World reclassification of the Gonatocerus group of genera (Hymenoptera: Mymaridae)
Author
Huber, John T.
text
Zootaxa
2015
3967
1
1
184
journal article
40912
10.11646/zootaxa.3967.1.1
17446e98-b166-4d8d-aa19-c2da27aa7b3d
1175-5326
288712
809A05D1-3BAD-4A32-8D56-C91A6B609D00
Cosmocomoidea
Howard
, stat. rev. (
=
ater
species group, of authors)
(
Figs 3–5
,
9–38
,
41–60
)
Cosmocomoidea
Howard, 1908
: 69
; Girault, 1929 [428]: 21 (implied synonymy under
Ooctonus
);
Ogloblin, 1959a
: 44
(subgenus under
Lymaenon
);
De
Santis, 1979
: 367
(subgenus under
Gonatocerus
);
Matthews, 1986
: 220
(species group [the
ater
group] within
Gonatocerus
); Triapitsyn
et al
., 2010: 94 (subgenus under
Gonatocerus
), 95 (key to Neotropical species);
Triapitsyn, 2013a
: 117
(subgenus under
Gonatocerus
), 118 (key to Palaearctic species),
Triapitsyn, 2013b
: 213
(key to Nearctic species).
Type
species
.
Cosmocomoidea morrilli
Howard
, by monotypy.
Diagnosis
. Within Gonatocerini, the species of
Cosmocomoidea
are distinguished by the following combination of features: subantennal sulci strongly converging and close together at mouth margin (
Fig. 43
); ocellar triangle with 2 setae (
Fig. 43
); pronutm with lobes abutting (
Fig. 49
); dorsellum rhomboidal, with anterior and posterior margins not parallel (
Fig. 49
); fore wing with microtrichia behind venation usually absent (
Fig. 19
) but, if present, only on one surface (
Fig. 20
) or less dense than beyond venation; propodeum with two submedian carinae (
Figs 17
,
49
).
Description
.
FEMALE
. Face with subantennal sulci present, strongly converging and close together, almost touching ventrally at mouth margin, the distance between them at mouth margin much less than their distance from preorbital sulcus (
Figs 10
,
14
a, 43, 48). Vertex with 2 setae between lateral ocelli (
Figs 9
,
43, 46
). Occiput entire, without a transverse suture dividing it into upper and lower portions (
Figs 11
,
14
e, 45) but in the
bucculentus
subgroup of
Huber (1988: 49)
and Triapitsyn
et al
. (2010: 95) with a circular or oval sulcus constricting the gena behind and below eye (
Figs 5
c, 46, 48). Funicle 8-segmented (
Figs 3
,
12, 13
,
15
a, b, 41, 42), the basal four segments often as long as apical four and at least one of the basal four with mps (apical four segments always have mps on some segments). Pronotum longitudinally divided, the lobes abutting medially (
Figs 5
,
9
,
17
,
49, 53
). Dorsellum rhomboidal, slightly to considerably longer medially than laterally, with anterior and posterior margins clearly not parallel (
Figs 5
,
17, 18
,
49
), rarely narrow and almost parallel. Pronotal spiracle about the same diameter as propodeal spiracle (
Figs 50, 52
). Propodeum with one pair of submedian carinae varying considerably in shape, thickness, length, and distance apart (
Figs 5
,
17
,
49, 54
), the carinae sometimes very short and/or faint and sometimes (
morrilli
subgroup of Triapitsyn
et al.
2010) with transverse wrinkles or carinae lateral to the submedian carinae (
Figs 49, 50
). Fore wing relatively wide, usually without microtrichia behind parastigma (
Figs 3
,
19
,
38
) but, if present, the microtrichia usually not as dense or as uniformly distributed as beyond venation (
Fig. 41
), except on dorsal surface in the
chusqueicolus
subgroup (Triapitsyn
et al
. 2010: 94) (
Figs 20
a, 42). Stigmal vein with apex oblique (
Figs 3
,
38
,
41
,
42
).
MALE
. Radicle short to very short, not or barely separated from scape by a faint line or sulcus. Scape short, at most about 3× as long as often small pedicel. Flagellum usually with relatively wide flagellomeres, sometimes with fl1 almost wide as long. Genitalia with apodeme of genital sternum longer than aedeagus; aedeagal apodemes not fused at junction with aedeagus and almost as long as apodeme of genital sternum; genitalia with fairly wide scooplike lateral projections (volsellae?) lateral to base of aedeagal apodemes (see examples in Triapitsyn 2010).
Discussion
. With almost 100 nominal species,
Cosmocomoidea
is the second most speciose genus in Gonatocerini. Specimens of most species are relatively large and often have the fore wing with faint to pronounced colour band or bands but some are small and superficially resemble some
Lymaenon
species.
Triapitsyn
et al
. (2010) divided the Neotropical
Cosmocomoidea
(treated as a subgenus of
Gonatocerus
) into two species groups, the
ater
and
masneri
groups. They subdivided the
ater
group into the
ater
,
bucculentus
,
chusqueicolus
, and
morrilli
subgroups. The two species they placed in the
masneri
group are transferred to
Yoshimotoana
, described below.
It was unfortunate that
Matthews (1986)
and
Huber (1988)
named the species here placed in
Cosmocomoidea
as the
ater
-group of
Gonatocerus
.
As
Triapitsyn (2013a)
explained, the
type
series of
C
.
ater
(Förster)
consists of four specimens representing at least three species belonging, in his classification, to two different subgenera,
G
. (
Cosmocomoidea
) and
G
. (
Lymaenon
). Triapitsyn upheld the
type
fixation by
Matthews (1986)
but noted it is a
lectotype
, not a
holotype
; this female definitely belongs to
Cosmocomoidea
. One male has a strap-like dorsellum and two rows of microtrichia behind the parastigma and definitely belongs to
Lymaenon
. A third specimen, a female, belongs to a second species of
Cosmocomoidea
. Förster’s fourth specimen is a male of uncertain identity. All four specimens are part of Förster’s original material, upon which he based his description and redescription of
ater
(
Förster 1841
;
Foerster 1847
—Foerster stated the number of specimens in this paper).
The gender of
Cosmocomoidea
is feminine (
Huber 2002
). Because almost all the included species were originally described in
Gonatocerus
or
Lymaenon
, which are masculine, the names of the species listed under
Cosmocomoidea
have been changed to agree in gender, where necessary. In some cases it was difficult to determine if the name is an adjective or a noun in apposition. When in doubt I assumed, in the absence of other evidence, that the name was a noun in apposition and therefore the ending remains unchanged from the original spelling.
Distribution
.
Cosmocomoidea
species occur in the Nearctic, Neotropical, Palaearctic and Oriental regions. In the latter region
Cosmocomoidea
species occur as far east as the
Philippines
. The genus is absent from the Ethiopian and Australian regions except for one Nearctic species deliberately introduced for biological control into
French Polynesia
(
Grandgirard
et al.
2008
), two other species accidentally introduced into the Hawaiian Islands (
Huber & Beardsley 2000
), and
C
.
novifasciatus
(Girault)
accidentally introduced into both the Hawaiian Islands and
Guam
(new records).
The greatest number and diversity of species occur in the Neotropical region. The Old World has relatively few species. In the Palaearctic region most species are dark brown or black and, especially in Europe, similar to each other but in the Oriental region they are usually more colourful and diverse.
Hosts and habitat
.
Cicadellidae (Proconini)
. A few species have been studied in detail because of their importance in biological control of the glassy-winged sharpshooter,
Homalodisca vitripennis
(Germar) (Cicadellidae)
(Triapitsyn 2006,
Grandgirard
et al
. 2008
). The greatest diversity of species is in forested habitats.
Included species:
Cosmocomoidea abbreviata
(Ogloblin)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Lymaenon abbreviatus
Ogloblin 1953
: 7
;
holotype
♀ in MLPA.
De
Santis in
De
Santis & Esquivel, 1967
: 50 [transferred to
Gonatocerus
(
Gonatocerus
)
]; Triapitsyn
et al
., 2010: 99 [(transferred to
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Argentina
, Misiones, Loreto.
Cosmocomoidea annulicornis
(Ogloblin)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Lymaenon annulicornis
Ogloblin, 1936
: 41
;
lectotype
♀ in MLPA (examined).
Lymaenon
(
Cosmocomoidea
)
annulicornis
Ogloblin, 1959a
: 50
(assigned to subgenus);
De
Santis, 1967: 10, 106 [(transferred to
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Argentina
, Misiones, Loreto.
Cosmocomoidea ashmeadi
(Girault)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Lymaenon dolichocerus ashmeadi
Girault, 1915
[261]: 8;
lectotype
♀ in USNM (examined). Girault, 1929 [428]: 25 (given species status); Triapitsyn
et al
., 2010: 103 [(placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
USA
, Texas, no locality given.
Cosmocomoidea atra
(Förster)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus ater
Förster, 1841
: 45
;
lectotype
♀ in NHMW (examined) (fixed by
Triapitsyn, 2013a
: 119
[incorrectly designated as
holotype
by
Matthews 1986
: 221
]).
Foerster, 1847
: 206
(transferred to
Rachistus
);
Dalla Torre, 1898
: 429
(transferred to
Gonatocerus
); Triapitsyn
et al
., 2010: 103 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Germany
, Aachen.
Cosmocomoidea empoascae
(Subba Rao)
.
Lymaenon empoascae
Subba Rao, 1966
: 195
;
holotype
♀ in IARI. Subba Rao, 1983: 135 (transferred to
Gonatocerus
).
TL:
India
, Punjab, Jullundur. Synonymy under
G
.
ater
by
Zeya & Hayat, 1995
: 70
.
Cosmocomoidea indica
(Subba Rao & Kaur)
.
Lymaenon indicus
Subba Rao & Kaur, 1959
: 229
;
holotype
♀ in IARI. Subba Rao 1983: 135 (transferred to
Gonatocerus
).
TL:
India
, New Delhi. Synonymy under
G
.
ater
by
Zeya & Hayat, 1995
: 70
.
Cosmocomoidea intermedia
(Boţoc)
.
Lymaenon intermedius
Boţoc, 1962
: 108
;
holotype
♀ in Boţoc collection (lost).
TL:
Romania
, Cluj-Napoca, Someş river floodplain. Synonymy under
G
.
ater
by
Matthews, 1986
: 222
; treated as valid species by
Pricop, 2010
: 113
. Synonymy under
G
.
ater
s.
l
.
by
Triapitsyn, 2013a
: 119
, but see his discussion on p. 129.
Cosmocomoidea nigroides
(Narayanan & Subba Rao)
.
Lymaenon nigroides
Narayanan & Subba Rao, 1961
: 656
;
holotype
♀ in IARI. Subba Rao, 1983: 136 (as
nigriodes
[sic], transferred to
Gonatocerus
).
TL:
India
, Delhi. Synonymy under
G
.
ater
by
Zeya & Hayat, 1995
: 70
.
Cosmocomoidea pannonica
(Soyka)
.
Gonatocerus pannonicus
Soyka, 1946
: 39
;
holotype
♀ in NHMW.
TL:
Austria
, Hundsheim. Synonymy under
G
.
ater
by
Matthews, 1986
: 221
and under
G
.
ater
,
s
.
l
.
by
Triapitsyn, 2013a
: 119
.
Cosmocomoidea populi
(Viggiani)
.
Lymaenon populi
Viggiani, 1969
: 40
;
holotype
♀ in DEZA, but see
Triapitsyn, 2013a
: 122
.
Graham, 1973
: 48
(transferred to
Gonatocerus
).
TL:
Italy
, Rome. Synonymy under
G
.
ater
by
Matthews, 1986
: 222
; synonymy accepted by
Triapitsyn, 2013a
: 119
, but see his discussion on p. 133.
Cosmocomoidea schmitzi
(Debauche)
.
Lymaenon schmitzi
Debauche, 1948
: 86
;
holotype
♀ in IRSNB.
Matthews, 1986
: 221
(transferred to
Gonatocerus
).
TL:
Belgium
, Héverlé. Synonymy under
G
.
ater
by
Matthews, 1986
: 221
.
Cosmocomoidea atriclava
(Girault)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus triguttatus atriclavus
Girault, 1917
[316]: 19;
holotype
♀ in USNM (examined).
Triapitsyn
et al
., 2002
: 35 [placed in
Gonatocerus
(
Cosmocomoidea
)
and given species status)].
TL:
Trinidad and Tobago
,
Trinidad
, Mitan.
Cosmocomoidea barbos
(Triapitsyn)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus
(
Cosmocomoidea
)
barbos
Triapitsyn
in Triapitsyn
et al
., 2010: 111;
holotype
♀ in UCRC.
TL:
Mexico
, Veracruz, Tuxpan.
Cosmocomoidea blefuscu
(Triapitsyn)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus
(
Cosmocomoidea
)
blefuscu
Triapitsyn
in Triapitsyn
et al
., 2010: 114;
holotype
♀ in UCRC.
TL:
Costa Rica
, Cartago, Parque Nacional Tapantí.
Cosmocomoidea blesticus
(Ogloblin)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Lymaenon blesticus
Ogloblin, 1957
: 39
;
holotype
♀ in MLPA.
De
Santis, 1967: 10, 103 (as
biesticus
[
sic
], transferred to
Gonatocerus
); Triapitsyn
et al
., 2010: 116 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Argentina
, Puerto Blest, Lago Nahuel Huapí, Río Negro.
Cosmocomoidea bonariensis
(Brèthes)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Foersterella bonariensis
Brèthes, 1922
: 129
;
lectotype
♀ in MACN.
De
Santis, 1967: 10, 103 (transferred to
Gonatocerus
); Triapitsyn
et al
., 2010: 118 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Argentina
, Buenos Aires, General Urquiza.
Cosmocomoidea enicmophila
(Huber)
.
Gonatocerus enicmophilus
Huber, 1988
: 79
;
holotype
♀ in USNM (examined).
TL:
USA
, California, Riverside Co., Riverside, UCR Campus. Synonymy under
G
.
bonariensis
by Triapitsyn
et al
., 2010: 118.
Cosmocomoidea necator
(Ogloblin)
.
Lymaenon necator
Ogloblin, 1939
: 241
;
De
Santis in
De
Santis & Esquivel, 1967
: 50 (transferred to
Gonatocerus
);
lectotype
♀ in MLPA (examined).
TL:
Argentina
, Buenos Aires, José C. Paz. Synonymy under
G
.
bonariensis
by Triapitsyn
et al
., 2010: 118.
Cosmocomoidea brachyura
(Ogloblin)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Lymaenon brachyurus
Ogloblin, 1938a
: 32
;
holotype
♀ in MLPA (examined).
De
Santis, 1967: 10, 103 (transferred to
Gonatocerus
); Triapitsyn
et al
., 2010: 122 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Argentina
: Buenos Aires, Tigre.
Cosmocomoidea bucculenta
(Huber)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus bucculentus
Huber, 1988
: 76
;
holotype
♀ in CNC (examined). Triapitsyn
et al
., 2010: 125 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
USA
, Arizona, Cochise Co.,
5 mi
. W. Portal, Southwestern Research Station.
Cosmocomoidea capitata
(Gahan)
,
comb. n.
from
Gonatocerus
.
Gonatoceros
[
sic
]
capitatus
Gahan, 1932
: 754
;
lectotype
♀ in USNM (examined).
TL:
USA
, Utah, Centerville.
Cosmocomoidea carahuensis
(Ogloblin)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Lymaenon carahuensis
Ogloblin, 1957
: 36
;
holotype
♀ in MLPA (examined).
De
Santis, 1967: 10, 104 (transferred to
Gonatocerus
); Triapitsyn
et al
., 2010: 127 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Argentina
: Neuquén, Pucará, Lago Lacar.
Cosmocomoidea caudata
(Ogloblin)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus caudatus
Ogloblin 1935
: 74
;
lectotype
♀ in MLPA (examined). Triapitsyn
et al
., 2010: 131 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Argentina
, Misiones, Loreto.
Cosmocomoidea chusqueicola
(Ogloblin)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Lymaenon chusqueicolus
Ogloblin, 1957
: 33
;
holotype
♀ in MLPA.
De
Santis, 1967: 10, 104 (transferred to
Gonatocerus
); Triapitsyn
et al
., 2010: 135 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Argentina
: Neuquén, Pucará, Lago Lacar.
Cosmocomoidea concinna
(Ogloblin)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Lymaenon concinnus
Ogloblin, 1936
: 46
;
lectotype
♀ in MLPA (examined).
De
Santis, 1967: 11, 104 (transferred to
Gonatocerus
); Triapitsyn
et al
., 2010: 137 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Argentina
: Misiones, Loreto.
Cosmocomoidea coxalis
(Ogloblin)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Lymaenon
(
Cosmocomoidea
)
coxalis
Ogloblin, 1959a
: 51
;
holotype
♀ in MLPA (examined).
De
Santis, 1979
: 368 (transferred to
Gonatocerus
[
Cosmocomoidea
]).
TL:
Ecuador
, Esmeraldas, San Mateo.
Cosmocomoidea crassicornis
(Viggiani)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Lymaenon crassicornis
Viggiani, 1969
: 44
;
holotype
♂ in DEZA.
Triapitsyn, 2013a
: 137
[placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Russia
, Adzarskoj RSSA, Sosna (but see
Triapitsyn, 2013a
: 137
for correct locality).
Cosmocomoidea cuscus
(Triapitsyn)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus
(
Cosmocomoidea
)
cuscus
Triapitsyn
in Triapitsyn
et al
., 2010: 141;
holotype
♀ in UCRC (examined).
TL:
Peru
, Cuzco, Picol,
3728 m
.
Cosmocomoidea deleoni
(Triapitsyn, Logarzo & Virla)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus deleoni
Triapitsyn, Logarzo & Virla
, in
Triapitsyn
et al
., 2008
: 5;
holotype
♀ in MPLA.
TL:
Argentina
, Buenos Aires.
Cosmocomoidea dolichocerus
(Ashmead)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatoceru
s
dolichocerus
Ashmead, 1887
: 192
;
holotype
♀ in USNM (examined).
Triapitsyn, 2013a
: 145 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
USA
, Florida, Duval Co., Jacksonville.
Cosmocomoidea elizavetae
(Triapitsyn)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Cosmocomoidea elizavetae
Triapitsyn, 2013b
: 214
;
holotype
♀ in USNM.
TL:
USA
, Illinois, Elizabethtown.
Cosmocomoidea ermi
(Triapitsyn)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Cosmocomoidea ermi
Triapitsyn, 2013b
: 215
;
holotype
♀ in UCRC.
TL:
USA
, California, Riverside Co., near Temecula, Sandia Creek near
De
Luz road.
Cosmocomoidea fasciata
(Girault)
,
comb. n.
from
Gonatocerus
.
Gonatocerus fasciatus
Girault, 1911
[96]: 265;
lectotype
♀ in USNM (examined).
TL:
USA
, Virginia, Arlington.
Cosmocomoidea flagellaris
(Ogloblin)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Lymaenon flagellaris
Ogloblin, 1959a
: 61
;
holotype
♂ in MLPA (examined).
De
Santis, 1979
: 12, 365 (transferred to
Gonatocerus
); Triapitsyn
et al
., 2010: 147 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Ecuador
, Pichincha, vicinity of Quito.
Cosmocomoidea flagellata
(Huber)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus flagellatus
Huber, 1988
: 78
;
holotype
♀ in USNM (examined). Triapitsyn
et al
., 2010: 148 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
USA
, Texas, Weslaco.
Cosmocomoidea garchamp
(Triapitsyn)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus
(
Cosmocomoidea
)
garchamp
Triapitsyn
in Triapitsyn
et al
., 2010: 150;
holotype
♀ in MPLA (examined).
TL:
Argentina
, Misiones, Loreto.
Cosmocomoidea garyi
(Manickavasagam & Rameshkumar)
,
comb. n.
from
Gonatocerus
.
Gonatocerus garyi
Manickavasagam & Rameshkumar, 2013a
: 89
;
holotype
♀ in EDAU.
TL:
India
, Karnataka, Bengaluru, IWST campus.
Cosmocomoidea gerasim
(Triapitsyn)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus
(
Cosmocomoidea
)
gerasim
Triapitsyn
in Triapitsyn
et al
., 2010: 153;
holotype
♀ in UCRC (examined).
TL:
Mexico
, Tamaulipas, Gómez Farías, Estación Los Cedros,
340 m
.
Cosmocomoidea gracilicornis
(Ogloblin)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Lymaenon gracilicornis
Ogloblin, 1936
: 50
;
lectotype
♀ in MLPA (examined).
De
Santis, 1967: 10, 104 (transferred to
Gonatocerus
); Triapitsyn
et al
., 2010: 154 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Argentina
, Misiones, Loreto.
Cosmocomoidea grandis
(Ogloblin)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Lymaenon grandis
Ogloblin, 1936
: 38
;
lectotype
♀ in MLPA (examined).
De
Santis, 1967: 10, 104 (transferred to
Gonatocerus
); Triapitsyn
et al
., 2010: 157 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Argentina
: Misiones, Loreto.
†
Cosmocomoidea greenwalti
(Huber)
,
comb. n.
from
Gonatocerus
.
Gonatocerus greenwalti
Huber
in
Huber & Greenwalt, 2011
: 486
;
holotype
♀ in USNM (examined).
TL:
USA
, Montana, Flathead River, Kishenehn Formation.
Cosmocomoidea gunathilagaraji
(Manickavasagam & Rameshkumar)
,
comb. n.
from
Gonatocerus
.
Gonatocerus gunathilagaraji
Manickavasagam & Rameshkumar, 2013a
: 92
;
holotype
♀ in EDAU.
TL:
India
, Karnataka, Bengaluru, IWST campus.
Cosmocomoidea hayati
(Zeya & Khan)
,
comb. n.
from
Gonatocerus
.
Gonatocerus hayati
Zeya & Khan, 2012
: 59
;
holotype
♀ in NPC.
TL:
India
, West Bengal, Islampur, Sibdargipara.
Cosmocomoidea helavai
(Yoshimoto)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus helavai
Yoshimoto, 1990
: 87
;
holotype
♂ in CNC (examined). Triapitsyn
et al
., 2010: 160 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Panama
, Cerro Campana,
850m
[
8.333ºN
81.967ºW
].
Cosmocomoidea heratyi
(Manickavasagam & Rameshkumar)
,
comb. n.
from
Gonatocerus
.
Gonatocerus heratyi
Manickavasagam & Rameshkumar, 2013a
: 97
;
holotype
♀ in EDAU.
TL:
India
, Karnataka, Bengaluru, IWST campus.
Cosmocomoidea impar
(Huber)
,
comb. n.
from
Gonatocerus
.
Gonatocerus impar
Huber, 1988
: 69
;
holotype
♀ in USNM (examined).
TL:
USA
, CA, Menifee Valley, hills on W. end, 1800'.
Cosmocomoidea inaudita
(Ogloblin)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Lymaenon inauditus
Ogloblin, 1936
: 36
;
lectotype
♀ in MLPA, (examined).
De
Santis, 1967: 10, 104 (transferred to
Gonatocerus
); Triapitsyn
et al
., 2010: 166 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Argentina
, Loreto, Misiones.
Cosmocomoidea incompta
(Huber)
,
comb. n.
from
Gonatocerus
.
Gonatocerus incomptus
Huber, 1988
: 67
;
holotype
♀ in USNM (examined).
TL:
USA
, California, Riverside Co., Riverside.
Cosmocomoidea inexpectata
(Huber)
,
comb. n.
from
Gonatocerus
.
Gonatocerus inexpectatus
Huber, 1988
: 75
;
holotype
♀ in CNC (examined).
TL:
Canada
, Ontario, Constance Bay.
Cosmocomoidea inflatiscapus
(Huber)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus inflatiscapus
Huber 1988
: 70
;
holotype
♀ in USNM (examined). Triapitsyn
et al
., 2010: 170 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
USA
, California, Barton Flats, hwy. 38, Hathaway Creek.
Cosmocomoidea kashipurensis
(Zeya & Khan)
,
comb. n.
from
Gonatocerus
.
Gonatocerus kashipurensis
Zeya & Khan, 2012
: 58
;
holotype
♀ in NPC.
TL
:
India
, Uttarakhand, Kashipur, Pipalia.
Cosmocomoidea kikimora
(Triapitsyn)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus
(
Cosmocomoidea
)
Triapitsyn, 2013a
: 141
;
holotype
♀ in ZIN.
TL
:
Russia
, Primorskiy Kray, Ussuriyskiy rayon, Gornotayozhnoye.
Cosmocomoidea kiskis
(Triapitsyn)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus
(
Cosmocomoidea
) Triapitsyn
in Triapitsyn
et al
., 2010: 172;
holotype
♀ in MPLA (examined).
TL:
Argentina
, Tucumán, Tafí Viejo.
Cosmocomoidea kodaiana
(Mani & Saraswat)
,
comb. n.
from
Gonatocerus
.
Ooctonus kodaianus
Mani & Saraswat, 1973
: 78
;
holotype
♀ in USNM (examined).
Zeya & Hayat, 1995
: 66 (transferred to
Gonatocerus
)
TL:
India
, Tamil Nadu, Anamalai-Kodaikanal Hills, Berijam.
†
Cosmocomoidea kootenai
(Huber)
,
comb. n.
from
Gonatocerus
.
Gonatocerus kootenai
Huber
in
Huber & Greenwalt, 2011
: 481
;
holotype
♀ in USNM (examined).
TL:
USA
, Montana, Flathead River, Kishenehn Formation.
Cosmocomoidea latipennis
(Girault)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus latipennis
Girault, 1911
[96]: 268;
holotype
♂ in USNM (examined).
Triapitsyn, 2013a
: 145 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
North
America
, no locality given.
Cosmocomoidea maxima
(Girault)
,
comb. n.
from
Gonatocerus
.
Gonatocerus maximus
Girault, 1911
[96]: 264;
holotype
♀ in INHS (lost).
TL:
USA
, Illinois, Pulaski. Synonymy under
G
.
latipennis
by
Huber, 1988
: 60.
Cosmocomoidea logarzoi
(Triapitsyn)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus
(
Cosmocomoidea
) Triapitsyn
in Triapitsyn
et al
., 2010: 174;
holotype
♀ in MPLA (examined).
TL:
Argentina
, Corrientes,
20 km
E. Corrientes, Centro Nacional de Actividades Agrícolas.
Cosmocomoidea lucida
(Dodd)
,
comb. n.
from
Gonatocerus
.
Gonatocerus lucidus
Dodd, 1919
: 161
;
holotype
♀ in BMNH (examined).
TL:
Indonesia
, Java, Tjibodas.
Cosmocomoidea maculipennis
(Ashmead)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Litus maculipennis
Ashmead 1900
: 265
;
holotype
♂ in BMNH (examined). Triapitsyn
et al
., 2010: 176 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
St. Vincent and the Grenadines
,
St. Vincent
Island
.
Cosmocomoidea marilandica
(Girault)
,
comb. n.
from
Gonatocerus
.
Gonatocerus marilandicus
Girault, 1917
[301]: 115;
holotype
♀ in USNM (examined).
TL:
USA
, Maryland, Glenn Dale.
Cosmocomoidea metanotalis
(Ogloblin)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Lymaenon metanotalis
Ogloblin, 1938a
: 35
;
lectotype
♀ in MLPA (examined).
De
Santis, 1967: 10, 104 (transferred to
Gonatocerus
); Triapitsyn
et al
., 2010: 179 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Argentina
, Misiones,
Puerto Rico
.
Cosmocomoidea monticola
(Zeya)
,
comb. n.
from
Gonatocerus
.
Gonatocerus monticolus
Zeya
in
Zeya & Hayat, 1995
: 68
;
holotype
♀ in BMNH (examined).
TL:
India
, Uttar Pradesh, Mussoorie.
Cosmocomoidea morgani
(Triapitsyn)
,
comb. n.
from
Gonatocerus
.
Gonatocerus morgani
S. Triapitsyn, 2006: 9
;
holotype
♀ in UCRC.
TL:
USA
, California, Orange Co., Irvine, Irvine Ranch.
Cosmocomoidea morrilli
Howard
,
comb. rev.
from
Gonatocerus
(
Cosmocomoidea
)
.
Cosmocomoidea morrilli
Howard, 1908
: 69
;
lectotype
♀ in USNM (examined). Girault, 1929 [428]: 21 (transferred to
Ooctonus
);
Ogloblin, 1959a
: 50
[(transferred to
Lymaenon
(
Cosmocomoidea
)
];
Bouček & Graham, 1972
: 127
(transferred to
Gonatocerus
);
De
Santis, 1979
: 12
, 367 [(placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
USA
, Florida, Orlando.
Cosmocomoidea mumu
(Triapitsyn)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus (Cosmocomoidea) mumu
Triapitsyn
in Triapitsyn
et al
., 2010: 185;
holotype
♀ in MLPA (examined).
TL:
Argentina
, Misiones, Loreto, Ruinas Jesuíticas.
Cosmocomoidea nasuta
(Ogloblin)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Lymaenon nasutus
Ogloblin, 1939
: 244
;
lectotype
♀ in MLPA (examined).
De
Santis, 1967: 10, 104 (transferred to
Gonatocerus
); Triapitsyn
et al
., 2010: 187 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Argentina
, Misiones, Loreto.
Cosmocomoidea nigriflagellum
(Girault)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Ooctonus nigriflagellum
Girault, 1914
[195]: 150;
holotype
♀ in ZMHB.
Mendonça Filho, 1972
: 39 (transferred to
Lymaenon
);
De
Santis, 1979
: 12, 366 (transferred to
Gonatocerus
); Triapitsyn
et al
., 2010: 189 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Paraguay
, San Bernardino.
Cosmocomoidea
h-lutea
(Ogloblin).
Lymaenon h-luteum
Ogloblin, 1938a
: 29
;
lectotype
♀ in MLPA (examined).
De
Santis, 1967: 10, 104 (transferred to
Gonatocerus
).
TL:
Argentina
: Buenos Aires, Villa Ballester. Synonymy under
nigriflagellum
by Triapitsyn
et al
., 2010: 189.
Cosmocomoidea nigrithorax
(Ogloblin)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Lymaenon nigrithorax
Ogloblin, 1953
: 2
;
lectotype
♀ in MLPA.
De
Santis, 1967: 104 [transferred to
Gonatocerus
(
Gonatocerus
)
]; Triapitsyn
et al
., 2010: 194 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Argentina
, Misiones, Monte Carlo.
Cosmocomoidea novifasciata
(Girault)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus novifasciatus
Girault, 1911
[96]: 266;
holotype
♀ in USNM (examined). Triapitsyn
et al
., 2010: 200 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
USA
, Tennessee, no locality given.
Cosmocomoidea oxypygus
(Foerster)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus oxypygus
Foerster, 1856
: 118
;
neotype
♀ in MHNG, designated by
Triapitsyn, 2013a
: 150
.
Triapitsyn, 2013a
: 150
[placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Europe, possibly
Germany
, Aachen (see
Triapitsyn, 2013a
: 150
, for discussion of locality).
Cosmocomoidea megalura
Mathot.
Lymaenon megalura
Mathot, 1969
: 8
;
holotype
♀ in IRSNB.
TL:
Belgium
, Louvain. Synonymy under
G
.
oxypygus
by
Triapitsyn, 2013a
: 150
.
Cosmocomoidea ovicenatus
(Leonard & Crosby)
.
Gonatocerus ovicenatus
Leonard & Crosby, 1915
: 545
;
lectotype
♀ in CUIC (examined).
TL:
USA
, New York, Ithaca. Synonymy under
G
.
oxypygus
by
Triapitsyn, 2013a
: 150
.
Cosmocomoidea parcepilosa
(Ogloblin)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Lymaenon parcepilosus
Ogloblin, 1957
: 35
;
holotype
♀ in MLPA.
De
Santis, 1967: 10, 105 (transferred to
Gonatocerus
); Triapitsyn
et al
., 2010: 202 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Argentina
: Neuquén, Pucará, Lago Lacar.
Cosmocomoidea portoricensis
(Dozier)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus portoricensis
Dozier, 1937
: 131
;
holotype
♀ in USNM (examined). Triapitsyn
et al
., 2010: 205 [placed in
Gonatocerus
(
Cosmocomoidea
)
.
TL:
Puerto Rico
, Isabela.
Cosmocomoidea quadrivittata
(Dozier)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus quadrivittatus
Dozier, 1932
: 81
;
holotype
♀ in USNM (examined). Triapitsyn
et al
., 2010: 207 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Haiti
, Port-au-Prince.
Cosmocomoidea quirogai
(Ogloblin)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Lymaenon quirogai
Ogloblin, 1936
: 44
;
holotype
♀ in MLPA (examined).
TL:
Argentina
, Misiones, Tedyú Cuaré, opposite San Ignacio.
Ogloblin, 1959a
: 50 [placed in
Lymaenon
(
Cosmocomoide
a)];
De
Santis, 1967: 10, 106 [(transferred to
Gonatocerus
(
Cosmocomoidea
)
].
Cosmocomoidea rakitovi
(Triapitsyn)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus
(
Cosmocomoidea
)
rakitovi
Triapitsyn
, in Triapitsyn
et al
., 2010: 212;
holotype
♀ in UCRC (examined).
TL:
Costa Rica
, Cartago, Parque Nacional Tapantí,
1290 m
.
†
Cosmocomoidea rasnitsyni
(Huber)
,
comb. n.
from
Gonatocerus
.
Gonatocerus rasnitsyni
Huber
in
Huber & Greenwalt, 2011
: 483
;
holotype
♀ in USNM (examined).
TL:
USA
, Montana, Flathead River, Kishenehn Formation.
Cosmocomoidea rogersi
(Matthews)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus rogersi
Matthews, 1986
: 222
;
holotype
♀ in BMNH.
Triapitsyn, 2013a
: 161 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
England
, Hampshire, Awbridge.
Cosmocomoidea rufescens
(Ashmead)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Polynema rufescens
Ashmead, 1904
: 521
;
holotype
♂ in USNM (examined).
Ogloblin, 1946
: 286 (transferred to
Gonatocerus
); Triapitsyn
et al
., 2010: 213 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Brazil
, Pernambuco.
Cosmocomoidea sahadevani
(Subba Rao & Kaur)
,
comb. n.
from
Gonatocerus
.
Lymaenon sahadevani
Subba Rao & Kaur, 1959
: 231
;
holotype
♀ in IARI.
Subba Rao & Hayat, 1983
: 136 (transferred to
Gonatocerus
).
TL:
India
, New Delhi.
Cosmocomoidea schajovskoii
(Ogloblin)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Lymaenon schajovskoii
(
Ogloblin, 1957: 37
)
;
holotype
♀ in MLPA (examined).
De
Santis, 1967: 10, 105 (transferred to
Gonatocerus
). Triapitsyn
et al
., 2010: 215 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Argentina
: Neuquén, Pucará, Lago Lacar.
Cosmocomoidea seminigra
(Ogloblin)
,
comb n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Lymaenon
(
Cosmocomoidea
)
Ogloblin, 1959a
: 53
;
holotype
♀ in MLPA.
De
Santis, 1979
: 12, 368 [transferred to
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Ecuador
, Esmeraldas, San Mateo.
Cosmocomoidea tremulae
(Bakkendorf)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Lymaenon tremulae
Bakkendorf, 1934: 30
;
lectotype
♀ in ZMUC (lost). Bakkendorf did not specify a
holotype
or state the number of specimens he examined so the
holotype
designation by
Matthews (1986)
is deemed a
lectotype
under ICZN Rule 74.6.
Bouček & Graham, 1978a
: 109 (transferred to
Gonatocerus
);
Triapitsyn, 2013a
: 163 [placed in
Gonatocerus
(
Cosmocomoidea
)
, and
lectotype
fixation].
TL:
Denmark
, Copenhagen, Fortunen Indelukke. Synonymy under
G
.
ovicenatus
by
Baquero & Jordana, 2002
: 16; treated as separate species by
Pricop, 2010
: 117.
Cosmocomoidea trialbifuniculatus
(Subba Rao)
,
comb. n.
from
Gonatocerus
.
Gonatocerus trialbifuniculatus
Subba Rao 1989
: 147
;
holotype
♀ in BMNH (examined).
TL:
India
, Karnataka,
25 km
W. Mudigere.
Cosmocomoidea triangulifera
(Ogloblin)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Lymaenon triangulifer
Ogloblin, 1959a
: 56
;
holotype
♀ in MLPA (examined).
De
Santis, 1979
: 367 (transferred to
Gonatocerus
); Triapitsyn
et al
., 2010: 221 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Ecuador
, Esmeraldas, San Mateo
Cosmocomoidea triguttata
(Girault)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus triguttatus
Girault, 1916
[287]: 297;
lectotype
♀ in USNM (examined). Triapitsyn
et al
., 2010: 222 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Trinidad
, Caroni.
Cosmocomoidea tuberculifemur
(Ogloblin)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Lymaenon tuberculifemur
Ogloblin, 1957
: 38
;
holotype
♀ in MLPA (examined).
De
Santis, 1967: 10, 105 (transferred to
Gonatocerus
); Triapitsyn
et al
., 2010: 225 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Argentina
: Neuquén, Pucará, Lago Lacar.
Cosmocomoidea uat
(Triapitsyn)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus uat
S. Triapitsyn
in Triapitsyn
et al
., 2006: 58,
holotype
♀ in UCRC (examined). Triapitsyn
et al
., 2010: 227 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Mexico
, San Luis Potosí, Ciudad Valles.
Cosmocomoidea udakamanda
(Mani & Saraswat)
,
comb. n.
from
Gonatocerus
.
Gonatocerus udakamandus
Mani & Saraswat, 1973
: 96
;
holotype
♀ in USNM (examined).
TL:
India
, Ootacamund, Doddabetta.
Cosmocomoidea virlai
(Triapitsyn, Logarzo & de León),
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus virlai
S. Triapitsyn, Logarzo
& de León in
Triapitsyn
et al
., 2007
: 62;
holotype
♀ in MLPA (examined). Triapitsyn
et al
., 2010: 229 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
Argentina
, Tucumán, Tafí Viejo.
Cosmocomoidea walkerjonesi
(Triapitsyn)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus walkerjonesi
S. Triapitsyn, 2006: 15
;
holotype
♀ in UCRC (examined). Triapitsyn
et al
., 2010: 231 [placed in
Gonatocerus
(
Cosmocomoidea
)
].
TL:
USA
, California, Ventura Co., Fillmore.
Cosmocomoidea woohoo
(Triapitsyn)
,
comb. n.
from
Gonatocerus
(
Cosmocomoidea
)
.
Gonatocerus
(
Cosmocomoidea
)
Triapitsyn, 2013a
: 168
;
holotype
♀ in CAS.
TL:
Russia
, Sakhalinskaya Oblast’, Sakhalin
Island
, ~
6 km
E. Sokol.