High level of endemism in Haiti’s last remaining forests: a revision of Modisimus (Araneae: Pholcidae) on Hispaniola, using morphology and molecules Author Huber, Bernhard A. Author Fischer, Nadine Author Astrin, Jonas J. text Zoological Journal of the Linnean Society 2010 2010-01-25 158 2 244 299 http://dx.doi.org/10.1111/j.1096-3642.2009.00559.x journal article 10.1111/j.1096-3642.2009.00559.x 0024-4082 5438272 MODISIMUS SEGUIN HUBER & FISCHER SP. NOV. ( Figs 27–29 , 47 , 70 , 88 , 98 , 148–151 , 199 ) Type: Male holotype from near Seguin, at 18°18.2 N , 72°17.5 W , Dept Sud-Est , Haiti ; 990 m a.s.l. , leaf litter in forest over coffee and banana plantations, 28 November 2007 ( B.A. Huber ), in ZFMK ( Haiti 4a) . Etymology: The species name refers to the type locality; it is used as a noun in apposition. Diagnosis: Medium-sized species with pale callus-like area posteriorly on male abdomen ( Figs 27, 28 ; similar to M. roumaini sp. nov. ), modified hairs of various lengths on male chelicerae ( Figs 88 , 149 ; similar to M. roumaini sp. nov. ); distinguished from M. roumaini sp. nov. by shape of epigynum ( Figs 47 , 150 ; wider scape), internal sclerites in female genitalia ( Figs 70 , 151 ), and numerous spines on male femora. Male ( holotype ): Total length, 3.1; carapace width, 1.35. Leg 1: 33.2 (8.3 + 0.6 + 8.3 + 13.6 + 2.4); tibia 2, 5.5; tibia 3, 4.4; tibia 4, 5.9. Tibia 1 L/d: 67. Habitus as in Figures 27–29 , carapace pale ochre-white, with wide brown lateral margins posteriorly, thoracic furrow also dark brown; ocular area light brown, clypeus with pair of lateral brown bands; sternum medially brown with light spot behind labium, and pale ochre-yellow to whitish laterally; legs light brown, tips of femora and tibiae whitish, darker rings subdistally on femora (indistinct on tibiae); abdomen bluish grey, dorsally (except heart area) and laterally densely covered with black spots, with some small light-bluish spots forming disrupted lines, with distinctive pale area posteriorly; genital area and area in front of spinnerets light brown, bluish spot in-between. Ocular area strongly elevated; thoracic furrow distinct. PME–PME, 150 Mm; PME diameter, 135 Mm; PME–ALE, 175 Mm; AME–AME, 20 Mm; AME diameter, 25 Mm. Sternum wider than long (0.9/ 0.7), unmodified. Chelicerae with ~30 modified hairs on each side, mediodistal hairs clearly longer than the others ( Figs 88 , 149 ). Palps as in Figure 148 , coxa with retrolateral apophysis, femur with rounded proximal and pointed distal ventral apophyses; procursus without dorsal spine-like process ( Fig. 98 ), curved towards femur distally, bulb with large, curved apophysis, and complex membranous subdistal projections. Legs with spines on femora 1 and 2; femur 1 with prolateral row (~30 spines not reaching tip) and retrolateral row (~20 very weak spines not reaching tip); femur 2 with four rows, prolateral (~25 spines, not reaching tip), prolateroventral (18 spines), retrolateroventral (20 spines), and retrolateral (~20 spines, almost until tip); all femora with many short vertical hairs; curved hairs on tibiae 1–3 and metatarsi 2–4; retrolateral trichobothrium of tibia 1 at 11%; prolateral trichobothrium missing on tibia 1, present on all other tibiae. Tarsus 1 with ~30 pseudosegments. Variation: Tibia 1 in four other males: 7.6, 7.9, 8.2, 8.5. Female: In general similar to male, but without callus-like area posteriorly on abdomen. Tibia 1 in six females : 5.9–6.6 (mean 6.2). Epigynum, with tongueshaped scape, and distinctive dark sclerites ( Figs 47 , 150 ); dorsal view as in Figures 70 and 151 . Distribution and habitat: Known from type locality only ( Fig. 199 ). This species was found in the same piles of dead banana leaf bases on the ground as M. epepye sp. nov. , but in the dryer upper parts. Material examined: Haiti : Dept Sud-Est , near Seguin, 1♂ , holotype above; same data, 5♂ , 7♀ and four juveniles ( ZFMK , Haiti 4/5); same data, 3♀ and two juveniles , in pure ethanol ( ZFMK , Haiti 81) .