Systematics and convergent evolution in three Australian genera of Pepsinae spider wasps (Hymenoptera: Pompilidae) Author Shimizu, Akira urn:lsid:zoobank.org:author:EB6A57A9-63D0-4433-802D-981E54132FFD Department of Biological Sciences, Tokyo Metropolitan University, Hachioji, Tokyo 192 - 0397, Japan & Research Institute of Evolutionary Biology, Inc., Setagaya-ku, Tokyo 158 - 0098, Japan & The University Museum, The University of Tokyo, Bunkyo-ku, Tokyo 113 - 0033, Japan. aquilashimizu7@gmail.com Author Pitts, James P urn:lsid:zoobank.org:author:89B4EC38-0687-4650-9F0E-AF2324459E10 Department of Biology, Utah State University, Logan, UT 84322 - 5305, USA. Author Rodriguez, Juanita urn:lsid:zoobank.org:author:D3E77535-16FA-472A-A686-CE84F8537B55 Australian National Insect Collection, CSIRO, Acton, ACT 2601, Australia. Author Wahis, Raymond Department of Functional and Evolutionary Entomology, Gembloux Agro-Bio Tech, University of Liège, Gembloux 5030, Belgium. Author Yoshimura, Jin Department of Biological Sciences, Tokyo Metropolitan University, Hachioji, Tokyo 192 - 0397, Japan & The University Museum, The University of Tokyo, Bunkyo-ku, Tokyo 113 - 0033, Japan & Department of International Health and Medical Anthropology, Institute of Tropical Medicine, Nagasaki University, Nagasaki 852 - 8523, Japan & Department of Environmental and Forest Biology, State University of New York College of Environmental Science and Forestry, Syracuse, NY 13210, USA & Marine Biosystems Research Center, Chiba University, Uchiura, Kamogawa, Chiba 299 - 5502, Japan. text Austral Entomology 2021 2021-03-17 60 2 301 316 journal article 7644 10.1111/aen.12530 871ea7e5-8005-4c5b-9af9-a247a638aa74 4627871 Evansiclavelia Pitts, Rodriguez & Shimizu , gen. nov . http://zoobank.org/urn:lsid:zoobank.org:act:A2BAB947-2745- 4D26-B8B6-FCB40A34C083 Type species: Evansiclavelia poecilopteryx Pitts, Rodriguez & Shimizu, sp. nov. (here designated). Diagnosis This genus differs from other genera of the Australian ‘Ctenocerini’ sensu Evans (1972) by the following female features: head wider than high ( Fig. 4a ); clypeus much narrower than LID, roughly rectangular in shape (lateral margin vertical and apical margin truncate), mostly flattened and polished; lower frons laterally to antennal sockets and below them deeply depressed with median bridge (frontal bridge) connecting frons and clypeus at the same level; eye broader than half of frons; malar space very long ( Fig. 4a ); gena in dorsal view abruptly receding posteriorly ( Fig. 4b ); and pronotum with anterior face vertical, flattened and polished, forming right angle with dorsum ( Fig. 4c ), latter truncate anteriorly, shallowly depressed anteromedially and posteriorly ( Fig. 4b ). Description Female Vertex moderately convex between eye tops ( Fig. 4a ), distance from eye top to vertex crest, in profile, less than tenth of eye height ( Fig. 4c ). Posterior ocelli at level of eye tops ( Fig. 4a ). Scape curved outward with lateral face concave ( Fig. 4b ). Uppermost portion of occipital suture very high, almost at level of vertex top ( Fig. 4d , arrow). Pronotum with shoulder swollen ( Fig. 4b ). Propodeum with declivity rather flattened but not clearly divided from dorsum ( Fig. 4c,e ). Fore femur thinner than mid femur. Dorsal apex of fore tibia with short stout decurved spines interiorly ( Fig. 4f ) and exteriorly ( Fig. 4g ). All femora, tibiae and undersides of tarsomere 5 lacking spines. Hind tibiae without integumental serrations dorsally. Orbicula very small with a few divergent, straight, weak setae. Tarsal claws unidentate. Transverse groove on S2 almost concealed by posterior margin of S1. S6 compressed laterally with sharp median carina. Male Unknown. Distribution Australia. Etymology The generic name is in honour of the late entomologist Dr H. E. Evans, who was an authority on systematics and behaviour of solitary wasps, especially of spider wasps; and ‘ Clavelia ’ a generic name in the subfamily Ctenocerinae . The name is considered feminine.