Tanaidaceans from Brunei, IV. The Families Kalliapseudidae, Pagurapseudopsidae, Parapseudidae and Apseudidae (Crustacea: Peracarida: Tanaidacea: Apseudomorpha), with descriptions of a new genus and six new species Author Bamber, Roger N. text Zootaxa 2013 3734 4 401 441 journal article 42872 10.11646/zootaxa.3734.4.1 f372aec6-39ab-4836-a1f3-d0eefa9efefd 1175-5326 223309 00DC3ED7-62FD-4D99-ABCC-0DC57D1A51E7 Phoxokalliapseudes gobinae (Bamber, 1999) Kalliapseudes (Kalliapseudes) gobinae Bamber 1999 , 181–184, figs 8–9. Bamber & Sheader 2005, 287 (partim). Phoxokalliapseudes gobinae Drumm & Heard 2011, 38–43, figs 22–24. Bamber et al . 2012, 87. Material : 1♂ , Sample BU3; 2♀♀ (one with oostegites), Sample B12; 2♀♀ (one brooding), Sample B13; all Station B, 04°51′06"N 114°35′26"E , 20 m depth, sand. 5♂♂, 4 brooding ♀♀, 9 neuters, sample DA1; 2♂♂, 2♀♀ (one brooding), sample DA2; both Station DA, 05°54′28.5"N 114°38′23.3"E , 20 m depth, sand. 6♀♀, 1 neuter, Station S14, 04°42′39"N 114°26′24.6"E , 12.7 m depth, sand. 3♀♀, Station S18, 04°41′30"N 114°27′13.5"E , 7.5 m depth, sand. 1♀, Station S17, 04°41′13"N 114°27′23"E , 6.5 m depth, sand. All coll. March 2009 . 2 specimens , 05°21′25"N 111°26′30"E ; 1 specimen , 05°21′08"N 111°26′25"; 4 specimens , 05°20′58"N 111°26′15"E ; 1 juvenile , 1 manca, 05°20′53"N 111°26′21"E ; 3♀♀ (1 brooding), 1 subadult ♂, 1♂ , 2 juveniles , 05°20′17"N 111°27′06"E . All ca 90 m depth, coll. September 2009 . 1 specimen , 05°22′10"N 111°27′00"E ; 2 specimens , 05°21′51"N 111°27′18"E ; 3 specimens , 05°21′59"N 111°27′07"E ; 2♀♀, 05°22′14"N 111°27′04"E ; 3 specimens , 05°22′02"N 111°26′52"E ; 3 specimens , 05°22′14"N 111°26′52"E ; 1 specimen , 05°21′56"N 111°27′22"E ; 3 specimens , 05°21′44"N 111°27′22"E ; 2 specimens , 05°21′44"N 111°27′11"E ; 5 specimens , 05°22′21"N 11°27′30"E ; 1♀, 1 juvenile , 05°22′44"N 111°27′52"E ; 4 specimens , 05°20′17"N 111°27′06"E ; 1 specimen , 05°22′42"N 111°28′46"E . All ca 70 m depth, coll. March 2012 . Remarks : Phoxokalliapseudes gobinae is relatively widespread off the South China Sea coasts of Brunei and Sabah (Bamber 1999; Bamber & Sheader 2005), in the latter case sympatric with Kalliapseudes bamberi Drumm & Heard , 2011, and is a dominant member of the macrobenthos (see Introduction). The present material extends its known depth range to from 6.5 to 90 m . As this species was relatively common, all the samples were analyzed for their population structure, along with data from the type collection (Bamber 1999), collected at 60 m off Brunei . At 20 to 90 m , densities were 7–11 individuals m -2. Densities in shallower-water samples were sparser, ranging from zero to 4 individuals m -2. Males and breeding females (females with either oostegites or brood pouches) were absent at stations shallower than 20 m . The male:female sex ratio at depths from 20 to 90 m ranged from 0.7 to 1.1, not significantly different from 1:1. Bamber et al . (2009) found a 1:1 sex ratio in Cristapseudes omercooperi (Larwood, 1954) in the Eastern Mediterranean; conversely, BłaŻewicz-Paszkowycz & Bamber (2012) found a male:female ratio of 0.4 for Kalliapseudes obtusifrons (Haswell, 1882) in the Bass Strait, Australia . Analysis of the variance and mean of sample densities was used to determine the common dispersion parameter ( k ) of the negative binomial distribution [ k = m 2/( s 2- m ), where m is the mean and s 2 the variance]. The densest populations, at 20 m depth, showed a highly aggregated distribution ( k = 0.22; 1/ k >0); populations shallower than 20 m showed an even distribution ( k = -0.3 to -0.4; 1/ k <0), albeit based on sparse data; populations deeper than 20 m showed a random distribution ( k = -1.4 to -2.5; 1/ k ≈0). Bamber & Marshall (in press) discussed the instability of the shallower inshore seabed off Brunei , which was attributed to physical disturbance from wave action, probably coupled with a greater input of organic matter below the wave-base. Total tanaidacean densities from 2 to 13 m depth were between one half and one-quarter of the average at 20 m depth. Phoxokalliapseudes gobinae thus appears to be widespread and breeding at depths from 20 m and deeper, while its occurrence in the less stable sediments in shallower waters is sparser, involving nonreproductive individuals.