Small snails in a big place: a radiation in the semi-arid rangelands in northern Australia (Eupulmonata, Camaenidae, Nanotrachia gen. nov.) Author Köhler, Frank Author Criscione, Francesco text Zoological Journal of the Linnean Society 2013 Zool. J. Linn. Soc. 2013-08-29 169 1 103 123 http://dx.doi.org/10.1111/zoj.12051 journal article 10.1111/zoj.12051 0024-4082 4890431 281CB6ED-063E-4CF0-89A2-80EB3761BCDB NANOTRACHIA CORONATA SP. NOV. Holotype Australia , WA , EK , Osmond Range , 17 km northeast of Palm Yard , narrow limestone and sandstone ridge, with steep open slopes, talus, and Spinifex , in litter under rocks, 17°12′13″S , 128°24′50″E (coll. V. Kessner , 29 August 2009 ); dissected specimen ( WAM S66301 ). Paratypes Same as holotype ; one wet ( WAM S49181 ), 11 dry ( WAM S49086 ) . Other material: Table 1 . Etymology In reference to the crown-like appearance of the peripheral keel because of the axial ribs continuing onto it, derived from ‘coronatus’ (Latin = bearing a crown), adjective of feminine gender. Description Shell ( Figs 4C , 5C, D ): Large; periphery shouldered, strongly keeled; spire weakly elevated; umbilicus open, wide, not concealed by columellar reflection; protoconch and teleoconch with dense, pointed periostracal projections and fine axial growth lines; teleoconch with weak, blunt, widely spaced axial ribs, not interrupted on keel; apertural lip reflected, moderately expanded; yellowish brown. Genitalia: Epiphallus very short, one-quarter of penial length; no muscle fibres connect epiphallus and penial apex. Vas deferens entering penial sheath almost halfway up. Penial wall thin, with two longi- tudinal pilasters, one wide, irregular, comprising half of penial wall, second pilaster thinner, irregularly shaped, winding, with regularly spaced, pointed corrugations; penial pore surrounded by subcircular ridge. Vagina slightly shorter than penis, half to twice as long as free oviduct (identical to N. carinata sp. nov. ; Fig. 6C ). Comparative remarks Largest species of genus. Shell similar to N. carinata sp. nov. , but axial ribs continuing on keel; N. levis sp. nov. differing by absence of keel; N. orientalis with stronger ribs, other congeners with weaker keel. Genitalia identical to N. carinata sp. nov.