Systematics of Phasia Latreille (Diptera: Tachinidae) Author Sun, Xuekui Author Marshall, Stephen A. text Zootaxa 2003 2003-08-29 276 1 1 320 https://biotaxa.org/Zootaxa/article/view/zootaxa.276.1.1 journal article 10.11646/zootaxa.276.1.1 1175­5334 5019510 4177F6E7-B276-438E-8E49-EE5ECD84D351 3.5.9 Phasia punctigera ( Townsend, 1891 ) (Figures I­46, II­5.4) Hyalomyia punctigera Townsend, 1891: 135 . Phoranthella morrisoni Townsend, 1915: 23 , nomen nudum . Phoranthella morrisoni Brooks, 1945: 673 (redescription). Phoranthella punctigera : Sabrosky and Arnaud 1965: 970 (catalog). Phasia punctigera : O’Hara and Wood 1998: 765 (new combination). DESCRIPTION Body length: 4­7 mm . MALE: HEAD. Head spherical or nearly so. Eyes separated by a distance as wide as single ocellus. Fronto­orbital plate black, silky grey pruinose. Frontal vitta black. Frontal vitta at base of antennae 1.7­2.0 times as wide as fronto­orbital plate anteriorly. Ocellar setae present but fine; outer vertical setae absent; inner vertical setae absent. Face black with grey pruinosity. Parafacial black with grey pruinosity, bare, nearly as wide as first flagellomere. Lower margin of face projecting, visible in profile, black with grey pruinosity. Vibrissa not differentiated; intervibrissal distance 1.1 times distance between vibrissa and eye on same side; facial ridge with bristles on lower than 1/4. Gena black with grey pruinosity; hairs white; height 0.1­0.12 times eye height. Lunule black and shining; sublunular bulla indistinct. Antenna black; first flagellomere 1.1 times as long as pedicel; arista thickened on basal 0.25. Length of oral opening 2­2.2 times its width. Occiput flattened, white pruinose; hairs black. Palpus black. THORAX . Mesoscutum thinly pruinose, without black longitudinal vitta and fine black hairs. 0+0(1) acrostichal seta; 0+1 dorsocentral seta; 1­2 postpronotal setae; presutural supra­alar seta absent, or present, but fine (rarely); 1 postsutural intra­alar seta; 2 notopleural setae; 1 supra­alar seta; 2 postalar setae. Pleuron thinly grey pruinose; hairs black; scale­like hair absent. Anepimeral setae hair­like, black; 1 katepisternal seta; 6­8 meral setae. Scutellum black, shining, without pruinosity, with two pairs of marginal setae; apical setae present and distinct; discal setae randomly arranged, not in rows. Subscutellum not very prominent. WING . Lower calypter hyaline white. Wing base without scalelike setae. Tegula black. Basicosta black. Wing hyaline; petiole of apical cell 0.35­0.5 times as long as preceding section of R 4+5 ; M meeting R 4+5 almost at right angle. Halter brown. LEGS . Fore femur black (swollen); hairs black. Fore tibia black, without bristles, apically with 1 pv and 1 d . Fore tarus black. Fore claws brown, apex black; as long as fifth tarsomere. Pulvilli black. Mid femur black; hairs black. Mid tibia black, with 1 ad , 1 p and 1 v , apically with 1 ad , 1 av , 1 pv , 1 p , 1 v and 1 d . Hind femur swollen on 2/5 posteriorly, black; hairs black. Hind tibia strongly arched, distinctly shorter than hind femur, black, with 3­4 ad and 2­3 pd . ABDOMEN . Abdominal tergites black; pruinosity present, but thin; longitudinal vitta absent. Relative length of abdominal tergites I+II:III:IV:V = 1.3:1:1:1.3; relative width I+II:III:IV:V = 0.5:1:1:0.7. Syntergite I+II shiny, not pruinose. Pruinosity of tergite III evenly. Pruinosity of tergites IV, V silvery; hair spots distinct. TERMINALIA . Syncercus V­notched posteriorly, not bent. Surstylus slightly bent dorsally, longer than cercus. Ejaculatory apodeme small, knob­like. Hypandrium longer than phallapodeme. Pregonite reduced, with two stout teeth. Postgonite short, as long as pregonite. Phallus slender, 1.3 times as long as hypandrium, not haired. Distiphallus well sclerotized. FEMALE . Wing hyaline. Eyes touching. Anterior spine­like setae of hind tibia absent. Sternite V extremely long, longer than total length of previous sternites. Sternite VII (sheath) shorter than sternite VI, shovel­like, narrow apically, bent, apex directed ventrally, sternite VII with linear wrinkles ventrally, median area membranous. Ovipositor bent upward. TYPE MATERIAL Phoranthella morrisoni Townsend. Holotype , [ USA ], Georgia (USNM, No. 19139, examined). OTHER MATERIAL EXAMINED CANADA . Quebec : Lac Lauzon, Mt. Tremblant Pk., 28.viii.1956 , H.C. Huckett ( 1♂ , USNM). USA . Alabama : Montgomery, 22.xi.1946 , H.R. Dodge ( 1♂ 1♀ , USNM). Arizona : Cochise Co., Southernwestern Research Station, 5mi , 5,400ft , W. Portal, 1­ 9.viii.1956 , E. Ordway, C. and M. Cazier ( 6♂ 1♀ , AMNH); Huachuca Mts., Sierra Vista ( 15mi S), Ramsey Cyn., 5,200ft , 7.iv.1967 , R.F. Sternitzky ( 2♂ , CNCI); Near Patagonia, 6.x.1951 , E.C. Zimmerman ( 1♀ , BMNH); Ramsey Cn., Huachuca Mts., 25.iii.1967 , 5,500ft , D.M. Wood ( 1♂ , CNCI). Arkansas : Perry Co., 19.v.1956 , ( 1♀ , USNM); Washington Co., 29.v.1956 ( 1♀ , USNM). California : Santa Clara Co., Baker ( 1♀ , BMNH). Colorado : Pingree Park, 17.viii.1932 , C.C. Fluke ( 1♂ , AMNH); Golden, Mt. Vernon Cn., 7,200ft , 31.vii.1961 , C.H. Mann ( 2♂ , CNCI); Nederland, Caribou , 8,700ft , 7.viii.1961 , J.E.R. Stainer ( 1♂ , CNCI); [?], ( 1♂ , BMNH). District of Columbia : Washington, D.C., Eastern Brch and Rock Creek, 2.vi.1917 and 22.x.1914 , C.H.T. Townsend and R.C. Shannon ( 1♂ 2♀ , USNM). Idaho : Boise, J.M. Aldrich ( 1♂ , USNM); Moscow, 1­12.ix.1908 , J.M. Aldrich ( 5♂ 5♀ , USNM); Waha, 30.v.1924 , A.L. Melander ( 1♀ , USNM). Indiana : La Fayett, 22.x. and 19.x.1915 , D.G. Hall and J.M. Aldrich ( 1♂ 1♀ , USNM); Vinta, 7­ 8.vi.1899 , Wickham ( 1♂ , BMNH). Kansas : Manhattan, 15.vi.1923 , W. V . Hedding ( 1♂ , KSUC). Maryland : Beltsville, 30.vii.1916 , W.R. Walton ( 1♀ , USNM); July ( 1♂ , BMNH); Grove Hill, 2.xi.1916 , F.L. Aster ( 1♀ , USNM); [?], ( 1♂ , USNM); Leonardtown, 10.vii.1952 , O.L. Cartwright ( 1♀ , USNM). Massachusetts : Woods Hole, 13.ix.1912 , C.T. Greene ( 1♀ , AMNH); N Andover, 16.vii.1911 , J.D. Tothill ( 1♂ , CNCI). Mississippi : Aberdeen, 4.v.1921 , H.C. Dozier ( 1♀ , USNM); Agri. Col., 30.xi.1921 , 22.iv.1922 , H.W. Allen ( 1♂ 1♀ , USNM); A & M. Col. 21.xi.1924 , 15.v.1925 , H.W. Allen ( 1♂ 1♀ , USNM). Nevada : [?], 6.vii., Baker ( 1♀ , USNM). New Jersey : Hornerstown, 14.v.1910 ( 1♀ , AMNH); Lona, 16.v.1909 ( 1♂ , USNM); Paterson, 4.x.1915 ( 1♂ , USNM). New Mexico : Las Cruces, 31.viii. and 11.viii, C.H.T. Townsend ( 2♂ , USNM, BMNH); Roosevelt Co., Portales, 8.iv.1988 , N. Jorgensen ( 1♀ , CNCI). North Carolina : Tuckasegee, 2,200ft , 30.viii.1957 , J.G. Chillcott ( 1♀ , CNCI); Valley of Black Mts., 25.viii.1906 , W. Beutenmuller ( 1♂ , AMNH); Wayne Co., 15.vi.1955 ., H. V . Weems, Jr. ( 2♂ , USNM). Oklahoma : Marshall Co., Willis, 2­12.vi.1986 , D.J. Peckham ( 70♂ 27♀ , USNM). Oregon : Independence, 13.vi.1934 , N.P. Iarson ( 1♀ , USNM). South Carolina : Fairfax, 21.v.1931 (2?), E.W. Howe ( 2♂ 1♀ , CUIC); St. George, 1.vi.1853 , O.L. Cartwright ( 2♂ , USNM). Texas : Austin, A.L. Melander ( 2♂ 1♀ , USNM); Brazos Co., College Station, iv­v. and x.1917 ­ 1966, H.J. Reinhard, D.M. Wood ( 48♂ 29♀ , CNCI, USNM, CUIC, TAMU); Burleson., 25.ix.1939 , H. Menusan ( 1♀ , CNCI); Comal River, 24.iii.1942 , A.L. Melander ( 1♂ , USNM); Dallas, Osten­Sacken ( 2♂ , DEIC); Dallas, Bishopp, 19.iv.1916 , J.M. Aldrich ( 2♂ 2♀ , USNM); Denton, 27­29, vi., C.R. Jones, F.C. Bishopp ( 2♀ , USNM); Donna, 22.x.1933 . J.W. Monk ( 1♀ , TAMN); Ft. Davis ( 21mi W), 22­25.vii.1956 and 26.v.1901 , E.G. Matthews etc. ( 3♂ , CNCI, USNM); Kerrville, 31.iii­3iv.1959 , J.F. McAlpine ( 21♂ 1♀ , CNCI); Nacogdoches Co., 16.v.1941 , H.J. Reinhard ( 2♂ , CNCI); Neuecest, 25.ix. 1896 , Marlatt ( 1♂ , USNM); Pleasanton, 22.iii.1937 ( 1♂ , TAMU); San Antonio, 8,16. ix.1942 and 18.v.1918 , A. Wetmore and A.L. Melander ( 2♂ 1♀ , USNM); Dimmit Co., 28.ii.1933 ( 1♂ , TAMU). Utah : Salt Lake, 24.vi.1922 , A.L. Lovett ( 1♀ , USNM). Virginia : Falls Church, 20.ix.1912 , C.T. Greene ( 1♀ , USNM); [?], 30.x.1926 , N.K. Bigerow ( 1♀ ,CNCI). Washington : Benton Co., Hanford Site, ALE, Rattlesnake Ridge, 29.viii.1994 , R.S. Zack ( 1♂ , CNCI); Yakima Co., Wenas Creek For. Camp. 20mi NW Selah, 2,550ft , 25.v. 1986 , N.E. Woodley ( 1♀ , USNM); [?], ( 1♂ , USNM); Pullman, 16.vii.1908 , W.M. Mann ( 1♂ , USNM). Doubtful State : "Osten­Sacken Collection ( 1♂ , DEIC)". MEXICO . Chih., 12mi NW Gran Morelos, 15.viii.1950 , R.F. Smith ( 1♂ , AMNH). HOST Unknown. NOTES The type of Hyalomyia punctigera was not located but the description fits this species better than any other. Phoranthella morrisoni Townsend was erected for a specimen determined by Coquillett as Phorantha occidentis (Walker) , but it remained undescribed until 1938 ( Brooks 1945: 673 ).