Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species Author Motamedinia, Behnam Author Skevington, Jeffrey H. 0000-0002-1445-9870 jhskevington@gmail.com Author Kelso, Scott 0000-0001-5559-9477 scott.kelso@canada.ca text Zootaxa 2021 2021-07-16 5002 1 1 103 journal article 10.11646/zootaxa.5002.1.1 1175-5326 5122655 5DC2A66A-3F04-42D0-8A8C-F0686054E556 Tomosvaryella subvirescens ( Loew, 1872 ) Figs 39A–E , 46D , 64 Pipunculus subvirescens Loew, 1872 .—syn. De Meyer (1996) : p. 95 . Diagnosis: This species can be recognized by the shape of surstyli in dorsal view, right surstylus quadratic-shaped with small inner projection, left surstylus elongated, rather rectangular-shaped, longer than right one ( Fig. 39A ); epandrium wider than long, slightly ovate-shaped ( Fig. 39A ); phallus trifid with wavy ejaculatory ducts ( Fig. 39D–E ); asymmetrical gonopods in ventral view, right higher than left one ( Fig. 39B ); hypandrium ovate-shaped with distinct hypandrial apodeme in ventral view ( Fig. 39B ). FIGURE 38. Male genitalia of Tomosvaryella subtransvaalensis sp. nov. , holotype (JSS51765, TAU) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view. FIGURE 39. Male genitalia of Tomosvaryella subvirescens (CNCD174944, CNC) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view. Specimens examined: IRAN : Sistan & Baluchestan : Saravan , 27°25’N , 62°17’E , 12.v.2013 , F. Hamzavi , pan trap , JSS51916 ( 1♂ , CNC ) ; ISRAEL : Ne’ot haKikkar, 30°56’N , 35°22’E , 1.v.1997 , A. Freidberg , JSS51694 ( 1♂ , TAU ) ; QATAR : Al Shamal distnict, Al Zubarah exclusion zone, Grassy meadow, 25°58’N , 51°02’E , 20.iii.2012 , K. P. Puliafico & A. M. Jensen , yellow & white pan traps , JSS52318–9 ( 2♂ , CNC ) ; UNITED ARAB EMIRATES : Bithnah , 25°06’N , 56°08’E , 4–26.iii.2006 , A. van Harten , light trap , JSS52240 ( 1♂ , CNC ); NARC , near Sweihan , 24°14’N , 55°16’E , 9–20.ix.2005 , A. van Harten , light trap , CNCD174944–7 ( 4♂ , CNC ); Wadi Maidaq , 25°11’N , 56°04’E , 7–14.iii.2006 , A. van Harten , pan trap , CNCD8951 ( 1♂ , USNM ); Wadi Wurayah farm, 25°14’N , 56°11’E , 17–24.iii.2009 , A. van Harten , Malaise trap ; CNCD137124–5 , CNCD137128 ( 3♂ , CNC ); 19.iv–19.v.2009 , A. van Harten , Malaise trap , CNCD137532 , CNCD137534 , CNCD137536 , CNCD137538 , CNCD137540 , CNCD137545 , CNCD137552 ( 7♂ , CNC ); 24–30.iii.2009 , A. van Harten , Malaise trap , CNCD136930 , CNCD136932 ( 2♂ , CNC ); 31.v–14.vi.2009 , A. van Harten , Malaise trap , CNCD137578 ( 1♂ , EAD ); Wadi Wurayah , 25°14’N , 56°11’E , 19– 26.iv.2009 , A. van Harten , Malaise trap , CNCD160583 ( 1♂ , CNC ); al-Rafa, 25°26’N , 55°31’E , 20–23.iv.2009 , A. van Harten , pan trap , CNCD189979 ( 1♂ , CNC ); near ad-Dhaid, 25°13’N , 55°35’E , 16–19.iii.2007 , J. Btelka , pan trap , CNCD159476 , CNCD159478–9 , CNCD159483–4 , CNCD159486 , CNCD159488 ( 7♂ , CNC ) . Distribution: Bahamas , Belize , Bermuda , Brazil , Canada , Chile , Colombia , Costa Rica , Cuba , Dominica , Egypt , Gabon , Iran , Israel , Jamaica , Mexico , Mozambique , Namibia , Nicaragua , Panama , Peru , Puerto Rico , Qatar , Thailand , the United States , United Arab Emirates ( Skevington 2020 ) ( Fig. 64 ). Note: DNA barcodes of T. subvirescens overlap with T. pilosiventris . See the notes under T. pilosiventris .