A revision of the genus Bromeloecia Spuler (Diptera: Sphaeroceridae: Limosininae) Author Yau, Tiffany Author Marshall, Stephen A. text Zootaxa 2018 2018-07-13 4445 1 1 115 journal article 29191 10.11646/zootaxa.4445.1.1 0ce95227-0383-471e-9cac-4cab7c0e34f0 1175-5326 1454109 95850B7B-4E31-4C9D-89BA-EE65BDB79BDD Bromeloecia ponsa Yau and Marshall sp. n. (Figs. 10.1–10.14) Description: Body length 1.1–1.5 mm (males), 1. 1–1.6 mm (females). Head: Length of anterior fronto-orbital seta 0.6x posterior seta. Flagellomere 1 with 1 long pit surrounding 5/6 of basal outer edge. Eye ovoid to almost circular; genal height 0.6x eye height. Thorax: Katepisternum with a row of 3–4 smaller anterior setae with dorsal most seta 0.5x and almost level with strong dorsal seta. Wing: Costa ends at 3/4 of wing length. Angle between costa and R4+5 90°. Distance between radial fork and rm 2x length of r-m; dm-cu equal in length to r-m and distance between crossveins no more than length of dm-cu. M-pseudovein slightly bent towards costa, bent posteriorly to meet level of basal portion in distal one third and ending close to wing margin. WIPs: Ripple pattern of cell r2+3 distinct, main length of pattern generally parallel to R2+3 and costa, distally dipped, narrowest cross section at mid length of CS3. Basal angle of distal spot wide, 125°. Male abdomen and terminalia: Syntergite 1+2 and T3 with long lateral setae, a distinctly long pair of divergent medial setae and multiple short marginal setae in between. Anteroventral arm of synsternite 6+7 similar to a laterally stretched out Ω-shape; posteroventral lobe long and curled inwards to abdominal cavity, with anteroventral edge sclerotized. Right side of epandrium without anteroventral process. Surstylus ventrally covered with mollisetae. Hypandrium Y-shaped, lateral extension plate starts at distal half of posteromedial hypandrial processes; hypandrial arms round. Ejaculatory apodeme finger-like, ejaculatory bulb simple. Epiphallus strongly bent ventrally. Distiphallus LFS slender; FDS folded, apex reaches back to base of WS; second distal SDS shoe-hornshaped. Female abdomen and terminalia: T6 as wide as T5, medially with a pair of widely separated setae. Each half of T8 triangular, with 1 large and 5 smaller setae. S8 rectangular. Epiproct glossy on lateral quarters. Cercus diamondshaped. Each spermatheca with 1 large apical and 1 small basal thimble-shaped pit. Type specimens: Holotype : : ECUADOR : Napo : Lago Papallacta , 00°20ʹ29ʺS, 078°10ʹ23ʺW, forest above lake, 3400 m , pans/dung, 4–8 Nov 1999 , S.A. Marshall , debu00135739 ( QCAZ ). Paratypes : ARGENTINA : Salta : Campo Quijano , FIT, 28 Feb 1992 , S. A. Marshall ( 1♀ , DEBU ) ; 30 km E Salta Campo Quijano, forest remnant, dung trap , 18–20 Feb 1992 , S. A. Marshall ( 10♂ , 12♀ , DEBU ) ; 30 km E Salta Campo Quijano, El Alisal, forest remnant, FIT, 18–20 Feb 1992 , S. A. Marshall ( 4♂ , 2♀ , DEBU ) ; La Cladera , forest, 1500 m , sweep roadside, 27 Feb 1992 , S. A. Marshall ( 1♂ , DEBU ) ; 22 km N La Cladera , El Ucumar , subtropical humid forest, 1550 m , Malaise FIT, 2–30 Dec 1987 , S. & J. Peck ( 1♂ , 6♀ , DEBU ) ; El Maray , 50 km W Chicoana , dry, 2500 m , pan traps , 18–28 Feb 1992 , S. A. Marshall ( 3♂ , 4♀ , DEBU ) . BOLIVIA : La Paz : Zongo road, 16°10ʹ04ʺS, 068°07ʹ49ʺW, alpine shrub, ~ 3800 m , on cow dung, 18 Apr 2001 , S. A. Marshall ( 3♂ , 1♀ , CBFC , DEBU ) ; Zongo Valley , 16°7ʹ57ʺS, 068°06ʹ56ʺW, forest, 2600 m , pan traps , 18 Apr 2001 , S. A. Marshall ( 1♀ , DEBU ) . ECUADOR : Carchi : Guandera For. Res. , 15 km E San Gabriel , 00°36ʹ00ʺN, 077°41ʹ35ʺW, 3400 m , sweeping forest trail, 1 Nov 1999 , S. A. Marshall ( 1♂ , DEBU ) ; Napo : 27 km NW Baeza , 2700 m , dung trap , 2–6 Mar 1976 , S. Peck ( 9♂ , 12♀ , QCAZ , DEBU ) ; 42 km NW Baeza , 3300 m , carrion trap , 2–6 Mar 1976 , S. Peck ( 3♂ , 3♀ , QCAZ , DEBU ) ; same as holotype (1♂, DEBU). FIGURES 10.1–10.5. Bromeloecia ponsa sp. n. , male habitus. 10.1, lateral (Ecuador); 10.2, dorsal (Argentina); 10.3a, head (Argentina); 10.3b, left flagellomere 1 posterior; 10.4, wing against white background (Argentina); 10.5, wing displaying WIP reflection against black background (Argentina). FIGURES 10.6–10.10. Bromeloecia ponsa sp. n. , abdominal structures (male: Ecuador and Bolivia). 10.6, terminalia left ventrolateral; 10.7, terminalia ventral; 10.8, epandrium posterior; 10.9, phallus and associated structures left ventrolateral; 10.10, phallus and associated structures left posterolateral. FIGURES 10.11–10.14. Bromeloecia ponsa sp. n. , abdominal structures (female: Argentina). 10.11, terminalia dorsal; 10.12, terminalia ventral; 10.13, spermathecae and spermathecal ducts; 10.14, terminalia lateral. Etymology: The species name refers to the strongly arched, bridge-shaped anteroventral arm of synsternite 6+7 (Latin ‘pons’: bridge). Comments: B. ponsa , like B. balaena , is found in the high elevations of the Andes, however its distribution stretches further south, to northern Argentina .