Revision of Eupsenella Westwood, 1874 (Hymenoptera, Bethylidae) Author Ramos, Magno S. Author Azevedo, Celso O. text Zootaxa 2012 2012-11-05 3539 1 1 80 http://dx.doi.org/10.11646/zootaxa.3539.1.1 journal article 10.11646/zootaxa.3539.1.1 1175­5334 10096759 0564EF02-FCF9-49A8-BDAC-F432876C16B8 Eupsenella djagaraga Ramos & Azevedo , sp. nov. ( Figs 22 , 67 , 112 ) Description. FEMALE ( holotype ). Body length 7.2 mm . LFW 3.67 mm . Coloration. Head dark castaneous almost black; scape, pedicel and flagellum castaneous; mandible light castaneous with teeth dark castaneous; palpi light castaneous; mesosoma dark castaneous almost black; tegula castaneous; wing venation dark castaneous; procoxa castaneous; mesocoxa, metacoxa and trochanters light castaneous; profemur dark castaneous; mesofemur light castaneous; metafemur castaneous; tarsi light castaneous; metasoma dark castaneous with light castaneous portions along its length. Head ( Fig. 22 ). Head longer than wide, globoid in profile; gena without concavity in profile. Mandible evenly wide, teeth sub-equal, lower tooth curved inward, lower margin strongly setose. Clypeus coriaceous-punctate, median clypeal lobe angulate, median concavity of median clypeal lobe absent, 010 LH, well marked off from frons, median carina present, 0.15 LH. Malar space conspicuous, 0.22 mandibular base. Antenna 1.67 mm ; first five antennal sections in ratio of about 10:9:9:9:9; flagellomere 1 1.0 longer than wide, flagellar pubescence dense, with few outstanding erect setae. Region between torulus and eye straight. Area between the anterior margin of eyes and demarkation of median clypeal lobe 0.07 LH. Eye protuberant, 0.55 LH, with sparse hairs, located more laterally, lower margin without strong depression. Frons coriaceous-punctate. WH 0.91 LH. WF 0.62 WH. WF 1.19 HE. OOL 1.0 WOT. VOL 0.27 HE. Distance from posterior ocellus to vertex crest 0.5 DAO. Ocelli large, DAO 0.13 WF, ocellar triangle not compact, distance between posterior ocelli 2.2 DAO, anterior ocellus not surpassing imaginary top eye line, posterior ocellus almost reaching vertex. Vertex straight, long setae, 0.39 LH, temple profile slightly convergent in dorsal view, with corner angulate, not projected postero-laterad. Hipostomal carina arched, low. Mesosoma ( Fig. 67 ). Pronotal disc confused-rugulose, 0.44 as long as wide, anterior corner rounded, posterior margin concave medially, pronotal disc without short longitudinal furrow. Parapsidal furrow wholly conspicuous. Notauli parallel to each other. Mesoscutum 1.1 length pronotal disc. Scutellum coriaceouspunctate, 1.0 length pronotal disc, almost flat in profile, posterior margin convex medially, scutellar pit almost circular, narrowed, distant each other 8.34 its own diameter. Dorsal axillar surface with fovea oval. Metanotum with median region high, coriaceous-puncticulate, as high as scutellum in profile, with fovea oval laterally. Propodeal disc wider than long; anterior margin without foveolate or transverse furrow; median carina 0.70 length propodeal disc, incomplete posterad, discal carina 0.29 length propodeal disc, converging posterad, straight; space between median and discal carina confused-rugulose. Mesopleuron with subtegular furrow oval; mesepimeral suture almost inconspicuous posterad; mesopleural epicoxal lobe inconspicuous. Propleuron with sequence of foveae from anterior margin of prosternum until cervical collar absent. Prosternum with median excavation complete, median suture present. Mesopectus with posterior depression heart-shaped; median suture evident. Wings. Macropterous. Forewing ( Fig. 112 ) with 1M cell subpentagonal; 1M and 1R1 cells not aligned basally; C cell widening distally; 1R1 cell present, distinctly longer than 2R1 cell, evenly wide; Rs b convex; 2R1 cell 0.15 LFW; distal margin of Rs c convex; r-rs vein strongly inclined; R1 b entirely on anterior margin of forewing; Cu b absent; Rs+M vein 1.0 Rs a ; M vein 0.83 Rs a ; Rs a subperpendicular to anterior margin of forewing; stigma rectangular; distal stigmal margin truncate. Hind wing with four distal hamuli, last hamulus discontinuous each other. Legs. Profemur swollen, 0.5 as wide as long. Metacoxa with median region 0.54 as wide as long. Metacoxal MALE unknown. Material examined. Holotype . AUSTRALIA , Q[ueens]l[an]d, 18.55S 146.10E , 880 m , S 2 M[oun]t[ain] Spec , 9 Mar [ch] 6 Apr [Il] 1995, M.Cermak , F[light] I[ntercept] Trap , JCU, 10 m , ANIC . Paratypes . AUSTRALIA , Q[ueens]l[an]d, 1 ♀ , 19.00S 146.11E , S1 M[oun]t[ain] Spec , 10 Jan [uary]–6 Feb [ruary] 1995, M.Cermak , F[light] I[ntercept] Trap , JCU, (West), ANIC ; 2 ♀ , 9 Mar[ch]–6 Apr[il] 1995, M.Cermak, F[light] I[ntercept] Trap, JCU, (East); 1 ♀ , 5 m , ANIC ; 1 ♀ , 6 Apr[il]– 8 May 1995 , M.Cermak, Malaise trap; 1 ♀ , M.Cermak , F[light] I[ntercept] Trap , JCU, (East), ANIC ; 1 ♀ , 8 May–5 June 1995 , M.Cermak , F[light] I[ntercept] Trap , JCU, (East), ANIC ; 1 ♀ , 5 June–3 July 1995 , M.Cermak , F[light] I[ntercept] Trap , JCU, (West), ANIC ; 1 ♀ , 6 Dec[ember] 1994–10 Jan [uary] 1995, M.Cermak, F[light] I[ntercept] Trap, JCU, 10 m ; 1 ♀ , ANIC ; 1 ♀ , 18.55S 146.10E , 880 m , S2 M[oun]t[ain] Spec, 9 Mar[ch]–6 Apr[il] 1995, M.Cermak, F[light] I[ntercept] Trap, JCU, 10 m ; 1 ♀ , 6 Apr[il]– 8 May 1995 , M.Cermak, Malaise trap; 1 ♀ , 3–31 July 1995 , M.Cermak, Malaise trap; 1 ♀ , 6 Dec[ember] 1994–10 Jan [uary] 1995, M.Cermak, F[light] I[ntercept] Trap, JCU, (West); 1 ♀ , S3 M[oun]t[ain] Spec , 8 May–5 June 1995 , M.Cermak , F[light] I[ntercept] Trap , JCU, (West), ANIC ; 1 ♀ , 17.37S 145.34E , 1000 m , BS3 Massey Cr[ee]k, 1 Dec[ember] 1994–3 Jan [uary] 1995, P.Zoborowski, Malaise trap; 1 ♀ , F[light] I[ntercept] Trap , ANIC ; 2 ♀ , 3 Jan [uary]– 4 Feb [ruary] 1995, Malaise trap , ANIC ; 1 ♀ , 17.28S 145.29E , 1150 m , BS1, Longlands Gap , 2 May–1 June 1995 , P.Zoborowski , Malaise trap , ANIC ; 2 ♀ , 1 Dec [ember] 1994–3 Jan [uary] 1995, P.Zoborowski , F[light] I[ntercept] Trap , JCU, (West); ANIC ; 2 ♀ , 1 Dec [ember] 1994–3 Jan [uary] 1995, P.Zoborowski , F[light] I[ntercept] Trap , ANIC ; 2 ♀ , 17.06S 145.36E , 1150 m , GS1, M[oun]t[ain] Haig , 17 Mar [ch]–6 Apr [il] 1995 P.Zoborowski , Malaise trap , ANIC ; 1 ♀ , 16.30S 145.16E , 9 Km N , M[oun]t[ain] Lewis , 30 Jan [uary] 1998, D.C.F.Rentz , StopA-10, ANIC ; N[ew] S[outh] W[ales], 1 ♀ , 28.22S 153.05E , Brindle C[ree]k, Border Rangers, NP , 3–14 Oct [ober] 1984, I.Naumann , J.Cardale , Malaise trap /ethanol, ANIC ; 1 ♀ , 28.29S 152.24E , Tooloom Plateau , 14 Km W Urbenville , 4–14 Oct [ober] 1984, I.Naumann , J.Cardale , Malaise trap /ethanol, ANIC ; Tas[mania], 1 ♀ , 41.14S 147.56E , 4 Km SE , Weldborough , 13 Jan [uary]–7 Feb [ruary] 1983, I.D.Naumann & J.C.Cardale , Malaise /ethanol, ANIC ; 1 ♀ , 4 Km E Rosebery , 16 Jan [uary]–1 Feb [ruary] 1983, I.D.Naumann & J.C.Cardale , Malaise /ethanol, ANIC ; 2 ♀ , Eugenama , 8 Apr [il] 1988, L. Hill , Euc [ alyptus ] viminalis , forest, ANIC . Distribution. Australia ( Queensland , New South Wales ). Variation. Body length 4.61–6.20 mm ; LFW 3.33–4.04 mm ; head with many hairs; clypeus slightly projected; metasoma dark castaneous. Remarks. This species is similar to E. bilingara sp. nov. by having the area between the anterior margin of eyes and demarkation of median clypeal lobe short, the head subtriangular in profile and vertex straight, the eyes large, the median clypeal lobe angulate. However this species has the median carina of propodeal disc not reaching propodeal declivity, discal carina of propodeal disc short, the Rs c of forewing with distal region subperpendicular, whereas E. bilingara sp. nov. has the median carina of propodeal disc reaching propodeal declivity, discal carina of propodeal disc long, the Rs c of forewing with distal region slightly convex.