<document id="98D7A79E5770388DD8527EBB1B439505" ID-DOI="10.11646/zootaxa.2851.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5286934" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629721501414" checkinUser="felipe" docAuthor="Blahnik, Roger J. &amp; Holzenthal, Ralph W." docDate="2011" docId="03BE8797007FFF9A98B1FB47FD8EC438" docLanguage="en" docName="zt02851p075.pdf" docOrigin="Zootaxa 2851 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2851.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Mortoniella punensis" docType="treatment" docVersion="5" lastPageNumber="47" masterDocId="FF87FFEF0053FFB49826FF89FFBAC62F" masterDocTitle="Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851" masterLastPageNumber="75" masterPageNumber="1" pageNumber="45" updateTime="1699101759394" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="3B1F6C865DC0FEE3D7110A8D60B31164">Revision of the austral South American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae) 2851</mods:title>
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<mods:namePart id="07B5260434D6AF3C360372E49A091EA3">Blahnik, Roger J.</mods:namePart>
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<subSubSection id="C30D650A007FFF9898B1FB47FD82C2C7" box="[151,568,1230,1256]" pageId="44" pageNumber="45" type="nomenclature">
<paragraph id="8BA83681007FFF9898B1FB47FD82C2C7" blockId="44.[151,568,1230,1289]" box="[151,568,1230,1256]" pageId="44" pageNumber="45">
<heading id="D0E081ED007FFF9898B1FB47FD82C2C7" bold="true" box="[151,568,1230,1256]" fontSize="11" level="1" pageId="44" pageNumber="45" reason="1">
<taxonomicName id="4C174D02007FFF9898B1FB47FD82C2C7" authority="(Flint, 1983)" baseAuthorityName="Flint" baseAuthorityYear="1983" box="[151,568,1230,1256]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="44" pageNumber="45" phylum="Tracheophyta" rank="species" species="punensis">
<emphasis id="B963EA93007FFF9898B1FB47FD82C2C7" bold="true" box="[151,568,1230,1256]" pageId="44" pageNumber="45">
<emphasis id="B963EA93007FFF9898B1FB47FE22C2C7" bold="true" box="[151,408,1230,1256]" italics="true" pageId="44" pageNumber="45">Mortoniella punensis</emphasis>
(
<bibRefCitation id="EF864B70007FFF98998EFB47FD8BC2C7" author="Flint" box="[424,561,1230,1256]" firstAuthor="Flint" pageId="44" pageNumber="45" pagination="1 - 100" refId="ref41926" refString="Flint, O. S., Jr. (1983) Studies of Neotropical caddisflies, XXXIII: New species from austral South America (Trichoptera). Smithsonian Contributions to Zoology, 377, 1 - 100." type="journal article" year="1983">Flint, 1983</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30D650A007FFF9898B1FB78FF5CC326" box="[151,230,1265,1289]" pageId="44" pageNumber="45" type="description">
<paragraph id="8BA83681007FFF9898B1FB78FF5CC326" blockId="44.[151,568,1230,1289]" box="[151,230,1265,1289]" pageId="44" pageNumber="45">
<figureCitation id="132C2A04007FFF9898B1FB78FF5CC326" box="[151,230,1265,1289]" captionStart="FIGURE 22" captionStartId="45.[151,250,1979,2002]" captionTargetBox="[269,1312,183,1954]" captionTargetId="figure-43@45.[259,1330,183,1971]" captionTargetPageId="45" captionText="FIGURE 22. Mortoniella punensis (Flint), male genitalia: A—lateral; B—segment IX and tergum X, dorsal; C—phallic ensemble, ventral; D—dorsal phallic spine, dorsal; E—ventral process of segment VI, lateral." figureDoi="http://doi.org/10.5281/zenodo.5287021" httpUri="https://zenodo.org/record/5287021/files/figure.png" pageId="44" pageNumber="45">Fig. 22</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30D650A007FFF9898B1FAB0FC2CC049" pageId="44" pageNumber="45" type="reference_group">
<paragraph id="8BA83681007FFF9898B1FAB0FB9EC360" blockId="44.[151,1060,1337,1390]" box="[151,1060,1337,1359]" pageId="44" pageNumber="45">
<treatmentCitationGroup id="AB0711AF007FFF9898B1FAB0FB9EC360" box="[151,1060,1337,1359]" pageId="44" pageNumber="45">
<treatmentCitation id="0AB61090007FFF9898B1FAB0FE44C360" author="Flint, O. S., Jr." box="[151,510,1337,1359]" page="9" pageId="44" pageNumber="45" year="1983">
<taxonomicName id="4C174D02007FFF9898B1FAB0FE44C360" ID-CoL="42KXZ" authority="Flint, 1983: 9" authorityName="Flint" authorityPageNumber="9" authorityYear="1983" box="[151,510,1337,1359]" class="Insecta" family="Glossosomatidae" genus="Mexitrichia" kingdom="Animalia" order="Trichoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="punensis">
<emphasis id="B963EA93007FFF9898B1FAB0FED6C360" box="[151,364,1337,1359]" italics="true" pageId="44" pageNumber="45">Mexitrichia punensis</emphasis>
<bibRefCitation id="EF864B70007FFF989954FAB0FE44C360" author="Flint" box="[370,510,1337,1359]" firstAuthor="Flint" page="9" pageId="44" pageNumber="45" pagination="1 - 100" refId="ref41926" refString="Flint, O. S., Jr. (1983) Studies of Neotropical caddisflies, XXXIII: New species from austral South America (Trichoptera). Smithsonian Contributions to Zoology, 377, 1 - 100." type="journal article" year="1983">Flint, 1983: 9</bibRefCitation>
</taxonomicName>
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; Rueda &amp; Gibon 2008: 223 [illustration; distribution].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA83681007FFF9898B1FAD1FBAEC341" blockId="44.[151,1060,1337,1390]" box="[151,1044,1368,1390]" pageId="44" pageNumber="45">
<treatmentCitationGroup id="AB0711AF007FFF9898B1FAD1FBAEC341" box="[151,1044,1368,1390]" pageId="44" pageNumber="45">
<taxonomicName id="4C174D02007FFF9898B1FAD1FE03C341" authority="(Flint)" baseAuthorityName="Flint" baseAuthorityYear="1983" box="[151,441,1368,1390]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="44" pageNumber="45" phylum="Tracheophyta" rank="species" species="punensis">
<emphasis id="B963EA93007FFF9898B1FAD1FED5C341" box="[151,367,1368,1390]" italics="true" pageId="44" pageNumber="45">Mortoniella punensis</emphasis>
(Flint)
</taxonomicName>
; 
<treatmentCitation id="0AB61090007FFF9899E5FAD1FCB8C341" author="Blahnik, R. J. &amp; Holzenthal, R. W." box="[451,770,1368,1390]" page="69" pageId="44" pageNumber="45" year="2008">
<bibRefCitation id="EF864B70007FFF9899E5FAD1FCB8C341" author="Blahnik &amp; Holzenthal" box="[451,770,1368,1390]" firstAuthor="Blahnik" page="69" pageId="44" pageNumber="45" pagination="1 - 72" refId="ref41691" refString="Blahnik, R. J. &amp; Holzenthal, R. W. (2008) Revision of the Mexican and Central American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae). Zootaxa, 1711, 1 - 72." type="journal article" year="2008">Blahnik &amp; Holzenthal 2008: 69</bibRefCitation>
</treatmentCitation>
[
<taxonomicName id="4C174D02007FFF989B37FAD0FCCEC341" baseAuthorityName="ME Mosely" baseAuthorityYear="1937" box="[785,884,1368,1390]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="44" pageNumber="45" phylum="Tracheophyta" rank="species" species="leroda">
<emphasis id="B963EA93007FFF989B37FAD0FC9CC341" box="[785,806,1369,1390]" italics="true" pageId="44" pageNumber="45">M</emphasis>
. 
<emphasis id="B963EA93007FFF989B15FAD1FCCEC341" box="[819,884,1368,1390]" italics="true" pageId="44" pageNumber="45">leroda</emphasis>
</taxonomicName>
species group].
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</paragraph>
<paragraph id="8BA83681007FFF9898B1FA12FC2CC049" blockId="44.[151,1436,1434,2034]" pageId="44" pageNumber="45">
<taxonomicName id="4C174D02007FFF9898B1FA12FE3CC39D" baseAuthorityName="Flint" baseAuthorityYear="1983" box="[151,390,1435,1459]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="44" pageNumber="45" phylum="Tracheophyta" rank="species" species="punensis">
<emphasis id="B963EA93007FFF9898B1FA12FE3CC39D" box="[151,390,1435,1459]" italics="true" pageId="44" pageNumber="45">Mortoniella punensis</emphasis>
</taxonomicName>
is most closely related to 
<taxonomicName id="4C174D02007FFF989A8BFA12FCA4C39D" baseAuthorityName="Rueda &amp; Gibon" baseAuthorityYear="2008" box="[685,798,1435,1458]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="44" pageNumber="45" phylum="Tracheophyta" rank="species" species="marini">
<emphasis id="B963EA93007FFF989A8BFA12FCA4C39D" box="[685,798,1435,1458]" italics="true" pageId="44" pageNumber="45">M. marini</emphasis>
</taxonomicName>
. As mentioned in the discussion of the 
<taxonomicName id="4C174D02007FFF989CFAFA12FADEC39D" baseAuthorityName="Flint" baseAuthorityYear="1983" box="[1244,1380,1435,1458]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="44" pageNumber="45" phylum="Tracheophyta" rank="species" species="punensis">
<emphasis id="B963EA93007FFF989CFAFA12FB49C39D" box="[1244,1267,1435,1458]" italics="true" pageId="44" pageNumber="45">M</emphasis>
. 
<emphasis id="B963EA93007FFF989D24FA12FADEC39D" box="[1282,1380,1435,1458]" italics="true" pageId="44" pageNumber="45">punensis</emphasis>
</taxonomicName>
subgroup, both species have curved, spine-like dorsolateral processes on the phallicata that hook over the paramere appendages, causing them to cross mesally, and also have upturned, flattened processes from the ventral margin of the phallicata. Both species also have a second apicolateral projection on each side of tergum X, in addition to the usual acute apicolateral projection. 
<taxonomicName id="4C174D02007FFF989A0DF9A2FD0EC06D" baseAuthorityName="Flint" baseAuthorityYear="1983" box="[555,692,1579,1602]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="44" pageNumber="45" phylum="Tracheophyta" rank="species" species="punensis">
<emphasis id="B963EA93007FFF989A0DF9A2FD0EC06D" box="[555,692,1579,1602]" italics="true" pageId="44" pageNumber="45">M. punensis</emphasis>
</taxonomicName>
is most readily diagnosed from 
<taxonomicName id="4C174D02007FFF989C01F9A2FB23C06D" baseAuthorityName="Rueda &amp; Gibon" baseAuthorityYear="2008" box="[1063,1177,1579,1602]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="44" pageNumber="45" phylum="Tracheophyta" rank="species" species="marini">
<emphasis id="B963EA93007FFF989C01F9A2FB23C06D" box="[1063,1177,1579,1602]" italics="true" pageId="44" pageNumber="45">M. marini</emphasis>
</taxonomicName>
by having a tergum X that is rounded and projecting apicomesally, rather than invaginated.
</paragraph>
</subSubSection>
<subSubSection id="C30D650A007FFF9A98E1F9FBFD8EC438" lastPageId="46" lastPageNumber="47" pageId="44" pageNumber="45" type="description">
<paragraph id="8BA83681007FFF9898E1F9FBFB15C0FD" blockId="44.[151,1436,1434,2034]" pageId="44" pageNumber="45">
Adult. Length of forewing: male 
<quantity id="4CEF9B64007FFF989A11F9FBFD35C0A5" box="[567,655,1650,1674]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="44" pageNumber="45" unit="mm" value="4.2">4.2 mm</quantity>
. Forewing with forks I, II, and III present, hind wing with fork II only. Spur formula 0:4:4. Overall color medium brown. Tibial spurs somewhat darker than legs, contrasting in color. Wing bar at anastamosis more or less distinctly marked with whitish setae, bar discontinuous.
</paragraph>
<paragraph id="8BA83681007FFF9A98E1F957FAF7C783" blockId="44.[151,1436,1434,2034]" lastBlockId="46.[151,1437,151,535]" lastPageId="46" lastPageNumber="47" pageId="44" pageNumber="45">
Male genitalia. Ventral process of segment VI laterally compressed, somewhat posteriorly directed, relatively wide basally, narrowed and subacute apically, length about 2 times width at base (
<figureCitation id="132C2A04007FFF989C61F88BFB15C135" box="[1095,1199,1794,1818]" captionStart="FIGURE 22" captionStartId="45.[151,250,1979,2002]" captionTargetBox="[269,1312,183,1954]" captionTargetId="figure-43@45.[259,1330,183,1971]" captionTargetPageId="45" captionText="FIGURE 22. Mortoniella punensis (Flint), male genitalia: A—lateral; B—segment IX and tergum X, dorsal; C—phallic ensemble, ventral; D—dorsal phallic spine, dorsal; E—ventral process of segment VI, lateral." figureDoi="http://doi.org/10.5281/zenodo.5287021" httpUri="https://zenodo.org/record/5287021/files/figure.png" pageId="44" pageNumber="45">Fig. 22E</figureCitation>
). Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin convexly rounded in dorsal 1/2, narrowing ventrally; segment deeply excised dorsomesally and ventromesally, forming lateral lobes, lobes separated dorsomesally by much less than 1/2 width of segment. Tergum X convexly rounded mesally, with short, projecting dorsolateral lobes and rounded ventrolateral lobes; dorsolateral lobes, as viewed dorsally, somewhat mesally curved, narrow basally, subacute apically. Inferior appendages short, without distinct ventromesal projection, lateral lobes rounded dorsally, not forming tapering dorsal projections. Mesal pockets of fused inferior appendages with apical processes relatively short, posteriorly curved. Paramere appendages short, narrow, uniform in width, acute apically, each mesally turned so that appendages from opposite sides cross over one another mesally. Dorsal phallic spine, as viewed laterally, somewhat widened in middle, rather gradually upturned from base, more strongly in apical 1/4th, tapering apically; in dorsal view, uniform in width in basal part, somewhat inflated before apical inflection, apex weakly bifid, apices acute. Phallicata with elongate narrow, sinuous dorsolateral processes, each curved over corresponding paramere appendage; laterally with broad, subtruncate, dorsallydirected lobes, each paralleling assemblage formed by paramere appendage and dorsolateral lobe of phallicata. Endophallic membrane apparently short and simple, without ventromesal spine; phallotremal spines absent.
</paragraph>
<caption id="DF686609007EFF9998B1F832FBF1C1DB" ID-DOI="http://doi.org/10.5281/zenodo.5287021" ID-Zenodo-Dep="5287021" httpUri="https://zenodo.org/record/5287021/files/figure.png" pageId="45" pageNumber="46" startId="45.[151,250,1979,2002]" targetBox="[269,1312,183,1954]" targetPageId="45">
<paragraph id="8BA83681007EFF9998B1F832FBF1C1DB" blockId="45.[151,1436,1979,2036]" pageId="45" pageNumber="46">
<emphasis id="B963EA93007EFF9998B1F832FE9EC1FD" bold="true" box="[151,292,1979,2002]" pageId="45" pageNumber="46">FIGURE 22.</emphasis>
<taxonomicName id="4C174D02007EFF999908F832FDE3C1FE" authority="(Flint)" baseAuthorityName="Flint" baseAuthorityYear="1983" box="[302,601,1979,2001]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="45" pageNumber="46" phylum="Tracheophyta" rank="species" species="punensis">
<emphasis id="B963EA93007EFF999908F832FDB1C1FE" box="[302,523,1979,2001]" italics="true" pageId="45" pageNumber="46">Mortoniella punensis</emphasis>
(Flint)
</taxonomicName>
, male genitalia: A—lateral; B—segment IX and tergum X, dorsal; C—phallic ensemble, ventral; D—dorsal phallic spine, dorsal; E—ventral process of segment VI, lateral.
</paragraph>
</caption>
<paragraph id="8BA83681007DFF9A98E1FE3EFE1BC7DB" blockId="46.[151,1437,151,535]" pageId="46" pageNumber="47">
<materialsCitation id="3B7F3CDC007DFF9A98E1FE3EFB23C7E0" box="[199,1177,439,464]" collectingDate="1973-10-11" collectionCode="OS" collectorName="La Angostura" country="Argentina" location="Tucuman" pageId="46" pageNumber="47" specimenCount="1" stateProvince="Tucuman">
Material examined. 
<emphasis id="B963EA93007DFF9A998CFE3EFD5BC7FF" bold="true" box="[426,737,439,464]" pageId="46" pageNumber="47">
<collectingCountry id="F3007611007DFF9A998CFE3EFDE3C7FF" box="[426,601,439,464]" name="Argentina" pageId="46" pageNumber="47">ARGENTINA</collectingCountry>
: 
<collectingRegion id="49D3F863007DFF9A9A41FE31FD66C7FF" box="[615,732,440,464]" country="Argentina" name="Tucuman" pageId="46" pageNumber="47">Tucuman</collectingRegion>
:
</emphasis>
Rt. 307, 
<collectorName id="26E25357007DFF9A9B6BFE31FC50C7E0" box="[845,1002,439,463]" pageId="46" pageNumber="47">La Angostura</collectorName>
, 
<date id="FFA91041007DFF9A9BD1FE3EFBD3C7E0" box="[1015,1129,439,463]" pageId="46" pageNumber="47" value="1973-10-11">
<collectingDate id="EFEDE9A9007DFF9A9BD1FE3EFBD3C7E0" box="[1015,1129,439,463]" pageId="46" pageNumber="47" value="1973-10-11">11.x.1973</collectingDate>
</date>
, 
<collectionCode id="ED06AE44007DFF9A9C53FE3EFB23C7E0" box="[1141,1177,439,463]" country="USA" name="Oregon State University" pageId="46" pageNumber="47">OS</collectionCode>
</materialsCitation>
<materialsCitation id="3B7F3CDC007DFF9A9C84FE3EFE27C7DB" collectionCode="NMNH" country="Argentina" pageId="46" pageNumber="47" specimenCount="1" specimenCount-male="1" stateProvince="Tucuman" typeStatus="paratype">
Flint — 
<specimenCount id="9D11FD08007DFF9A9D22FE3EFAEAC7E0" box="[1284,1360,439,463]" pageId="46" pageNumber="47" type="male">1 male</specimenCount>
(
<typeStatus id="54AC8823007DFF9A9D47FE3EFF70C7DB" pageId="46" pageNumber="47" type="paratype">paratype</typeStatus>
, pinned) (
<collectionCode id="ED06AE44007DFF9A991FFE55FE2FC7DB" box="[313,405,476,500]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="46" pageNumber="47" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA83681007DFF9A98E1FE76FD8EC438" blockId="46.[151,1437,151,535]" box="[199,564,511,535]" pageId="46" pageNumber="47">
Distribution. 
<collectingCountry id="F3007611007DFF9A997AFE76FE75C438" box="[348,463,511,535]" name="Argentina" pageId="46" pageNumber="47">Argentina</collectingCountry>
, 
<collectingCountry id="F3007611007DFF9A99FCFE76FD8AC438" box="[474,560,511,535]" name="Bolivia" pageId="46" pageNumber="47">Bolivia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>