A Revision of the Neotropical Goblin Spider Genus Neoxyphinus Birabén, 1953 (Araneae, Oonopidae)
Author
Abrahim, Naiara
Author
Brescovit, Antonio D.
Author
Rheims, Cristina A.
Author
Santos, Adalberto J.
Author
Ott, Ricardo
Author
Bonaldo, Alexandre B.
text
American Museum Novitates
2012
2012-05-16
2012
3743
1
76
http://www.bioone.org/doi/abs/10.1206/3743.2
journal article
10.1206/3743.2
0003-0082
5360343
Neoxyphinus keyserlingi
(Simon)
,
new combination
Figures 6, 18, 24, 30, 40, 48, 313–344; map 3
Dysderina keyserlingi
Simon, 1907: 252
(male [reported female]
holotype
from
Therezopolis
[
Teresópolis
, state of
Rio de Janeiro
,
Brazil
],
Brunet
col., PBI_
OON 568
, in
Muséum d’Histoire Naturelle
,
Paris, AR
5773, examined by
N.I. Platnick
).
Dysderina rugosa
Bristowe, 1938: 69
(female
holotype
from
Brazil
[probably southeastern
Brazil
],
W.S. Bristowe
col., 1923, PBI_
OON 567
, deposited in the
Natural History Museum
,
London
, examined by
N.I. Platnick
). NEW SYNONYMY.
DIAGNOSIS: This species is readily recognized by the evenly granulated carapace surface (figs. 315, 333, 338) and by a wrinkled sternal median field (figs. 334, 339) in both males and females. Males are unique by the embolus with a prolateral sulcus and apical copulatory opening (figs. 48, 327). Females resemble those of
N. hispidus
and
N. furtivus
by the wide epigynal atrium, with an anteriorly positioned median element, differing by the carapace microsculpture and by the bilobated epigynal median element (fig. 344).
MALE (PBI_OON 14482, IBSP 62132): Total length 2.16. Carapace dark red-brown, posterolateral surface without spikes, surface of elevated portion of pars cephalica and sides strongly granulate, lateral margin without tubercles. Clypeus margin unmodified, straight in frontal view. Eyes: ALE separated by their radius to diameter. Sternum as long as wide, dark red-brown, median concavity absent, with a median wrinkled field, furrows deep, anterior margin with continuous transverse groove, setae evenly scattered, originating from small tubercles. Book lung covers large, ovoid. Pedicel tube medium sized. Dorsal scutum dark red-brown, middle and sides surface smooth, anterior half without projecting denticles. Postepigastric scutum dark red-brown. Legs orange-brown. Spine formula: tibia I v5-3-2, metatarsus I v2-2-1; tibia II v2-4-1, metatarsus II v2-2-0. Genitalia: sperm pore situated at level of anterior spiracles. Palp proximal segments, cymbium, and bulb yellow. Embolus with a prolateral sulcus and apical copulatory opening, without prolateral prong or prolateral lamella (figs. 323–331).
FEMALE (PBI_OON 14483, IBSP 62115): Total length 2.33. Spine formula: tibia I v4-4-2, metatarsus I v2-2-1; tibia II v4-4-1, metatarsus II v3-2-0. Epigynal atrium wide, with an anteriorly positioned, bilobated median element (fig. 344).
DISTRIBUTION: Known from southeastern
Brazil
(map 3).
OTHER MATERIAL EXAMINED:
BRAZIL
:
Espírito Santo
:
Linhares
: Reserva Florestal Vale do
Rio Doce
,
19°23′30″S
40°04′20″W
,
June 1997
, J. Raizer (PBI_OON 14692, IBSP 14881)
1♂
.
São Paulo
:
São Luis
do Paraitinga
: Núcleo Santa Virginia, Parque Estadual Serra do Mar,
23°20′28″S
45°08′49″W
,
Apr. 10, 2005
, M.U. Prado (PBI_OON 14688, IBSP 58438)
1♀
;
May 22, 2005
(PBI_OON 14687, IBSP 58436; PBI_OON 14689, IBSP 58439)
2♀
.
São Paulo
: Parque do Estado,
23°32′25″S
46°37′48″W
, all collected by J.R. Valvassori,
Feb. 28–Mar. 06, 2002
(PBI_OON 11262, IBSP 59156)
1♂
,
Jan. 21–28, 2003
(PBI_OON 11283, IBSP 68150; PBI_ OON 11306, IBSP 68132)
2♂
,
1♀
,
Apr. 07–14, 2003
(PBI_OON 11276, IBSP 68129; PBI_OON 11278, IBSP 68128; PBI_OON 11281, IBSP 68153; PBI_OON 11284, IBSP 68134; PBI_OON 11298, IBSP 68131; PBI_ OON 11301, IBSP 68152; PBI_OON 11304, IBSP 68151)
8♂
,
1♀
,
July 16–23, 2003
(PBI_OON 11285, IBSP 68133; PBI_OON 11293, IBSP 59166; PBI_OON 11299, IBSP 68154; PBI_OON 11300, IBSP 68126)
5♂
,
1♀
,
Oct. 18–25, 2003
(PBI_OON 11277, IBSP 68130; PBI_OON 11279, IBSP 68125; PBI_OON 11280, IBSP 68139; PBI_OON 11282, IBSP 68146; PBI_OON 11286, IBSP 68143; PBI_OON 11287, IBSP 68144; PBI_
Figs. 313–322.
Neoxyphinus keyserlingi
(Simon)
, male.
313.
Habitus, dorsal view.
314.
Same, lateral view.
315.
Carapace, dorsal view.
316.
Sternum and mouthparts, ventral view.
317.
Carapace, anterior view.
318.
same, posterior view.
319.
Abdomen, anterior view.
320.
Same, dorsal view.
321.
Same, ventral view.
322.
Same, lateral view.
Figs. 323–331.
Neoxyphinus keyserlingi
(Simon)
, male.
323.
Left male palp, prolateral view.
324.
Same, ventral view.
325.
Same, retrolateral view.
326.
Left embolus, prolateral view.
327.
Right embolus, apical view.
328.
Left embolus, retrolateral view.
329.
Same, prolateral view.
330.
Same, ventral view.
331.
Same, retrolateral view.
OON 11288,
IBSP 68147
;
PBI
_OON 11289,
IBSP 68145
;
PBI
_OON 11290,
IBSP 68138
;
PBI
_OON 11291,
IBSP 68148
;
PBI
_OON 11292,
IBSP 68137
;
PBI
_OON 11295,
IBSP 68149
;
PBI
_OON 11296,
IBSP 68127
;
PBI
_OON 11297,
IBSP 68140
;
PBI
_OON 11302,
IBSP 68141
;
PBI
_OON 11303,
IBSP 68142
;
PBI
_OON 11305,
IBSP 68135
)
17♂
,
4♀
;
Cidade Universitária Armando de Salles Oliveira
–CUASO,
Universidade
de
São Paulo
,
23°33′S
46°43′W
, all collected by
D.F. Candiani
, 1999 (
PBI
_OON 11223,
IBSP 69001
;
PBI
_OON 11224,
IBSP 68990
;
PBI
_OON 11227,
IBSP 68986
;
PBI
_OON 11228,
IBSP 69017
;
PBI
_OON 11230,
IBSP 68983
;
PBI
_OON 11232,
IBSP 68994
;
PBI
_OON 11234,
IBSP 69021
;
PBI
_OON 11235,
IBSP 68992
;
PBI
_OON 11237,
IBSP 69016
;
PBI
_OON 11238,
IBSP 68988
;
PBI
_OON 11239,
IBSP 69022
;
PBI
_OON 11240,
IBSP
Figs. 332–335.
Neoxyphinus keyserlingi
(Simon)
, male.
332.
Habitus, lateral view.
333.
Carapace, dorsal view.
334.
Sternum and mouthparts, ventral view.
335.
Mouthparts, posterior view.
69005;
PBI
_OON 11241,
IBSP 69003
;
PBI
_OON 11242,
IBSP 68989
;
PBI
_OON 11243,
IBSP 69013
;
PBI
_ OON 11244,
IBSP 68985
;
PBI
_OON 11246,
IBSP 69023
;
PBI
_OON 11247,
IBSP 68997
;
PBI
_OON 11248,
IBSP 69025
;
PBI
_OON 11249,
IBSP 68996
;
PBI
_OON 11250,
IBSP 68999
;
PBI
_OON 11254,
IBSP 69009
;
PBI
_OON 11255,
IBSP 69008
;
PBI
_OON 11256,
IBSP 69026
;
PBI
_OON 11258,
IBSP 69020
)
14♂
,
12♀
,
June 11–18, 1999
(
PBI
_OON 11229,
IBSP 69024
;
PBI
_OON 11231,
IBSP 68995
;
PBI
_OON 11233,
IBSP 69012
;
PBI
_OON 11245,
IBSP 69015
;
PBI
_OON 11252,
IBSP 68993
)
2♂
,
3♀
,
Aug. 16–23, 1999
(
PBI
_OON 11273,
IBSP 42179
;
PBI
_OON 11274,
IBSP 42178
;
PBI
_OON 11275,
IBSP 42181
;
PBI
_OON 14691,
IBSP 42180
)
4♂
,
1♀
,
Nov. 16–23, 1999
(
PBI
_OON 14509,
IBSP 42192
)
1♂
,
Dec. 12–19, 1999
(
PBI
_OON 11222,
IBSP 68982
;
PBI
_OON 11225,
IBSP 69011
;
PBI
_OON 11226,
IBSP 69000
;
PBI
_OON 11236,
IBSP 68984
;
PBI
_OON 11251,
IBSP 68991
;
PBI
_OON 11253,
IBSP 69014
;
PBI
_OON 11257,
IBSP 69002
)
8♂
;
Mata da Biologia
,
Universidade
de
São Paulo
, all collected by
D.F. Candiani
, 1999 (
PBI
_OON 14485,
IBSP 69007
;
PBI
_OON 14490,
IBSP 69018
;
PBI
_OON 14516,
IBSP 69010
;
PBI
_OON 14522,
IBSP 69006
;
PBI
_OON 14552,
IBSP 68987
)
2♂
,
5♀
,
Jun. 11–18, 1999
(
PBI
_OON 14489,
IBSP 69004
;
PBI
_OON 14502,
IBSP 69019
)
1♂
,
1♀
,
Dec. 12–19, 1999
(
PBI
_OON 14521,
IBSP 68998
)
2♂
;
Horto Oswaldo
Cruz, Campus Instituto Butantan, D.F. Candiani et al., 2000
–2001 (
PBI
_OON 11261,
IBSP 117031
)
2♂
,
2♀
;
Jardim Angela
,
Reservatório Guarapiranga
,
23°43′51″S
46°44′23″W
,
R.P. Indiicatti
,
Sep. 09, 1999
–
May 12, 2000
(
PBI
_OON 11259,
IBSP 68041
;
PBI
_OON 11260,
IBSP 68042
)
13♂
,
14♀
,
Nov. 06–12, 1999
(
PBI
_OON 11569,
IBSP 131858
)
1♂
,
1♀
,
I. Cizauskas
and
C.R.M Garcia
,
Apr. 07–13, 2005
(
PBI
_OON 11307,
IBSP 62098
)
1♂
;
Parque
Ilha dos Eucaliptos
,
Reservatório Guarapiranga
,
23°44′14″S
46°44′09″W
, all collected by
I. Cizauskas
and
C.R.M. Garcia
,
Oct.
Figs. 336–344.
Neoxyphinus keyserlingi
(Simon)
, female.
336.
Habitus, dorsal view.
337.
Same, lateral view.
338.
Carapace, dorsal view.
339.
Sternum and mouthparts, ventral view.
340.
Carapace, posterior view.
341.
Abdomen, dorsal view.
342.
Same, lateral view.
343.
Same, ventral view.
344.
Epigynum, dorsal view.
07–13, 2003 (PBI_OON 11310, IBSP 62133; PBI_OON 11313, IBSP 62141; PBI_OON 11315, IBSP 62117; PBI_OON 11318, IBSP 62119; PBI_OON 11319, IBSP 62159; PBI_OON 11321, IBSP 62125; PBI_OON 11324, IBSP 62140; PBI_OON 11325, IBSP 62161; PBI_OON 11329, IBSP 62160; PBI_OON 11333, IBSP 62100; PBI_OON 11334, IBSP 62153; PBI_OON 14553, IB62152)
4♂
,
9♀
,
Apr. 07–13, 2004
(
PBI
_OON 11309,
IBSP 62144
;
PBI
_OON 11314,
IBSP 62103
;
PBI
_OON 11320,
IBSP 62148
;
PBI
_OON 11322,
IBSP 62138
;
PBI
_OON 11323,
IBSP 62137
;
PBI
_OON 11326,
IBSP 62102
;
PBI
_OON 11327,
IBSP 62128
;
PBI
_ OON 11328,
IBSP 62145
;
PBI
_OON 11330,
IBSP 62146
;
PBI
_OON 11332,
IBSP 62154
;
PBI
_OON 11335,
IBSP 62147
;
PBI
_OON 14482,
IBSP 62132
;
PBI
_OON 14483,
IBSP 62115
;
PBI
_OON 14484,
IBSP 62155
;
PBI
_OON 14486,
IBSP 62151
;
PBI
_OON 14487,
IBSP 62105
;
PBI
_OON 14488,
IBSP 62112
;
PBI
_OON 14491,
IBSP 62131
;
PBI
_OON 14492,
IBSP 62150
;
PBI
_OON 14493,
IBSP 62129
;
PBI
_OON 14508,
IBSP
62149;
PBI
_OON 14518,
IBSP 62123
;
PBI
_OON 14520,
IBSP 62107
;
PBI
_OON 14495,
IBSP 62110
;
PBI
_ OON 14496,
IBSP 62134
;
PBI
_OON 14497,
IBSP 62158
;
PBI
_OON 14498,
IBSP 62124
;
PBI
_OON 14499,
IBSP 62114
;
PBI
_OON 14500,
IBSP 62142
;
PBI
_OON 14501,
IBSP 62099
;
PBI
_OON 14503,
IBSP 62121
;
PBI
_OON 14504,
IBSP 62162
;
PBI
_OON 14506,
IBSP 62120
;
PBI
_OON 14507,
IBSP 62104
;
PBI
_OON 14511,
IBSP 62106
;
PBI
_OON 14512,
IBSP 62101
;
PBI
_OON 14513,
IBSP 62130
;
PBI
_OON 14514,
IBSP 62109
;
PBI
_OON 14515,
IBSP 62156
;
PBI
_OON 14517,
IBSP 62135
;
PBI
_OON 14519,
IBSP 62111
;
PBI
_ OON 14523,
IBSP 62139
;
PBI
_OON 14524,
IBSP 62126
;
PBI
_OON 14551,
IBSP 62113
)
35♂
,
25♀
,
Oct. 07–13, 2004
(
PBI
_OON 11311,
IBSP 62118
;
PBI
_OON 11312,
IBSP 62157
;
PBI
_OON 11317,
IBSP 62122
)
3♀
,
Apr. 07–13, 2005
(
PBI
_OON 11316,
IBSP 62163
;
PBI
_OON 11331,
IBSP 62116
;
PBI
_OON 11336,
IBSP 62143
)
3♂
.
Itapevi
:
Condomínio Residencial TransUrb
,
23°32′57″S
46°56′02″W
, all collected by
V.C. Onofrio
,
June 25, 1999
(
PBI
_OON 11263,
IBSP 67562
;
PBI
_OON 11264,
IBSP 67594
;
PBI
_OON 11265,
IBSP 67580
;
PBI
_OON 11267,
IBSP 67573
;
PBI
_OON 14694,
IBSP 67608
)
3♂
,
3♀
,
Nov. 20, 1999
(
PBI
_OON 11269,
IBSP 67600
)
1♀
,
Dec. 17, 1999
(
PBI
_OON 11266,
IBSP 67583
)
1♀
.
Miracatu
:
Fazenda Itereí
,
24°03′S
47°13′W
,
Equipe Biota
,
Sep. 20, 2001
(
PBI
_OON 11559,
IBSP 136432
;
PBI
_OON 11560,
IBSP 136433
;
PBI
_OON 11561,
IBSP 136431
;
PBI
_OON 11562,
IBSP 136429
;
PBI
_OON 11563,
IBSP 136428
;
PBI
_OON 11564,
IBSP 136427
;
PBI
_OON 11565,
IBSP 136430
)
6♂
,
5♀
.
Peruibe
:
Estação Ecológica de Juréia–Itatins
,
24°18′47″S
47°00′03″W
,
Mar. 17–21, 1997
,
A.D. Brescovit
et al. (
PBI
_OON 14693,
IBSP 9605
)
1♂
,
2♀
.
Iporanga
:
Parque Estadual Turístico do Alto Ribeira
–PETAR,
24°28′30″S
48°39′30″W
,
May 02, 2002
,
R. Andrade
(
PBI
_OON 14690,
IBSP 44806
)
1♂
(
PETAR
,
Núcleo Santana
),
Equipe Biota
,
Dec. 08–12, 2002
(
PBI
_OON 11566,
IBSP 136520
;
PBI
_OON 11567,
IBSP 136521
;
PBI
_OON 11568,
IBSP 136519
)
3♂
.