<document id="BCBB4B0D482F9BF831567744CC319AB2" ID-DOI="10.11646/zootaxa.5655.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="15822077" ID-ZooBank="4411E6C2-D4C5-4340-AF6B-FDDBB0F7E1A1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1751801736697" checkinUser="plazi" docAuthor="Perina, Giulia, Camacho, Ana I., Morgan, Liesel, Lawrie, Angus, Floeckner, Stephanie &amp; Guzik, Michelle T." docDate="2025" docId="0399C326FFADFFC453BB04CFFBFBFB2F" docLanguage="en" docName="zootaxa.5655.1.1.pdf" docOrigin="Zootaxa 5655 (1)" docSource="https://doi.org/10.11646/zootaxa.5655.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Atopobathynella Schminke 1973" docType="treatment" docVersion="2" lastPageNumber="23" masterDocId="FFA0BB5EFFBBFFD2532C055CFFA9FFDC" masterDocTitle="New species of Atopobathynella, Kimberleybathynella and Hexabathynella (Parabathynellidae, Bathynellacea) from the arid zones of Western Australia" masterLastPageNumber="103" masterPageNumber="1" pageNumber="23" updateTime="1751803507751" updateUser="ExternalLinkService">
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<mods:title id="A2554180FF4AD48D92C95280870BE710">New species of Atopobathynella, Kimberleybathynella and Hexabathynella (Parabathynellidae, Bathynellacea) from the arid zones of Western Australia</mods:title>
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<mods:namePart id="90DF6BCC28B1C8D20C82B0FE9CC3384A">Perina, Giulia</mods:namePart>
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<mods:affiliation id="FEA10FB046834EC7AC03208BD5988B35">Collections &amp; Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia &amp; Biologic Environmental Survey, East Perth, Western Australia 6004, Australia liesel @ biologicenv. com. au; https: // orcid. org / 0009 - 0001 - 1458 - 6490 steph @ biologicenv. com. au; https: // orcid. org / 0009 - 0000 - 5089 - 7096 &amp; Subterranean Research and Groundwater Ecology (SuRGE) Group, Trace and Environmental DNA (TrEnD) Laboratory, School of Molecular and Life Sciences, Curtin University, Perth 6102, WA, Australia angus. lawrie @ curtin. edu. au; https: // orcid. org / 0000 - 0002 - 6766 - 1191</mods:affiliation>
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<mods:namePart id="208DA21FFC6D74F4329D0A9965B807A7">Camacho, Ana I.</mods:namePart>
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<mods:affiliation id="DE7778DB15912218012460DFC9FAE01C">Museo Nacional de Ciencias Naturales (CSIC). Dpto. Biodiversidad y Biología Evolutiva. C / José Gutiérrez Abascal 2. 28006 - Madrid, Spain mcnac 22 @ mncn. csic. es; https: // orcid. org / 0000 - 0003 - 0596 - 7678</mods:affiliation>
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<mods:namePart id="AD63CCD9E1D1DBE229E0A3A2BFF1CB8E">Morgan, Liesel</mods:namePart>
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<mods:affiliation id="5EA59075169E66C849099A7C5B32F42E">Biologic Environmental Survey, East Perth, Western Australia 6004, Australia liesel @ biologicenv. com. au; https: // orcid. org / 0009 - 0001 - 1458 - 6490 steph @ biologicenv. com. au; https: // orcid. org / 0009 - 0000 - 5089 - 7096</mods:affiliation>
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<mods:namePart id="A3134B231996ECEEAA3431D8E5D68362">Lawrie, Angus</mods:namePart>
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<mods:affiliation id="E8EC45B04B8F050A79E860BA3A53264F">Subterranean Research and Groundwater Ecology (SuRGE) Group, Trace and Environmental DNA (TrEnD) Laboratory, School of Molecular and Life Sciences, Curtin University, Perth 6102, WA, Australia angus. lawrie @ curtin. edu. au; https: // orcid. org / 0000 - 0002 - 6766 - 1191</mods:affiliation>
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<mods:namePart id="5590A1B11B93ECEB7281ABCFD2573C06">Floeckner, Stephanie</mods:namePart>
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<mods:affiliation id="B1AFCA341AEA25A4CABA2E4DF70DBFB8">Biologic Environmental Survey, East Perth, Western Australia 6004, Australia liesel @ biologicenv. com. au; https: // orcid. org / 0009 - 0001 - 1458 - 6490 steph @ biologicenv. com. au; https: // orcid. org / 0009 - 0000 - 5089 - 7096</mods:affiliation>
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<mods:namePart id="FBF146159AA0F8ACD3D5DB31511DE7E8">Guzik, Michelle T.</mods:namePart>
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<mods:affiliation id="BCAB22839A33D6B22C87594DF7369655">Collections &amp; Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia &amp; School of Biological Sciences, The University of Adelaide, SA 5005, Australia michelle. guzik @ adelaide. edu. au; https: // orcid. org / 0000 - 0002 - 4947 - 9353 * Corresponding author: giulialittlepear @ gmail. com; https: // orcid. org / 0000 - 0002 - 0349 - 3803 &amp; Collections &amp; Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia</mods:affiliation>
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<taxonomicName id="4C3009B3FFADFFC453BB04CFFD82FE71" ID-CoL="3574" ID-ENA="489763" authority="Schminke, 1973" authorityName="Schminke" authorityYear="1973" box="[151,555,403,430]" class="Malacostraca" family="Parabathynellidae" genus="Atopobathynella" kingdom="Animalia" order="Bathynellacea" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B944AE22FFADFFC453BB04CFFD82FE71" bold="true" box="[151,555,403,430]" pageId="22" pageNumber="23">
<emphasis id="B944AE22FFADFFC453BB04CFFEF7FE72" bold="true" box="[151,350,403,430]" italics="true" pageId="22" pageNumber="23">Atopobathynella</emphasis>
<bibRefCitation id="EFA10FC1FFADFFC4524904CFFD82FE71" author="Schminke, H. K." box="[357,555,403,429]" pageId="22" pageNumber="23" pagination="1 - 192" refId="ref42972" refString="Schminke, H. K. (1973 a) Evolution, System und Verbreitungs geschichte der Familie Parabathynellidae (Bathynellacea, Malacostraca). Mikrofauna des Meeresbodens, 24 (87), 1-192." type="journal article" year="1973">Schminke, 1973</bibRefCitation>
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<taxonomicName id="4C3009B3FFADFFC453BB048EFE13FE36" authority="Noodt, 1965" authorityName="Noodt" authorityYear="1965" box="[151,442,466,490]" class="Malacostraca" family="Parabathynellidae" genus="Parabathynella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Bathynellacea" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B944AE22FFADFFC453BB048EFE9CFE36" box="[151,309,466,490]" italics="true" pageId="22" pageNumber="23">Parabathynella</emphasis>
<bibRefCitation id="EFA10FC1FFADFFC45217048EFE13FE36" author="Noodt, W." box="[315,442,466,490]" pageId="22" pageNumber="23" pagination="77 - 186" refId="ref42047" refString="Noodt, W. (1965) Naturliches System und Biogeographie der Syncarida. Gewasser und Abwasser, 37 / 38, 77-186." type="journal article" year="1965">Noodt, 1965</bibRefCitation>
</taxonomicName>
(synonymised by 
<bibRefCitation id="EFA10FC1FFADFFC45158048EFCB3FE36" author="Schminke, H. K." box="[628,794,466,490]" pageId="22" pageNumber="23" pagination="1 - 192" refId="ref42972" refString="Schminke, H. K. (1973 a) Evolution, System und Verbreitungs geschichte der Familie Parabathynellidae (Bathynellacea, Malacostraca). Mikrofauna des Meeresbodens, 24 (87), 1-192." type="journal article" year="1973">Schminke, 1973</bibRefCitation>
)
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<emphasis id="B944AE22FFADFFC453BB04ADFE8AFDD5" bold="true" box="[151,291,497,521]" pageId="22" pageNumber="23">
<typeStatus id="548BCC92FFADFFC453BB04ADFF65FDD5" box="[151,204,497,521]" pageId="22" pageNumber="23">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C3009B3FFADFFC4520604ADFD71FDD5" authority="(Noodt, 1965)" baseAuthorityName="Noodt" baseAuthorityYear="1965" box="[298,728,497,521]" class="Malacostraca" family="Parabathynellidae" genus="Atopobathynella" kingdom="Animalia" order="Bathynellacea" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="valdiviana">
<emphasis id="B944AE22FFADFFC4520604ADFDEAFDD5" box="[298,579,497,521]" italics="true" pageId="22" pageNumber="23">Atopobathynella valdiviana</emphasis>
(
<bibRefCitation id="EFA10FC1FFADFFC4517C04ADFD78FDD5" author="Noodt, W." box="[592,721,497,521]" pageId="22" pageNumber="23" pagination="77 - 186" refId="ref42047" refString="Noodt, W. (1965) Naturliches System und Biogeographie der Syncarida. Gewasser und Abwasser, 37 / 38, 77-186." type="journal article" year="1965">Noodt, 1965</bibRefCitation>
)
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</paragraph>
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<taxonomicName id="4C3009B3FFADFFC453EB074CFDF2FDFB" authority="Noodt, 1965" authorityName="Noodt" authorityYear="1965" box="[199,603,527,552]" class="Malacostraca" family="Parabathynellidae" genus="Parabathynella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Bathynellacea" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="valdiviana">
<emphasis id="B944AE22FFADFFC453EB074CFE7FFDF4" box="[199,470,528,552]" italics="true" pageId="22" pageNumber="23">Parabathynella valdiviana</emphasis>
<bibRefCitation id="EFA10FC1FFADFFC452F0074CFDF2FDFB" author="Noodt, W." box="[476,603,527,552]" pageId="22" pageNumber="23" pagination="77 - 186" refId="ref42047" refString="Noodt, W. (1965) Naturliches System und Biogeographie der Syncarida. Gewasser und Abwasser, 37 / 38, 77-186." type="journal article" year="1965">Noodt, 1965</bibRefCitation>
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</paragraph>
<paragraph id="8B8F7230FFADFFC453BB0772FE0EFD9A" blockId="22.[151,801,466,582]" box="[151,423,558,582]" pageId="22" pageNumber="23">
<emphasis id="B944AE22FFADFFC453BB0772FE80FD9A" bold="true" box="[151,297,558,582]" pageId="22" pageNumber="23">Other species</emphasis>
: see 
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<subSubSection id="C32A21BBFFADFFC453BB0728FBFBFB2F" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="8B8F7230FFADFFC453BB0728FD06FD52" blockId="22.[151,1437,628,1267]" box="[151,687,628,654]" pageId="22" pageNumber="23">
<emphasis id="B944AE22FFADFFC453BB0728FED6FD52" bold="true" box="[151,383,628,654]" pageId="22" pageNumber="23">Amended diagnosis</emphasis>
(after 
<bibRefCitation id="EFA10FC1FFADFFC452E60728FD0AFD52" author="Perina, G. &amp; Camacho, A. I. &amp; Danks, M. &amp; White, N. &amp; Guzik, M. T." box="[458,675,628,654]" pageId="22" pageNumber="23" pagination="2228326" refId="ref42229" refString="Perina, G., Camacho, A. I., Danks, M., White, N. &amp; Guzik, M. T. (2023 b) Two new species of Atopobathynella (Parabathynellidae, Bathynellacea) from the Pilbara region, Australia. Systematics and Biodiversity, 21 (1), 2228326. https://doi.org/10.1080/14772000.2023.2228326" type="journal article" year="2023">
Perina 
<emphasis id="B944AE22FFADFFC451350729FDF8FD52" box="[537,593,628,654]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
2023b
</bibRefCitation>
).
</paragraph>
<paragraph id="8B8F7230FFADFFC453EB07C4FBFBFB2F" blockId="22.[151,1437,628,1267]" pageId="22" pageNumber="23">
Antennula 6-segmented often showing sexual dimorphism on the inner margin of the second article, with female bearing one simple seta, and male an antennal organ with one or two setae that can be modified. Antenna one-segmented, with three smooth setae (one inner proximal and two apical) and an apical seta. Labrum heterodont with numerous teeth. Mandible with 
<emphasis id="B944AE22FFADFFC451160659FD62FCC2" box="[570,715,773,798]" italics="true" pageId="22" pageNumber="23">pars incisiva</emphasis>
with three teeth (in one case four); lobe with row of five claws (
<emphasis id="B944AE22FFADFFC4538D0674FE9CFC9E" box="[161,309,808,834]" italics="true" pageId="22" pageNumber="23">Borstenlobus</emphasis>
). Distal endite of maxillula with five or six claws. Maxilla four-segmented (in two cases three-segmented) with setal formula: 2-2/4-n-1/7 (the third article can have a variable number “n” of setae, while the fourth article can have up to 7 setae). Six or seven pairs of thoracopods. Epipod present or absent in thoracopod I, always present in thoracopods II to VII. Exopod of thoracopods I to VII one-segmented. Exopod of thoracopods II to VII usually with two terminal setae and one ventral subterminal one (one species has two ventral setae, one species has none); endopod of thoracopods I to VII four-segmented, fourth article tiny, thoracopods II to VII with one or two very long claws on the fourth article; setal formula of endopod II to VII articles 1 to 3: 0+0/0+1/0+1. Male thoracopod VIII small, semicircular in lateral view; protopod massive, anteriorly with dentate lobe, and often with a small protrusion; outer lobe drawn out into a conical projection directed anteriorly; basipod triangular or round/ paddle-like, as large as outer lobe, with one or two setae of uncertain interpretation that could represent endopod and/or exopod. Pleopod I consisting of one seta. Sympod of the uropod with homonomous or inhomonomous row of several spines; endopod with dagger-like extension and two or three setae at its base (only one seta in one species); exopod of uropod often with ventromedial seta and 2 to 4 terminal/subterminal setae.
</paragraph>
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