Land snail diversity in the monsoon tropics of Northern Australia: revision of the genus Exiligada Iredale, 1939 (Mollusca: Pulmonata: Camaenidae), with description of 13 new species
Author
Criscione, Francesco
Author
Law, Margot Louisa
Author
Köhler, Frank
text
Zoological Journal of the Linnean Society
2012
Zool. J. Linn. Soc.
2012-11-26
166
4
689
722
http://dx.doi.org/10.1111/j.1096-3642.2012.00863.x
journal article
10.1111/j.1096-3642.2012.00863.x
0024-4082
5409181
AC086999-76A8-4F93-9A51-5B5B213A50E6
EXILIGADA BRABYI
SP. NOV.
(
FIGS 5B
,
7D–E
)
Exiligada negriensis
–
Solem, 1984: 673
, plate 57F; figure 175 (a–c) (in part).
Holotype
:
AUSTRALIA
,
NT
,
VRD
, east of
Duncan Hwy
,
37.7 km
north-west of
Kirkimbie Station
H
/S, large, open, and very rugged limestone formation, in talus, under spinifex,
17°26
′
59
′′
S
,
129°03
′
20
′′
E
(coll.
V. Kessner
, R. Hokkanen
2.ix.2009
), wet (
WA
– 81/09,
WAM
S83168
).
Paratypes
:
Same
data as for holotype, 27 dry, two wet (
WAM
S49159
);
Duncan Rd
, old
Ord River
homestead,
Forrest Creek
(Ck),
70 km
north of
Nicholson
homestead, under limestone rocks and concrete of old structures,
17°23
′
56
′′
S
,
128°52
′
01
′′
E
(coll.
R. Crookshanks
,
29.i.2005
), four dry, two wet (
AM
C.443279)
.
Etymology:
In honour of Michael Braby, for his most valuable help with fieldwork in the
Victoria
River District, noun in genitive case.
Description:
Shell (
Figs 5B
,
7D, E
). Moderately large, moderately elevated, moderately thick; micropustulations absent; background colour yellowish horn fading to whitish towards shell base, with one faint to conspicuous, continuous, brown-reddish peripheral band, and few regularly spaced, narrower spiral bands, more conspicuous on body whorl, absent on shell base; no maculations.
Genitalia. Epiphallus as long as penis. Vas deferens entering penial sheath within anterior third. Penis five times longer than wide, not coiled inside sheath, with thick wall, inner penial wall with thick, regularly shaped, sometimes interrupted main longitudinal pilaster restricted to posterior half, with several thin longitudinal folds anteriorly. Vagina half as long as penis, as long as free oviduct.
Bursa
copulatrix with no well-differentiated end (
Solem, 1984
: fig. 175, ‘
E. negriensis
’).
Remarks:
Shell colour pattern not readily distinguishable from that of
E. qualis
.
Solem’s (1984: 676
, fig. 175) description of genital anatomy of ‘
E. negriensis
’ is here attributed to
E. brabyi
. Penis length relative to width (pl/pw) average, epiphallus length relative to penis (el/pl) half of average. Inner penial wall sculpture very similar to that of
E. qualis
and
Exiligada unistriata
, from which it can be distinguished by the posteriorly situated, main longitudinal pilaster being interrupted and by a larger number of anterior folds. Vagina length relative to penis and free oviduct (vl/pl, vl/ol) shorter than average.