like, ejaculatory bulb simple. Epiphallus strongly bent ventrally. Distiphallus LFS slender; FDS folded, apex reaches back to base of WS; second distal SDS shoe-hornshaped.
Female abdomen and terminalia: T6 as wide as T5, medially with a pair of widely separated setae. Each half of T8 triangular, with 1 large and 5 smaller setae. S8 rectangular. Epiproct glossy on lateral quarters. Cercus diamondshaped. Each spermatheca with 1 large apical and 1 small basal thimble-shaped pit.
Type
specimens:
Holotype
:
♂
:
ECUADOR
:
Napo
:
Lago Papallacta
, 00°20ʹ29ʺS, 078°10ʹ23ʺW, forest above lake,
3400 m
, pans/dung,
4–8 Nov 1999
,
S.A. Marshall
, debu00135739 (
QCAZ
).
Paratypes
:
ARGENTINA
:
Salta
:
Campo Quijano
, FIT,
28 Feb 1992
,
S. A. Marshall
(
1♀
,
DEBU
)
;
30 km
E
Salta
Campo Quijano, forest remnant,
dung trap
,
18–20 Feb 1992
,
S. A. Marshall
(
10♂
,
12♀
,
DEBU
)
;
30 km
E
Salta
Campo Quijano, El Alisal, forest remnant, FIT,
18–20 Feb 1992
,
S. A. Marshall
(
4♂
,
2♀
,
DEBU
)
;
La Cladera
, forest,
1500 m
, sweep roadside,
27 Feb 1992
,
S. A. Marshall
(
1♂
,
DEBU
)
;
22 km
N
La Cladera
,
El Ucumar
, subtropical humid forest,
1550 m
,
Malaise
FIT,
2–30 Dec 1987
,
S. & J. Peck
(
1♂
,
6♀
,
DEBU
)
;
El Maray
,
50 km
W
Chicoana
, dry,
2500 m
,
pan traps
,
18–28 Feb 1992
,
S. A. Marshall
(
3♂
,
4♀
,
DEBU
)
.
BOLIVIA
:
La Paz
:
Zongo
road, 16°10ʹ04ʺS, 068°07ʹ49ʺW, alpine shrub, ~
3800 m
, on cow dung,
18 Apr 2001
,
S. A. Marshall
(
3♂
,
1♀
,
CBFC
,
DEBU
)
;
Zongo Valley
, 16°7ʹ57ʺS, 068°06ʹ56ʺW, forest,
2600 m
,
pan traps
,
18 Apr 2001
,
S. A. Marshall
(
1♀
,
DEBU
)
.
ECUADOR
:
Carchi
:
Guandera For. Res.
,
15 km
E
San Gabriel
, 00°36ʹ00ʺN, 077°41ʹ35ʺW,
3400 m
, sweeping forest trail,
1 Nov 1999
,
S. A. Marshall
(
1♂
,
DEBU
)
;
Napo
:
27 km
NW
Baeza
,
2700 m
,
dung trap
,
2–6 Mar 1976
,
S. Peck
(
9♂
,
12♀
,
QCAZ
,
DEBU
)
;
42 km
NW
Baeza
,
3300 m
,
carrion trap
,
2–6 Mar 1976
,
S. Peck
(
3♂
,
3♀
,
QCAZ
,
DEBU
)
; same as holotype (1♂, DEBU).
FIGURES 10.1–10.5.
Bromeloecia ponsa
sp. n.
, male habitus. 10.1, lateral (Ecuador); 10.2, dorsal (Argentina); 10.3a, head (Argentina); 10.3b, left flagellomere 1 posterior; 10.4, wing against white background (Argentina); 10.5, wing displaying WIP reflection against black background (Argentina).
FIGURES 10.6–10.10.
Bromeloecia ponsa
sp. n.
, abdominal structures (male: Ecuador and Bolivia). 10.6, terminalia left ventrolateral; 10.7, terminalia ventral; 10.8, epandrium posterior; 10.9, phallus and associated structures left ventrolateral; 10.10, phallus and associated structures left posterolateral.
FIGURES 10.11–10.14.
Bromeloecia ponsa
sp. n.
, abdominal structures (female: Argentina). 10.11, terminalia dorsal; 10.12, terminalia ventral; 10.13, spermathecae and spermathecal ducts; 10.14, terminalia lateral.
Etymology:
The species name refers to the strongly arched, bridge-shaped anteroventral arm of synsternite 6+7 (Latin ‘pons’: bridge).
Comments:
B. ponsa
, like
B. balaena
, is found in the high elevations of the Andes, however its distribution stretches further south, to northern
Argentina
.