Revision of the Nearctic genus Tritoxa Loew (Diptera: Ulidiidae) Author Sinclair, Bradley J. Canadian National Collection of Insects & Canadian Food Inspection Agency, OPL-Entomology, K. W. Neatby Bldg., C. E. F., 960 Carling Ave., Ottawa, ON, K 1 A 0 C 6, Canada Author Macleod, Alyssa M. 0000-0002-7048-8481 Department of Natural Resource Sciences, McGill University, Macdonald Campus, Ste-Anne-de-Bellevue, QC, H 9 X 3 V 9, Canada alyssa. m. macleod @ gmail. com; https: // orcid. org / 0000 - 0002 - 7048 - 8481 alyssa.m.macleod@gmail.com Author Wheeler, Terry A. text Zootaxa 2021 2021-01-29 4920 3 359 379 journal article 8337 10.11646/zootaxa.4920.3.3 3f20c135-c448-4c8c-a716-993e4f6d50d0 1175-5326 4478080 20794FBC-E83E-419F-945D-4F94C6F9A47C Tritoxa californica sp. nov. ( Figs 6 , 14, 15 , 24 ) urn:lsid:zoobank.org:act: 0350D4DB-7407-4C19-941A-31B9951299EB Type material. HOLOTYPE ♂, labelled: “ CALIF : Plumas Co. / Little Long Valley / Creek [ 39°52′06″N 120°42′08″W ], 6000′ 6 mi. E./ Spring Garden/ VIII-4-10-1977 ”; “ Malaise Trap / 8A-6P/ M. Wasbauer / Collector”; “ HOLOTYPE / Tritoxa / californica Sinclair ,/ MacLeod & Wheeler” ( CSCA ) . PARATYPES : USA . California : El Dorado Co. , Pollack Pines [ 38°45′41″N 120°35′12″W ], 2.ix.1986 , Jackson trap , peach, D. Bolster (1♂, CSCA ) ; Nipa Co. , Angwin , 38.575°N 122.453°W , 17.ix.2008 , McPhail trap , T . Samansky (1♂, 1♀ , CSCA ) ; Same data as holotype (1♂, 2♀ , CSCA ) ; Santa Cruz Co. , Big Basin SP [ 37°10′21″N 122°13′21″W ], 13.ix.1966 , E.L. Sleeper (1♂, CAS ) ; Trinity Co. [ 40.66°N 123.12°W ], 13.vi.1934 (1♂, CAS ) ; Tuolumne Co. , 2.7 mi E Smoky Jack [ 37°49′3.72″N 119°42′45.61″W ], Summer 1962, Frick trap on Prunus emarginata , D.P. Allen (1♂, CSCA ) . Possible additional material. USA . California : El Dorado Co. , Placerville [ 38°43′47″N 120°47′55″W ] , 26.vi.1953 , P.H. Arnaud (1♂, USNM ) [genitalia lost]; Sagehen Ck Field Stn., 15 km N Truckee , 39°25.89′N 120°14.61′W , 16.viii.1999 , J. Savage ( 1♀ , LEM ); Nevada Co., Sagehen Ck [ 39°25′57″N 120°14′13″W ] , 2.vii.1968 , R . W. Pinger ( 1♀ , USNM ) . Diagnosis. This species has the general appearance of Tritoxa cuneata , but is distinguished by the short, oblique discal hyaline crossband not extending beyond level of crossvein r-m in cell dm, inner surstylus with tight cluster of 4–5 thick prensisetae and phallus is shorter, with two loops. Description. Entirely yellowish brown to brown, abdomen darker brown bands. Head: ocellar triangle dark brown to black, microtrichose; frons yellowish brown to brown; parafrontal silvery microtrichia stripe very narrow, limited to extreme margin of eye, continuous with very narrow microtrichose parafacial; gena slightly darker brown below narrowest part of lower eye margin; postocular microtrichia slightly wider than parafrontal microtrichose stripe in lateral view, extending from base to middle of eye; face concolourous with face laterally, slightly paler medially; supracervical setulae very pale. Antenna with postpedicel greyish brown, yellowish brown on inner and medially surfaces. Proboscis with yellowish brown palpus. Thorax: yellowish brown; scutum mostly thinly microtrichose without pair of vittae; lateral margin of scutum and postpronotal lobe broadly shiny; scutellum concolourous with scutum; mediotergite without distinct stripe; pleura shiny, anepisternum and katepisternum lightly clothed in whitish microtrichia when viewed obliquely; whitish microtrichia above fore coxa; 2 anepisternal setae. Wing ( Fig. 6 ) entirely brown, slightly tapered apically, with 3 hyaline crossbands; some faded or paler regions between bands; costal cell without hyaline band at humeral break, with narrow hyaline band at extreme apex of cell in line with discal hyaline crossband; oblique subbasal hyaline crossband very short, extending to basal fifth of cell dm; discal hyaline crossband short, continuous from R 1 , reaching posterior margin of cell dm, not extending beyond level of crossvein r-m in cell dm; subapical hyaline crossband nearly straight, anterior end not extending proximal to crossvein r-m; crossvein dm-m straight to slightly arched; anal lobe developed; calypter with white margin. Abdomen: yellowish brown. Male terminalia ( Figs 14, 15 ): inner surstylus with tight cluster of 4–5 thick prensisetae, inner margin concave; outer surstylus slender, slightly tapered and arched, slightly longer than length of inner surstylus, with narrow apex ( Fig. 10 ); subepandrial sclerite with strong dark setae. Cercus with short setae; apical margin slightly pointed. Phallus short, with 2 loops. Distribution. This species is known exclusively from northern California ( Fig. 24 ). Remarks. See comments under T. cuneata .