<document id="CE5C9F346F474B925720572D018B3E63" ID-DOI="10.5733/afin.049.0210" ID-ISSN="2305-2562" ID-Zenodo-Dep="7917381" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683578209636" checkinUser="felipe" docAuthor="Londt, Jason G. H." docDate="2008" docId="039087DEFFB76D4DBFB29C8367D0AF7D" docLanguage="en" docName="AfrInvertebr.49.2.171-226.pdf" docOrigin="African Invertebrates 49 (2)" docSource="http://www.bioone.org/doi/abs/10.5733/afin.049.0210" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.4:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="4" docTitle="Trichardis rueppelii" docType="treatment" docVersion="5" lastPageNumber="199" masterDocId="FFA9FFA6FFAC6D51BE01987D6467AB6A" masterDocTitle="A review of Afrotropical Trichardis Hermann, 1906, and the description of the first Oriental representative of the genus (Diptera: Asilidae: Laphriinae)" masterLastPageNumber="171" masterPageNumber="171" pageNumber="198" updateTime="1699499062618" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="6942D0E4E1FEB8AE5F815E547D3BE7D0">A review of Afrotropical Trichardis Hermann, 1906, and the description of the first Oriental representative of the genus (Diptera: Asilidae: Laphriinae)</mods:title>
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<mods:namePart id="E5EE0E3940687A9BCBDDBB7FAD2E5093">Londt, Jason G. H.</mods:namePart>
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<mods:number id="D53A94982ACE06A211D62234074A402D">49</mods:number>
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<paragraph id="8B8636C8FFB76D4ABFB29C83671FAE7D" blockId="27.[435,888,1278,1303]" box="[435,888,1278,1303]" pageId="27" pageNumber="198">
<heading id="D0CE81A4FFB76D4ABFB29C83671FAE7D" box="[435,888,1278,1303]" centered="true" fontSize="11" level="2" pageId="27" pageNumber="198" reason="3">
<taxonomicName id="4C394D4BFFB76D4ABFB29C83671FAE7D" authority="(Wiedemann, 1828)" baseAuthorityName="Wiedemann" baseAuthorityYear="1828" box="[435,888,1278,1303]" class="Insecta" family="Asilidae" genus="Trichardis" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="198" phylum="Arthropoda" rank="species" species="rueppelii">
<emphasis id="B94DEADAFFB76D4ABFB29C8366F5AE7C" box="[435,658,1278,1302]" italics="true" pageId="27" pageNumber="198">Trichardis rueppelii</emphasis>
(
<bibRefCitation id="EFA84B39FFB76D4ABCA09C836717AE7D" author="WIEDEMANN, C. R. W." box="[673,880,1278,1303]" pageId="27" pageNumber="198" pagination="1828 - 1830" refId="ref35989" refString="WIEDEMANN, C. R. W. 1828. Aussereuropaische zweiflugelige Insekten als Fortsetzung des Meigenschen Werkes v. 1. Pt 8. Familie: Rauberfliegen (Asilici). Hamburg: Schulzischen Buchhandlung, 1828 - 1830." type="book chapter" year="1828">Wiedemann, 1828</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3236543FFB76D4ABEB49D506601AE1A" pageId="27" pageNumber="198" type="reference_group">
<paragraph id="8B8636C8FFB76D4ABEB49D50661AAE2A" blockId="27.[181,1142,1324,1671]" box="[181,637,1324,1345]" pageId="27" pageNumber="198">
<treatmentCitationGroup id="AB2911E6FFB76D4ABEB49D50661AAE2A" box="[181,637,1324,1345]" pageId="27" pageNumber="198">
<taxonomicName id="4C394D4BFFB76D4ABEB49D506508AE2B" ID-CoL="8V48X" authority="Ruppelii" authorityName="Ruppelii" box="[181,367,1325,1345]" class="Insecta" family="Asilidae" genus="Dasypogon" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="198" phylum="Arthropoda" rank="genus">
<emphasis id="B94DEADAFFB76D4ABEB49D506508AE2B" box="[181,367,1325,1345]" italics="true" pageId="27" pageNumber="198">Dasypogon Rüppelii</emphasis>
</taxonomicName>
: 
<treatmentCitation id="0A9810D9FFB76D4ABF7A9D51661FAE2A" author="WIEDEMANN, C. R. W." box="[379,632,1324,1344]" page="569" pageId="27" pageNumber="198" year="1828">
<bibRefCitation id="EFA84B39FFB76D4ABF7A9D51661FAE2A" author="WIEDEMANN, C. R. W." box="[379,632,1324,1344]" pageId="27" pageNumber="198" pagination="1828 - 1830" refId="ref35989" refString="WIEDEMANN, C. R. W. 1828. Aussereuropaische zweiflugelige Insekten als Fortsetzung des Meigenschen Werkes v. 1. Pt 8. Familie: Rauberfliegen (Asilici). Hamburg: Schulzischen Buchhandlung, 1828 - 1830." type="book chapter" year="1828">Wiedemann 1828: 569–570</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8B8636C8FFB76D4ABEB49D386672AE32" blockId="27.[181,1142,1324,1671]" box="[181,533,1348,1369]" pageId="27" pageNumber="198">
<taxonomicName id="4C394D4BFFB76D4ABEB49D386676AE32" authority=": Oldroyd 1980: 352" authorityName=": Oldroyd" authorityPageNumber="352" authorityYear="1980" box="[181,529,1348,1369]" class="Insecta" family="Asilidae" genus="Laphria" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="198" phylum="Arthropoda" rank="species" species="rueppelii">
<emphasis id="B94DEADAFFB76D4ABEB49D386531AE33" box="[181,342,1349,1369]" italics="true" pageId="27" pageNumber="198">Laphria rueppelii</emphasis>
: Oldroyd 1980: 352
</taxonomicName>
.
</paragraph>
<paragraph id="8B8636C8FFB76D4ABEB49D206601AE1A" blockId="27.[181,1142,1324,1671]" box="[181,614,1372,1393]" pageId="27" pageNumber="198">
<treatmentCitationGroup id="AB2911E6FFB76D4ABEB49D206601AE1A" box="[181,614,1372,1393]" pageId="27" pageNumber="198">
<treatmentCitation id="0A9810D9FFB76D4ABEB49D206605AE1A" author="GELLER-GRIMM, F." box="[181,610,1372,1393]" page="214" pageId="27" pageNumber="198" year="1999">
<taxonomicName id="4C394D4BFFB76D4ABEB49D206605AE1A" authority=": Geller-Grimm 1999: 214" authorityName=": Geller-Grimm" authorityPageNumber="214" authorityYear="1999" box="[181,610,1372,1393]" class="Insecta" family="Asilidae" genus="Trichardis" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="198" phylum="Arthropoda" rank="species" species="rueppellii">
<emphasis id="B94DEADAFFB76D4ABEB49D206516AE1B" box="[181,369,1373,1393]" italics="true" pageId="27" pageNumber="198">Trichardis rueppellii</emphasis>
: 
<bibRefCitation id="EFA84B39FFB76D4ABF7C9D216605AE1A" author="GELLER-GRIMM, F." box="[381,610,1372,1392]" pageId="27" pageNumber="198" pagination="205 - 217" refId="ref35212" refString="GELLER-GRIMM, F. 1999. Raubfliegen-Typenmaterial des Senckenberg-Museums in Frankfurt am Main, das uberwiegend von Wiedemann und Jaennicke bearbeitet wurde (Insecta, Diptera, Asilidae). Senckenbergiana biologica 78 (1 / 2): 205 - 217." type="journal article" year="1999">Geller-Grimm 1999: 214</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C3236543FFB76D4DBEB49D0367D0AF7D" lastPageId="28" lastPageNumber="199" pageId="27" pageNumber="198" type="description">
<paragraph id="8B8636C8FFB76D4ABEB49D0366D8AEBD" blockId="27.[181,1142,1324,1671]" pageId="27" pageNumber="198">
Redescription (based on 
<typeStatus id="5482886AFFB76D4ABFCE9D036656AEFD" box="[463,561,1406,1431]" pageId="27" pageNumber="198" type="holotype">holotype</typeStatus>
in fair condition: both antennae broken off beyond pedicels, hind margins of wings damaged and not showing extent of costal vein, terminal tarsomeres mostly missing, damaged or dirty):
</paragraph>
<paragraph id="8B8636C8FFB76D4ABEB49D9B6028AD55" blockId="27.[181,1142,1324,1671]" pageId="27" pageNumber="198">
<emphasis id="B94DEADAFFB76D4ABEB49D9B6495AE94" box="[181,242,1510,1534]" italics="true" pageId="27" pageNumber="198">Head</emphasis>
: Pale orange, silver pruinescence, setae white. Antennae orange, both broken off beyond pedicel, white setose. Mystax white. Ocellar tubercle with 4 macrosetae and a few fairly big setae. Proboscis orange with orange-brown distal half, palpi orange.
</paragraph>
<paragraph id="8B8636C8FFB76D4DBEB49E33648CAA8D" blockId="27.[181,1142,1324,1671]" lastBlockId="28.[144,1105,206,1047]" lastPageId="28" lastPageNumber="199" pageId="27" pageNumber="198">
<emphasis id="B94DEADAFFB76D4ABEB49E336561AD0C" box="[181,262,1614,1638]" italics="true" pageId="27" pageNumber="198">Thorax</emphasis>
: Brown-orange, dorsal parts red-brown, setae white. Postpronotum entirely pruinose, mesonotum dark red-brown centrally, orange laterally, extensively silver pruinose except for central parts [may be worn smooth through handing?], setae white. Scutellum pruinose except for hind margin.Anepisternum lacking posterior macroseta, extensively pruinose (somewhat apruinose anteroventrally [may be worn]). Proepimeron, katepisternum and anepimeron pruinose. Legs: Uniformly orange, lengths of pulvilli and empodia difficult to study as terminal tarsomeres dirty, damaged or missing. Hind femur orange, length:height ratio 4.1:1 (i.e. slender), ventral tubercles not evident. Hind tibia lacking ventrodistal spur. Wing: 6.3× 
<quantity id="4CC19B2DFFB06D4DBC7299F366A9AACD" box="[627,718,398,423]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="28" pageNumber="199" unit="mm" value="2.4">2.4 mm</quantity>
. Costal veins broken and missing beyond wing tips [due to damaged hind margins]. Membrane entirely without microtrichiae.
</paragraph>
<paragraph id="8B8636C8FFB06D4DBE91998B672AA925" blockId="28.[144,1105,206,1047]" pageId="28" pageNumber="199">
<emphasis id="B94DEADAFFB06D4DBE91998B649EA964" box="[144,249,502,526]" italics="true" pageId="28" pageNumber="199">Abdomen</emphasis>
: Uniformly brown-orange, terga silver pruinose except for small areas anterolaterally and centrally [may be worn], white setose (setulae longish). T2 orange, pruinose except for small anterolateral and central [may be worn] areas.
</paragraph>
<paragraph id="8B8636C8FFB06D4DBE919A216505A9BA" blockId="28.[144,1105,206,1047]" pageId="28" pageNumber="199">
<materialsCitation id="3B513C95FFB06D4DBE919A216505A9BA" collectionCode="SMFD" collectorName="Ruppelii Wd. &amp; Dr. F. Herman'" country="Eritrea" location="ERITREA" pageId="28" pageNumber="199" specimenCount="1" typeStatus="holotype">
<typeStatus id="5482886AFFB06D4DBE919A216483A91A" box="[144,228,604,624]" pageId="28" pageNumber="199" type="holotype">Holotype</typeStatus>
(examined): 
<collectingCountry id="F32E7658FFB06D4DBF5F9A2165A6A91A" box="[350,449,604,624]" name="Eritrea" pageId="28" pageNumber="199">ERITREA</collectingCountry>
: 
<emphasis id="B94DEADAFFB06D4DBFCA9A206582A91A" box="[459,485,604,624]" italics="true" pageId="28" pageNumber="199">^‘</emphasis>
Abyssinia [no locality given] / Dr Rüppell.’, ‘136’ [blue with black frame], ‘Typus’ [red with black frame], ~ ‘Dipt. / 105’ [white], ‘ 
<taxonomicName id="4C394D4BFFB06D4DBCE59A086725A9E3" box="[740,834,629,649]" class="Insecta" family="Asilidae" genus="Trichardis" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B94DEADAFFB06D4DBCE59A086725A9E3" box="[740,834,629,649]" italics="true" pageId="28" pageNumber="199">Trichardis</emphasis>
</taxonomicName>
/ 
<collectorName id="26CC531EFFB06D4DBD579A0867B5A9E2" box="[854,978,628,649]" pageId="28" pageNumber="199">
<emphasis id="B94DEADAFFB06D4DBD579A0867C4A9E3" box="[854,931,629,649]" italics="true" pageId="28" pageNumber="199">Ruppelii</emphasis>
Wd.
</collectorName>
/ det. / 
<collectorName id="26CC531EFFB06D4DBA1D9A0964B9A9CA" pageId="28" pageNumber="199">Dr. F. Herman’</collectorName>
(
<collectionCode id="ED28AE0DFFB06D4DBEE99AF1654DA9CA" box="[232,298,652,672]" pageId="28" pageNumber="199">SMFD</collectionCode>
). Note: Wiedemann (1928) gives the provenance as ‘Aus Nubien’. This suggests the Nubian Desert which is in Eritrea and not Sudan, as listed by Oldroyd (1980). For the present it is not possible to provide a type-locality.
</materialsCitation>
</paragraph>
<paragraph id="8B8636C8FFB06D4DBE919AA367D0AF7D" blockId="28.[144,1105,206,1047]" pageId="28" pageNumber="199">
Taxonomic status: Morphologically the 
<typeStatus id="5482886AFFB06D4DBC599AA366DDA99D" box="[600,698,734,759]" pageId="28" pageNumber="199" type="holotype">holotype</typeStatus>
closely agrees with the description provided above for the 
<taxonomicName id="4C394D4BFFB06D4DBFA69A836645A87C" authorityName=": Engel" authorityYear="1924" box="[423,546,766,790]" class="Insecta" family="Asilidae" genus="Trichardis" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="199" phylum="Arthropoda" rank="species" species="leucocoma">
<emphasis id="B94DEADAFFB06D4DBFA69A836645A87C" box="[423,546,766,790]" italics="true" pageId="28" pageNumber="199">leucocoma</emphasis>
</taxonomicName>
type except for a few small details as follows. Pruinescence of head and thorax is not as strong or extensive; the legs are uniformly orange and totally lack red-brown parts. Bearing in mind that the 
<taxonomicName id="4C394D4BFFB06D4DBD689B4367A9A83C" authorityName=": Oldroyd" authorityYear="1980" box="[873,974,830,854]" class="Insecta" family="Asilidae" genus="Laphria" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="199" phylum="Arthropoda" rank="species" species="rueppelii">
<emphasis id="B94DEADAFFB06D4DBD689B4367A9A83C" box="[873,974,830,854]" italics="true" pageId="28" pageNumber="199">rueppelii</emphasis>
</taxonomicName>
<typeStatus id="5482886AFFB06D4DBDD59B436051A83D" box="[980,1078,830,855]" pageId="28" pageNumber="199" type="holotype">holotype</typeStatus>
is a female and that some sexual dimorphism is evident in 
<taxonomicName id="4C394D4BFFB06D4DBCF59B23670AA81C" authorityName=": Engel" authorityYear="1924" box="[756,877,862,886]" class="Insecta" family="Asilidae" genus="Trichardis" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="199" phylum="Arthropoda" rank="species" species="leucocoma">
<emphasis id="B94DEADAFFB06D4DBCF59B23670AA81C" box="[756,877,862,886]" italics="true" pageId="28" pageNumber="199">leucocoma</emphasis>
</taxonomicName>
everything points to 
<taxonomicName id="4C394D4BFFB06D4DBE919B03656EA8FC" authorityName=": Engel" authorityYear="1924" box="[144,265,894,918]" class="Insecta" family="Asilidae" genus="Trichardis" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="199" phylum="Arthropoda" rank="species" species="leucocoma">
<emphasis id="B94DEADAFFB06D4DBE919B03656EA8FC" box="[144,265,894,918]" italics="true" pageId="28" pageNumber="199">leucocoma</emphasis>
</taxonomicName>
being a synonym of 
<emphasis id="B94DEADAFFB06D4DBFFF9B03660DA8FC" box="[510,618,894,918]" italics="true" pageId="28" pageNumber="199">
<taxonomicName id="4C394D4BFFB06D4DBFFF9B036600A8FC" authorityName=": Oldroyd" authorityYear="1980" box="[510,615,894,918]" class="Insecta" family="Asilidae" genus="Laphria" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="199" phylum="Arthropoda" rank="species" species="rueppelii">rueppelii</taxonomicName>
.
</emphasis>
However, while I am reasonably sure that this will be the future taxonomic outcome, I refrain from establishing the synonymy until male specimens agreeing with 
<taxonomicName id="4C394D4BFFB06D4DBC239BC366FCA8BC" authorityName=": Engel" authorityYear="1924" box="[546,667,958,982]" class="Insecta" family="Asilidae" genus="Trichardis" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="199" phylum="Arthropoda" rank="species" species="leucocoma">
<emphasis id="B94DEADAFFB06D4DBC239BC366FCA8BC" box="[546,667,958,982]" italics="true" pageId="28" pageNumber="199">leucocoma</emphasis>
</taxonomicName>
are found in 
<collectingCountry id="F32E7658FFB06D4DBD319BC367E7A8BD" box="[816,896,958,983]" name="Eritrea" pageId="28" pageNumber="199">Eritrea</collectingCountry>
. This conservative approach also ensures retention of the well-known name, 
<emphasis id="B94DEADAFFB06D4DBD399BA367DEA89C" box="[824,953,990,1014]" italics="true" pageId="28" pageNumber="199">
<taxonomicName id="4C394D4BFFB06D4DBD399BA367D2A89C" authorityName=": Engel" authorityYear="1924" box="[824,949,990,1014]" class="Insecta" family="Asilidae" genus="Trichardis" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="199" phylum="Arthropoda" rank="species" species="leucocoma">leucocoma</taxonomicName>
,
</emphasis>
until further investigations of the Palaearctic 
<taxonomicName id="4C394D4BFFB06D4DBFFA9B836608AF7C" box="[507,623,1022,1046]" class="Insecta" family="Asilidae" genus="Trichardis" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="B94DEADAFFB06D4DBFFA9B836608AF7C" box="[507,623,1022,1046]" italics="true" pageId="28" pageNumber="199">Trichardis</emphasis>
</taxonomicName>
fauna have been undertaken.
</paragraph>
</subSubSection>
</treatment>
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