Taxonomic revision of a darkling beetles genus Anaxius (Tenebrionidae: Pedinini: Helopinina) Author Kamiński, Marcin J. Author Schoeman, Colin S. text Zootaxa 2018 2018-08-03 4455 3 471 485 journal article 29087 10.11646/zootaxa.4455.3.4 c9bf5059-abe4-414e-a7c2-f3f9e75f2979 1175-5326 1457467 69209FF7-80B0-409D-A257-0B88C65E903F Genus Anaxius Fåhraeus, 1870 Anaxius Fåhraeus, 1870 : 307 .- Koch 1958 : 208 . Type species . Anaxius obesus Fåhraeus, 1870 ; by monotypy. Diagnosis . According to Koch’s (1958) hypothesis, this genus is close to Aptila Fåhraeus, 1870 and Asidodema Koch, 1958 based on pronotal shape (subparallel sided; flat disc) and male terminalia (relatively short tegmen; clavae as large as penis). All three genera were previously classified within a single subtribe Aptilina. From Aptila and Asidodema , Anaxius can be distinguished by having widened protarsi and the apical segment of the maxillary palpus enlarged in males. Moreover, it differs from Aptila by the lack of scale-like vestiture on the upper surface of the body, while the relatively reduced prosternum differentiates it from Asidodema (see Koch 1958 ). Species included (7) . bloubergensis sp. nov. , campbellae Koch, 1958 , limpopoensis sp. nov. , meletsensis sp. nov. , obesus Fåhraeus, 1870 , prozeskyi Koch, 1958 , pseudoloensus sp. nov. Distribution . Representatives of this genus have been collected in the following ecoregions of Namibia , Mozambique , South Africa and Zimbabwe ( Fig. 5 ): Drakensberg montane grasslands, woodlands, and forests; Highveld grasslands; Kalahari xeric savanna; Southern Africa bushveld; and Southern Miombo woodlands. Anaxius was recorded from localities distributed between 874 and 1544 meters in elevation ( Fig. 6 ). Note . During the present investigation 49 Anaxius specimens were studied. Among this number a total of three females was identified. Because of this the following key includes only the male morphology.