Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae) Author Ballantyne, Lesley A. Author Lambkin, Christine text Zootaxa 2009 2009-02-04 1997 1 188 journal article 1175­5334 Pygatyphella marginata ( Ballantyne, 1968 ) Luciola ( Pygatyphella ) marginata Ballantyne, 1968:116 . Holotype . Male. PAPUA NEW GUINEA : Northern Pr. , 8.77S , 148.24E , Popondetta , 28.ii.1966 ( QM ). Diagnosis. A moderate sized ( 8.5–10 mm long) species with pale brown, pale margined elytra; distinguished from most pale coloured Pygatyphella by the absence of distinct elytral markings at base and apex. Remarks. Ballantyne (1968) described this species from 3 specimens .