Revision of the Australian species of Figulus MacLeay, 1819 (Coleoptera: Lucanidae)
Author
Monte, Cinzia
Author
Zilioli, Michele
Author
Bartolozzi, Luca
text
Zootaxa
2016
4189
3
447
484
journal article
37842
10.11646/zootaxa.4189.3.2
c55b8fbe-5e88-4dae-89fb-38df76c0b839
1175-5326
166170
3B503E4F-7748-4AC4-965E-B010E225255B
Figulus lilliputanus
Westwood, 1855
(
Figs. 8–11
)
Figulus lilliputanus
Westwood, 1855
: 219
;
Thomson 1862
: 432
;
Parry 1863
: 450
;
Parry 1864b
: 100
;
Harold 1868
: 965
;
Macleay 1871
: 173
;
Deyrolle 1874
: 414
;
Parry 1875
: 22
;
Felsche 1898
: 78
;
Van Roon 1910
: 52
;
Mjöberg 1916
: 13
; Arrow 1938: 53, 60, 61;
Arrow 1943
: 139
;
Didier & Séguy 1953
: 180
;
Benesh 1950
: 53
;
Benesh 1955
: 64
, 66;
Benesh 1960
: 26
;
Klausnitzer 1982
: 15
;
Bomans 1986
: 9
(
lectotype
designation);
Maes 1992
: 36
, 38;
Moore & Cassis 1992
: 10
;
Araya 2001
: 13
;
Krajcik 2001
: 11
;
Krajcik 2003
: 79
;
Klausnitzer & Sprecher-Uebersax 2008
: 29
;
Fujita 2010
: 395
;
Maes 2014
.
Figulus clivinoides
Thomson, 1862
: 402
, 432; synonymised by
Parry 1863
: 450
.
Figulus
(
Figulinus
)
lilliputanus
:
Schoolmeesters 2015
.
Type material
.
Figulus lilliputanus
Westwood
, lectotype (designation by
Bomans 1986
): “S Austral [handwritten]”, “
Figulus
/
Liliputanus
(sic!) / Tr. Ent. Soc. n.s. III, pl. 12, fig. 5 [handwritten]”, “TYPE / WESTWOOD / Trans. Ent. / Soc. 1855 / P.219 T.12 f.5 / Coll. Hope Oxon. [printed, handwritten]”, “TYPE COL: 333
1
/
2
/
Figulus
/
lilliputanus
/ Westw. / HOPE DEPT. OXFORD [printed, handwritten]”, [OUMNH];
Figulus lilliputanus
Westwood
, paralectotype: “S Austral [handwritten]”, “W [printed]”, “TYPE / WESTWOOD / Trans. Ent. Soc./ 1855 P.219. / T.12 fig.5 / Coll. Hope Oxon. [printed, handwritten]”, “TYPE COL: 333
2
/
2
/
Figulus
/
lilliputanus
/ Westw. / HOPE DEPT. OXFORD [printed, handwritten]”, [OUMNH];
Figulus clivinoides
Thomson
:
Lectotype here designated
:
Lectotype
: “Th. / Type [printed]”, “
Clivinoides
/ Type Thoms / Austr [handwritten]”, “Ex Musaeo / James Thomson [printed]”, “MUSÉUM PARIS / 1952 / COLL. R. OBERTHÜR [printed]”, “H.Bomans det., 1973 / =
Figulus
/
lilliputanus
West. [printed, handwritten]”, [MNHN].
Other material.
Figulus lilliputanus
Westwood
(Ƌ): “
Lilliputanus
/ West. Ent. J. /
v. 3
Feb [handwritten]”, “Ex Musaeo / Parry [printed]”, “MUSÉUM PARIS / 1952 / Coll. R. OBERTHÜR [printed]”, [MNHN];
Figulus lilliputanus
Westwood
(Ƌ): “Ex Musaeo / Parry [printed]”, “MUSÉUM PARIS / 1952 / Coll. R. OBERTHÜR [printed]”, [MNHN];
Figulus lilliputanus
Westwood
(♀): “
Lilliputanus
/ West. /
Clivinoides
/ Thompson [handwritten]”, “Ex Musaeo / Parry [printed]”, “MUSÉUM PARIS / 1952 / Coll. R. OBERTHÜR [printed]”, [MNHN]. Note: this material was indicated as syntypes by S. Boucher (MNHN) in the loan form, but their status as type material is uncertain, as they have no red label or clear indication of being part of the original type series.
Diagnosis
.
This is the smallest species of Australian
Figulus
and differs from all other Australian species by its antenna with only 9 antennomeres. The irregular transversal rows of dense and deep punctures with short erected setae directed downwards on the ventrites 2‒4 are secondary sexual characters, as in other
Figulus
species (
Bomans 1989
).
Redescription.
Measurements
: size range (
n
= 165): TL: 6.0–10.0 mm; PL: 1.6
1–2.49 mm
; EL:
3.41–5.43 mm
; PW:
1.69–2.97 mm
; EW: 1.74–3.0 mm.
Male (
Fig. 8
). Body elongate, with subparallel sides, dark brown or reddish brown, glossy.
Head
: transverse, about twice as wide as long. Frontoclypeal region slightly concave at the anterior margin coarsely punctate with an evident median elevation and two large, glossy areas situated before the eyes. Labrum broad, short, with anterior margin slightly bilobed and with long, golden setae. Vertex slightly concave in the middle, feebly inclined anteriorly, with two small, circular elevations. Two longitudinal, slightly elevated carinae are present along the inner margin of eyes. Eyes small, divided into 2 parts by the canthi, the upper part smaller than the lower with distinct facets. Canthus slightly protruding, rounded laterally and angled behind; upper surface slightly depressed and smooth, with sparse, minute punctures on the lateral margin. Head surface densely punctate with superficial, large punctures smallest on canthi, anterior margin of the frons, longitudinal carinae and transversal elevations of vertex. Mandibles shorter than head, rounded laterally, apices acute; upper surface with external margin slightly raised. Internal margin of mandibles with a medial strong tooth, with apex blunted directed inwards. Antenna with 9 antennomeres, with scape as long as the remaining antennomeres combined; club of last three antennomeres, which are flatter and broader than the remaining antennomeres of the funicle.
Mentum
subsquare (
Fig. 9
), slightly concave medially; anterior margin slightly bilobed; base smooth inclined forwards, with sides slightly elevated at mid-length. Surface smooth, with mixed large punctures and wrinkles.
Thorax
: pronotum subsquare with subparallel sides; posterior width less than anterior width; anterior margin almost straight, with a small, blunt median tubercle; posterior margin slightly bisinuate; anterior angles produced with anterior margin rounded; posterior angles largely rounded, with slightly crenulate margin. Superficial longitudinal medial groove on pronotal disc, with some large, circular punctures. Pronotal surface glossy with dense, small, superficial punctures, smaller along the anterior margin of prothorax and with patch of large, close punctures at sides. Long, golden setae directed upwards along the lateral pronotal margin. Scutellum long, wedgeshaped; surface smooth and glossy. Elytra about twice as long as pronotum. Base concave, subparallel sides, apex regularly rounded. Humeral angles acute, with a small tubercle. Elytral surface glossy, with 9 visible subparallel striae, the dorsal striae deeper than the lateral striae; each stria formed by a row of large punctures well separated from each other, except for the first stria, deeper than others and where the punctures are confluent. Stria 1 the only stria than reaches apex; stria 2 ending slightly before; stria 3 curved outwards, converging with stria 6; striae 4 and 5 converging at declivity. Interstriae much larger than striae, slightly convex and smooth, with a longitudinal, irregular row of small punctures. Elytral margins and apex strongly punctate, with short, erect, red setae. Wings present. Legs: thin and straight. Protibiae with strong, elongate, apical fork, slightly dilated at the point where it articulates a long, strong spur; lateral margin with 3‒4 small teeth, increasing in size distally. Mesotibiae with 4 long, acute apical teeth, and 2 apical spurs, 1 larger than the other; lateral margin with strong submedial tooth, preceded by 1‒2 smaller teeth; ventral lateral margin of mesotibiae with 2‒3 small teeth. Metatibiae with 4 acute apical teeth, and 2 long apical spurs, 1 larger than the other; lateral margin with strong submedial tooth. Prosternum subpentagonal in shape behind the coxae; surface with small punctures at anterior edge and with smaller punctures medially and at posterior edge. Metasternal disc smooth, with a long medial groove, deep at posterior edge; sides of metasternum with large and dense, horse-shoe shaped, superficial hollows.
Abdomen
: first abdominal ventrite with some large and dense, horse-shoe shaped, superficial hollows with a long, golden seta in the middle. Ventrites 2‒4 with some deep longitudinal depressions along the anterior margin, with short, golden setae emerging from the anterior margin; they are not present in the anteromedial area where there are 1‒3 irregular transversal rows of dense and deep punctures, each with a short seta, erected and directed downwards. Last ventrite with some punctures, which are smaller near the posterior margin.
FIGURES 8‒11.
Figulus lilliputanus
Westwood.
8, dorsal view of adult (scale bar: 5 mm); 9, mentum; 10, male genitalia, dorsal view (scale bar: 0.5 mm); 11, male genitalia, lateral view.
Male genitalia
: penis tubular (
Figs. 10–11
), short and narrow, moderately curved in lateral view, strongly bent proximally close to articulation with basal piece, almost completely covered by a dorsal translucid extension of the basal piece. Permanently everted internal sac drawn out into a narrow colorless or yellow flagellum, slightly longer than the tegmen (basal piece + parameres), with two subsequent forkings in the basal half, tapering towards the apex (
Fig. 10
). Parameres slightly shorter than basal piece, with marginal fringe of setae entirely enclosed in the cuticle (not represented in figure); ventrally the base of the parameres ends in a long spine-shaped apophysis articulated with basal piece (
Figs. 10–11
). Basal piece ventrally largely membranous, dorsally sclerotized with a moderately large (about one-half of basal piece length), translucent cuticular area along the midline. Two long, slender struts run along basal piece, not reaching its base.
Female.
All external morphological characters as in male, except for sternites 2‒4 smooth and glabrous, without irregular, transverse punctation or short setae.
Female genitalia
: hemisternites elongate, broader anteriorly, narrower posteriorly; anteriorly broad up to about half of their length; outer edge notched; medial edge convex throughout length. Bursa copulatrix represented by a slight expansion of the wall of spermathecal duct near junction with the median oviduct. Spermathecal duct short, ending with a curve and long expansion, proximally colorless, distally pigmented.
Intraspecific variation.
The specimens examined do not show significant morphological variability, except for the body size (see “Measurements” section).
Additional material examined.
Australia
[state unknown]:
8 specimens
,
29.IX.1929
,
E.Sutton Collection
, donated
Dec.1964
(
QMB
)
;
1 specimen
, ex
Musaeo Ed. Brown
,
Muséum Paris
1952
Collection Oberthür
(
MNHN
)
;
1 specimen
, ex
Musaeo W.
Möllenkamp,
Muséum Paris
1952
Collection R.
Oberthür (
MNHN
)
; 2 specimens, det. A.M.Lea (MEL);
5 specimens
,
Nat. Mus.
Victoria
, COL-85173, COL-85174, COL-85175, COL-85176, and COL- 8517 (
NMVM
)
; 1 specimen (GTC);
1 specimen
, ex
Musaeo Mniszech
,
Muséum Paris
1952
Collection Oberthür
(
MNHN
)
;
1 specimen
, Nouvelle Hollande,
R.I.Sc.
N.B. 21.418,
Collection P.
de Moffarts, 4764,
Comp.
au
type
,
H.E.Bomans
BMNH
(E) 1999-248, 404764 (handwritten) (
NHM
)
.
AUSTRALIAN
CAPITAL
TERRITORY
:
1 specimen
,
Black Mountain
,
1.II.1968
,
light trap
,
M.S.Upton
(
ANIC
)
.
NEW SOUTH WALES
:
1 specimen
,
New
South Wales
,
Macleay Museum University
of
Sydney
2006
Australia
, 11384,
H.E.Bomans
BMNH
(E) 1999-248, 413884 (handwritten) (
NHM
)
;
1 specimen
,
New
South Wales
,
Nat. Mus
.
Victoria
, COL-85784 (
NMVM
)
;
1 specimen
,
Albury
,
16.II.1970
,
R.Parrot
,
Collection R.E.
Parrot (
MEL
); 1Ƌ,
Avoca Station
off
Pomona
road
20 km
N of Wentworth
,
13.VII.1993
, COL-86865 (NMVM); 1 specimen, Ballimore, F.H.Taylor (ANIC);
1♀
,
1 specimen
,
N Belmore Road
,
3.2 km
SSW of
Busbys Flat
,
26.I.
1999
, 170 m, at mv light,
S.Watkins
,
S.Watkins Collection
donated 2001 (
ANIC
)
;
1Ƌ,
Bungabbee State Forest
, SE
Kyogle,
29.I.1997
, mv light,
S.G.Watkins
, 97:199
Ƌ, S.G.Watkins Collection
, donated 2001 (
ANIC
);
1 specimen
,
Como
,
Dr. K.K.Spence, K.K.Spence Collection, K
293062 (AMS)
;
1 specimen
,
Coonabarabran
,
XII.1929
,
H.J.Carter
(
ANIC
)
;
1♀
,
Fortis Creek
National Park
,
South
boundary, N of
Grafton
,
4.I.
2000
, 50 m, at mv light,
S.Watkins Collection
donated 2001 (
ANIC
)
;
2ƋƋ,
Fortis Creek
National Park
,
South
boundary, N of
Grafton
,
17.XII.
1998
, 50 m, mv light, 98:837Ƌ and 98:838
Ƌ, S.Watkins Collection
donated 2001 (
ANIC
)
;
2 specimens
,
Maitland
,
26.XII.1962
,
A Neboiss
COL-85777 (
NMVM
)
;
1 specimen
,
Mogriguy
,
Goonoo State Forest
,
13.II.1981
,
R.H.M., R.H.Mulder Collection
, K 293063 (AMS)
;
1 specimen
,
Mullaly
,
XII.1929
,
H.J.Carter
(
ANIC
)
;
2 specimens
,
Sydney
,
Hyde Park
,
XII.1985
, K 293074 (AMS)
; 2 specimens, Wagga, 1937, C.Oke, COL-85913 (NMVM);
2 specimens
,
Wagga
, 1938,
C.Oke
,
Presented
by
C.G.Oke
1.IX.1948
, COL-85883 (
NMVM
)
.
QUEENSLAND
:
1 specimen
,
Queensland
, Nat. Mus
.
Victoria
, COL-85916 (
NMVM
);
1 specimen
,
Queensland
, Nouvelle Hollande,
Fry Collection
1905-100,
BMNH
(E) # 607642 (
NHM
)
;
1 specimen
,
Blackdown Tableland
, via
Dingo
,
XII.1978
,
Maywald
&
Czechura
(
QMB
); 1Ƌ,
Boggomoss
15 in
field,
Taroom District
,
25°28'S
,
150°01'E
,
12.XI.1996
, P.Lawless, 191, QM Reg. No T 34260 (QMB); 1Ƌ, Boggomoss number 8 via Taroom, 25°27'S, 150°02'E,
12.XI.1996
,
Queensland
Mus.
survey, day hand collection, 0 39, QM Reg. No T 34259 (
QMB
)
;
1Ƌ,
2 specimens
, Boyne Island, via
Gladstone
,
23°55'S
,
151°20'E
, 1995,
C.Q. University
, 5022 (
QMB
)
;
1Ƌ, Bribie Island,
XII.1925
,
H.Hacker
(
QMB
);
1 specimen
,
Brisbane
,
29.X.1917
and 1 specimen,
4.III.1920
, H.Hacker (QMB); 2 specimens, Brisbane (QMB);
2 specimens
,
Brisbane
,
Mount Coot-tha
,
I.1966
,
C.Oke
COL-85890 (
NMVM
)
;
1 specimen
,
Cape York
, 3851 (
QMB
)
;
6 specimens
,
Clermont
,
XII.1929
,
Dr. K.K.Spence, K.K.Spence Collection, K
293070 K 293071, and K 293072 (AMS)
;
2 specimens
,
Cooktown
,
I.1971
,
J.G.Brooks
,
J.G.Brooks Bequest
1976 (
ANIC
)
;
1 specimen
,
Dawson District
,
Barnard Collection
,
Muséum Paris
1952
Collection Oberthür
(
MNHN
)
;
1 specimen
,
Dawson River
,
Macleay Museum University
of
Sydney
2006, 11383 (handwritten),
H.E.Bomans
1999-248, 411383 (
NHM
)
;
1Ƌ,
2♀♀
,
Dunmore State Forest
,
27°38'S
,
150°58'E
,
26.XII.1998
, 98:883 Ƌ, mv light,
S.Watkins Collection
donated 2001 (
ANIC
)
;
1 specimen
,
Garradunga Stone Creek
,
15.I.1997
,
J.Hasenpusch
, R.deKeyzer
Insect Collection, K
293325 (AMS)
;
1 specimen
,
15 km
S of
Greenvale
,
9.I.1911
,
B.B.Howery
(
ANIC
); 1Ƌ,
Gurgeena Plateau
,
25°27'S
,
151°23'E
,
8.VI.
1999
, 360 m, open forest, 7741,
G.B.Monteith
&
G.I.Thompson
(
QMB
);
1 specimen
,
Hann River
,
15°11'S
,
143°52'E
(GPS)
,
18.XII.1993
‒
14.I.1994
,
flight intercept trap
,
P.Zborowski
,
E.D. Edwards
(
ANIC
);
1 specimen
,
14 km
ENE
Heathlands
,
11°41'S
,
142°42'E
,
8.XII.1992
, at light rainforest, P. Zoborowski & W.Dressler (ANIC); 1 specimen, Iron Range, W Claudie Rg, 12°45'S, 143°14'E,
3.XII.
1985
, 50 m, rainforest, stick brushing, G.B.Monteith, QM Berlesate No 690 (QMB);
1 specimen
, same locality,
3.XII.
1985
, 50 m, rainforest,
Monteith
&
Cook
(
QMB
)
;
1 specimen
,
Iron Range National Park
,
Gordon Creek
(
QMB
)
;
1 specimen
,
Iron Range
,
28.X.1981
,
J.Hasenpusch
, R.deKeyzer
Insect Collection, K
293323 (AMS); 1Ƌ,
Iron Range
,
Cape York
Peninsula
,
VI‒VII.1977
, G.B.Monteith (QMB);
1 specimen
,
Kuranda
,
13.I.1987
, R.deKeyzer
Insect Collection, K
293322 (AMS)
; 1 specimen, Maryborough, E.W.Fischer, ex Collection Australian Museum (ANIC);
2 specimens
,
Moreton Bay
,
K 15076
(handwritten), K 293064 (AMS)
;
1 specimen
,
Moreton Bay
, ex
Musaeo Ed. Brown
,
Muséum Paris
1952
Collection Oberthür
(
MNHN
)
; 1 specimen, Nipps, COL-85915 (NMVM);
1 specimen
,
Normanton
,
II.1982
, R.deKeyzer
Insect Collection, K
293324 (AMS)
;
1 specimen
,
Rockhampton
, ex
Musaeo Ed. Brown
,
Muséum Paris
1952
Collection Oberthür
(
MNHN
)
;
1 specimen
,
Rocky River
,
Silver Plains
, E
Coast
,
Cape York
Peninsula,
6.I.1960
,
J.L.Wassel
(
ANIC
)
; 2 specimens, Stanthorpe, E.Sutton, F.E.Wilson Collection, COL-85885 (NMVM);
5 specimens
,
Stanthorpe
,
29.VII.‒4.IX.1924
,
E.Sutton Collection
, donated
Dec.1964
(
QMB
)
;
2 specimens
,
Station Creek
, N
Queensland
,
30.XII.1964
,
J.G.Brooks
(
ANIC
)
;
1Ƌ,
2 specimens
,
Station Creek
,
30.XII.1969
,
J.G.Brooks
,
J.G.Brooks Bequest
1976 (
ANIC
)
;
1 specimen
,
Toomba
,
19°58'S
,
145°34'E
,
14‒16.XII.2007
,
Site
1, 390 m
, 14652, mv light, vinescrub edge/paddock,
Monteith
&
Wright, QM
Reg. No T147162 (
QMB
)
; 1 specimen, Townsville, K.K.Spence, K.K.Spence Collection, K 293069 (AMS);
3 specimens
,
9 km
ENE of
Mount Tozer
,
12°43'S
,
143°17'E
,
5‒10.VII.1986
, rotten log,
T.Weir
&
A.Calder
(
ANIC
)
;
1Ƌ,
Woodstock
,
3.VII.1992
,
B.P.Moore
(
LBC
)
.
SOUTH AUSTRALIA
:
2 specimens
,
South Australia
, ex
Macleay Museum University
of
Sydney
2006
Australia
(
GTC
,
LBC
)
;
1Ƌ,
South Australia
,
R. Mus. Hist. Nat. Belg.
, I.G. 12.595 (
IRSNB
)
; 1 specimen, Adelaide, BMNH(E) # 607646 (NHM);
2 specimens
,
Adelaide Hills
,
V.1930
,
C.Oke
,
Presented
by
C.G.Oke
1.IX.1948
, COL- 85892 (
NMVM
)
;
1 specimen
,
Adelaide
, ex
Musaeo Ed. Brown
,
Muséum Paris
1952
Collection Oberthür
(
MNHN
)
; 1 specimen, Adelaide, Griffith 0 4 (ANIC); 1 specimen, Adelaide,
K 28038
(handwritten), K 293073 (AMS); 1 specimen, Barossa, A.H.Elston, A.H.Elston Collection, K 293075 (AMS);
1 specimen
,
Buckaringa Gorge
,
Flinders Ra. C.
30 km
NNW of Quorn
,
19.XII.1985
,
Reid
,
Gullan
&
Lewis
,
at light trap
(
ANIC
)
.
VICTORIA
: 4ƋƋ,
Ballarat
,
VIII.1937
,
C.Oke
,
Presented
by
C.G.Oke
1.IX.1948
, COL-85875 and COL-85884 (
NMVM
)
; 1Ƌ, 1♀, Hattah, C.Oke COL-85914 (NMVM);
2 specimens
,
Lake Hattah
,
XI.1924
,
C.Oke
, COL-85785 and COL-85786 (
NMVM
)
;
1 specimen
,
Mount Hope
,
27.IX.1947
,
C.Oke
,
Presented
by
C.G.Oke
1.IX.1948
, COL-85881 (
NMVM
)
;
2 specimens
,
Piangil
,
20.VII.1919
,
C.Oke
, COL-85912 (
NMVM
)
;
1 specimen
,
Sea Lake
,
III.1912
,
J.C.
G., COL- 85894 (
NMVM
)
;
1 specimen
,
Sea Lake
,
Goudie
, 583, COL-85901 (
NMVM
)
.
WESTERN
AUSTRALIA
:
1 specimen
,
Bakers Hill
,
27.III.1973
,
light trap
, D.Weeldenburg (ANIC);
4 specimens
,
Blackwood River
(
ANIC
)
; 2 specimens, Bridgetown, L.J.Newman, F.E.Wilson Collection, COL-85850 (NMVM);
1 specimen
,
Champion Bay
, ex
Collection
V-de-P-,
Sig. K.
vV
. Steinwehr Dec./
Nov.1935
,
Museum Koenig Bonn
, 5368,
H.E.Bomans
BMNH
(E) 1999-248, 405368 (handwritten) (
NHM
)
; 3ƋƋ, Gingin, H.J.Carter Collection P.
20.4.22
., COL-85891 (NMVM);
1 specimen
,
Gray River
,
6.III.1983
, at light,
K. & E.Carnaby
(
ANIC
)
;
1 specimen
,
2 mi
.
NW
Margaret River
,
23.XII.1970
, mv lamp,
G.A.Holloway
&
H.Hughes
, K 293085 (AMS)
;
1 specimen
,
Nannup
,
26.XII.2004
,
D.Lord
, R.deKeyzer
Insect Collection, K
293326 (AMS); 1Ƌ,
Pipehead Dam
,
15 miles
SSE of Armadale
,
26.I.1967
, M.S.Upton (ANIC); 3 specimens, Wilga,
I.1973
and 1 specimen,
21.IV.1973
, K. & Carnaby (ANIC).
Distribution.
Australia
:
Australian
Capital
Territory
,
New South Wales
,
Queensland
,
Western
Australia
,
Victoria
,
South
Australia
. The species is widely distributed in
Australia
, with the exception of the
Northern
Territory
.
Natural history data.
According to label data, adults were collected thoughout the year, mostly from December to April. Most of them were collected at light trap with mercury vapor light; a single specimen was captured in a flight intercept trap (Hann River,
XII.1993
–
I.1994
). Few other specimens were found in rotten logs, in vine scrub. The elevation ranges from
50 to
390 m
. The larvae and host plant(s) are unknown.
Remarks.
Concerning
Figulus lilliputanus
Westwood
and
Figulus clivinoides
Thomson,
Bomans (1986: 9)
wrote that:
“- Il n’est pas absolument certain que ces deux insects soient synonymes. L’examen des types permet de constater que le premier est nettament plus petit que le second, 6,2 et
7,5 mm
pour
8 mm
. De plus
lilliputanus
a été découvert en Australie méridionale et occidentale, et
clivinoides
ou du moins l’espèce la plus grande, en Nouvelle Galle du Sud et au Queensland
” [It is not sure that the two species are synonyms. Examination of types shows that the first is clearly smaller than the second, 6.2 and
7.5 mm
instead of
8 mm
. Moreover
F. lilliputanus
has been discovered in southern and western Australia, and
F. clivinoides
, or at least the largest species, in New South Wales and Queensland]. We dissected the male labeled "Type" of
F. clivinoides
, and we compared the genitalia with those of
F. lilliputanus
. We do not see any significant difference between
F. clivinoides
and
F. lilliputanus
, both in external morphology and in genitalia characters (
e.g.
, in the flagellum shape). Therefore we consider
F. clivinoides
as a junior synonym of
F. lilliputanus
(
new synonymy
).
We found in the collection of NHM a specimen labeled “Tasmania” (A.Simson leg., 8116, 4765, H.E.Bomans), but this locality is most probably erroneous, since we have never seen any other specimens of this species from Tasmania.