High level of endemism in Haiti’s last remaining forests: a revision of Modisimus (Araneae: Pholcidae) on Hispaniola, using morphology and molecules Author Huber, Bernhard A. Author Fischer, Nadine Author Astrin, Jonas J. text Zoological Journal of the Linnean Society 2010 2010-01-25 158 2 244 299 http://dx.doi.org/10.1111/j.1096-3642.2009.00559.x journal article 10.1111/j.1096-3642.2009.00559.x 0024-4082 5438272 MODISIMUS ANGULATUS HUBER & FISCHER SP. NOV. ( Figs 21, 22 , 40 , 129 , 195 ) Type: Male holotype from near La Toma ( 18°27.5 N , 70°07.2 W ), San Cristóbal Prov., Dominican Republic ; degraded forest, 70 m a.s.l. , near ground, 7 November 2005 ( B.A. Huber ), in ZFMK ( DR 3 ) . Etymology: The species name refers to the dorsally angular abdomen. Diagnosis: Medium-sized species with pair of small projections on male chelicerae, provided with weakly modified hairs ( Fig. 129 ; similar to M. femoratus and M. jima sp. nov. ), epigynum very similar to M. femoratus . Distinguished from M. femoratus and M. jima sp. nov. by pattern on sternum (lateral borders of brown marks disrupted; Fig. 22 ), dorsally angular abdomen ( Fig. 21 ), male cheliceral projections slightly wider apart ( Fig. 129 ), and clypeus pattern in males indistinct or even missing. Male ( holotype ): Total length, 2.3; carapace width, 0.9. Leg 1, 20.0 (4.9 + 0.4 + 4.9 + 8.2 + 1.6); tibia 2, 3.0; tibia 3, 2.4; tibia 4, 2.8. Tibia 1 L/d: 55. Habitus as in Figures 21 and 22 , carapace pale ochre-yellow, ocular area darker posteriorly, clypeus with very indistinct pair of brown marks, sternum laterally brown, with a distinctively disrupted lateral pattern ( Fig. 22 ), and whitish medially; legs light brown, and femora and tibiae with light tips and dark subdistal rings; abdomen pale greenish grey, with black spots dorsally and laterally, genital area and area in front of spinnerets light brown. Ocular area elevated; thoracic furrow distinct. PME–PME, 70 Mm; PME diameter, 90 Mm; PME–ALE, 105 Mm; AME–AME, 10 Mm; AME diameter, 15 Mm. Sternum wider than long (0.6/0.4), unmodified. Chelicerae as in Figure 129 , very similar to M. femoratus and M. jima sp. nov. , but frontal apophyses slightly more widely spaced. Palps very similar to M. femoratus (cf. Fig. 124 ). Legs with short spines on femur 1 (two rows, of ~45 and 15 spines each) and femur 2 (two rows, of ~35 spines each), with many short vertical hairs on all femora; curved hairs on tibiae and metatarsi 2 and 3; retrolateral trichobothrium on tibia 1 at 11%; prolateral trichobothrium missing on tibia 1, present on other tibiae; tarsus 1 with ~25 pseudosegments. Variation: Tibia 1 in three other males: 4.7–4.9. Female: In general similar to male; tibia 1 in ten females 3.0–3.4, mean 3.2. Epigynum, similar M. femoratus ( Fig. 40 ); dorsal view not visibly different from M. femoratus (cf. Figs 63 , 126 ). Distribution and habitat: Known from two localities in the south-central Dominican Republic ( Fig. 195 ). In the Santo Domingo botanical garden, this species was found in low vegetation and under the roof of a shelter. Material examined: Dominican Republic : San Cristóbal Prov. , near La Toma , 1♂ , holotype above, together with 1♀ ( ZFMK , DR 3 ) . Distrito Nacional , Santo Domingo , Jardín Botánico ( 18°29.7 N , 69°57.2 W ), forest along brook, 50 m a.s.l. , low vegetation, near ground, 27 November 2005 ( B.A. Huber ), 3♂ , 7♀ and one juvenile ( ZFMK , DR 110 ) ; Santo Domingo , Jardín Botánico , under roof of shelter, 18–22 March 1984 ( H. & L. Levi ), 1♂ and 2♀ ( MCZ 26.666 ) . ‘ Boca Caica’ (= Distrito Nacional : Boca Chica ?) 5–7 March 1955 ( A. M. Nadler ), 4♀ ( AMNH ) .