A New Oligocene Flora From Ludvíkovice Near Děčín (České Středohoří Mts., The Czech Republic)
Author
Teodoridis, Vasilis
Author
Kvaček, Zlatko
Author
Radoň, Miroslav
Author
Raprich, Vladislav
Author
Bruch, Angela A.
text
Fossil Imprint
2020
2020-12-30
76
2
297
314
http://dx.doi.org/10.37520/fi.2020.024
journal article
10.37520/fi.2020.024
2533-4069
5387858
Genus
Carya
NUTT., 1818
Carya fragiliformis
(STERNB.) KVAČEK et H.WALTHER
in
Walter and
Kvaček 2007
/
Carya quadrangula
(KIRCHH.)
J.-F.
LEROY
, 1955
Pl. 3, Figs 6–9, Pl. 4, Fig. 1
1825
Phyllites fragiliformis
STERNB.
, p. 42, index iconum, pl. 50, fig. 1.
1936
Caryojuglans quadrangular
KIRCHH.
, pp. 82–85, text-fig. 13, pl. 12, fig. 36a–l.
1955
Carya quadrangular
(KIRCHH.)
J.-F.LEROY, pp. 162–163, text-fig. 81.
2007
Carya fragiliformis
(STERNB.) KVAČEK et H.WALTHER
in Walter and
Kvaček
, p. 110, text-fig. 6b, pl. 11, figs 1–3, pl. 23, figs 8–10.
M a t e r i a l. Incomplete leaflets and one incomplete endocarp, impressions PA 1627, PA 1644, PA 1659, PA 1660, PA 1682 (fruit).
D e s c r i p t i o n. Detached incomplete leaflets, lanceolate to oblanceolate,
35 to 63 mm
long,
12 to 26 mm
wide, apex acute to acuminate, base cuneate with fragmentary petiolule up to
5 mm
, often asymmetrical, margin irregularly serrate, teeth triangular, sharp, bent admedially, venation craspedodromous, to semicraspedodromous, midvein straight or curved, secondaries opposite in the basal part then alternate, originating at 40° to 60°, tertiary veins percurrent, venation of the higher orders regular polygonal reticulate, areolation well-developed, 3 to 4 sided. Incomplete endocarp widely elliptic in outline, apex acute, upper surface and distinct longitudinal ribs are not visible, base not preserved,
22 mm
long and
18 mm
wide, thickness of the nutshell
2.8 mm
, the primary and secondary septum fragmentary. Seed body (kernel) oval incomplete, 2 lateral symmetric parts preserved.
R e m a r k s. The material corresponds to the
holotype
of this fossil species based on a single specimen from Žichov (
Kvaček
and Straková 1997
,
Walther and
Kvaček 2007
). The hickory leaves and/or leaflets described as
C. fragiliformis
or
C. serrifolia
(GÖPP.) KRÄUSEL
are relatively common elements from Oligocene localities in Central Europe, known from e.g., Bechlejovice (
Kvaček
and Walther 2004
), Suletice-Berand (
Kvaček
and Walther 1995
), Seifhennersdorf (
Walther and
Kvaček 2007
) and Matrý (
Kvaček
et al. 2018
). According to
Walther and
Kvaček (2007: 110)
,
C. serrifolia
, which in most cases was assigned to Oligocene – early Miocene material from Central Europe, may represent a descendant of this lineage in the late Neogene.
Koutecký et al. (2019)
associated this type of foliage with fruits of
C. quadrangula
(KIRCHH.)
J.-F. LEROY and wood of
Eucaryoxylon crystallophorum
MÜLL.
- STOLL et MÄDEL following the whole plant concept. The fruit find from Ludvíkovice morphologically corresponds to
C. quadrangula
. On the other hand, some poorly preserved leaf material may show affinity to
Engelhardia orsbergensis
(P.WESSEL et C.O.WEBER) JÄHNICHEN, MAI et H.WALTHER.