Ultrastructure of attachment specializations of hexapods (Arthropoda): evolutionary patterns inferred from a revised ordinal phylogeny Author R. G. Beutel Author S. N. Gorb text J. Zool. Syst. Evol. Research 2001 39 177 207 journal article 28754 10.1046/j.1439-0469.2001.00155.x 95b3d62f-8fcc-4088-873f-b33f4a63d3fc 2360323 7 Odonata The tarsus is three­segmented and the distal segment is longer than the others. The paired claws are usually armed with a hook ( Watson and O’Farell 1991 ). Specific adhesive structures are absent from dragonflies and damselflies.