<document id="8D34FA8D7EEF75F7DF1B5AD506FF6CAB" ID-DOI="10.37828/em.2025.87.7" ID-ISSN="2336-9744" ID-Zenodo-Dep="16958844" ID-ZooBank="066C2CD1-5023-41CB-8F2E-667532F6F199" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1756269000471" checkinUser="felipe" docAuthor="László, Gyula M." docDate="2025" docId="038A879EFFB08C61FF67FB2B5010FF97" docLanguage="en" docName="EcolMontenegrina.87.135-191.pdf" docOrigin="Ecologica Montenegrina 87" docSource="https://doi.org/10.37828/em.2025.87.7" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.3:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="3" docTitle="Meganola bispermutata Hacker 2012" docType="treatment" docVersion="2" lastPageNumber="154" masterDocId="FFB3FFE6FFA38C72FFDBFF9C5210F802" masterDocTitle="Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini)" masterLastPageNumber="191" masterPageNumber="135" pageNumber="154" updateTime="1756274687144" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="D2F3BCD333F72D100809850642CEB3E8">Integrative taxonomic review of the Meganola monofascia van Son species group sensu Hacker et al. (Lepidoptera, Noctuoidea, Nolidae, Nolinae, Nolini)</mods:title>
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<mods:namePart id="A4983EA9589AA27512EB63038745E9A7">László, Gyula M.</mods:namePart>
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<heading id="D0D481E4FFB08C61FF67FB2B50D4FCD3" box="[188,708,1206,1233]" centered="true" fontSize="11" level="2" pageId="19" pageNumber="154" reason="2">
<emphasis id="B957EA9AFFB08C61FF67FB2B50D4FCD3" bold="true" box="[188,708,1206,1233]" pageId="19" pageNumber="154">
The 
<taxonomicName id="4C234D0BFFB08C61FF2EFB2A5004FCD3" ID-CoL="DHKT7" authorityName="Hacker" authorityYear="2012" box="[245,532,1206,1233]" class="Insecta" family="Nolidae" genus="Meganola" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="154" phylum="Arthropoda" rank="species" species="bispermutata">
<emphasis id="B957EA9AFFB08C61FF2EFB2A5004FCD3" bold="true" box="[245,532,1206,1233]" italics="true" pageId="19" pageNumber="154">Meganola bispermutata</emphasis>
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species group
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<emphasis id="B957EA9AFFB08C61FF67FB66509FFD17" bold="true" box="[188,655,1274,1301]" pageId="19" pageNumber="154">Characterisation of the species group</emphasis>
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The facies of the taxa of this species group are confusingly similar to those of the 
<taxonomicName id="4C234D0BFFB08C61FB1EFAA35768FD5A" authorityName="sensu Hacker et al." authorityYear="2012" baseAuthorityName="van Son" baseAuthorityYear="1933" box="[1221,1400,1342,1368]" class="Insecta" family="Nolidae" genus="Meganola" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="154" phylum="Arthropoda" rank="species" species="monofascia">
<emphasis id="B957EA9AFFB08C61FB1EFAA35768FD5A" box="[1221,1400,1342,1368]" italics="true" pageId="19" pageNumber="154">M. monofascia</emphasis>
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complex, featuring a well-developed, blackish, broad-based triangular medio-costal patch and a variably developed medial fascia of the forewing. The shared diagnostic male genitalia characters of the species group are the relatively narrow, elongate, apically evenly rounded valva, the variably long, apically pointed harpe and the variably long aedeagus possessing a characteristic rounded carina with a small subapical ventral tooth. In the female genitalia, the species of this complex display a very short sclerotised sinus vaginalis, a heavily sclerotised distal section of the ductus bursae and a single, short and rounded signum bursae accompanied distally by a longitudinal scobinated band.
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<emphasis id="B957EA9AFFB08C61FF67F9F15361FE85" bold="true" box="[188,369,1645,1671]" pageId="19" pageNumber="154">Taxon content</emphasis>
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<taxonomicName id="4C234D0BFFB08C61FF67F90C50D6FEAB" authority="Hacker, 2012" authorityName="Hacker" authorityYear="2012" box="[188,710,1679,1705]" class="Insecta" family="Nolidae" genus="Meganola" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="154" phylum="Arthropoda" rank="subSpecies" species="bispermutata" subSpecies="bispermutata">
<emphasis id="B957EA9AFFB08C61FF67F90C5031FEAB" box="[188,545,1679,1705]" italics="true" pageId="19" pageNumber="154">M. bispermutata bispermutata</emphasis>
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<taxonomicName id="4C234D0BFFB08C61FF67F92E50BCFEC9" authority="Hacker, 2012" authorityName="Hacker" authorityYear="2012" box="[188,684,1713,1739]" class="Insecta" family="Nolidae" genus="Meganola" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="154" phylum="Arthropoda" rank="subSpecies" species="bispermutata" subSpecies="cryptochra">
<emphasis id="B957EA9AFFB08C61FF67F92E5016FEC9" box="[188,518,1713,1739]" italics="true" pageId="19" pageNumber="154">M. bispermutata cryptochra</emphasis>
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<emphasis id="B957EA9AFFB08C61FD68F92D5117FEC9" bold="true" box="[691,775,1713,1739]" pageId="19" pageNumber="154">stat. n.</emphasis>
<taxonomicName id="4C234D0BFFB08C61FF67F9485013FEEF" ID-CoL="DHKVJ" authority="Hacker, 2012" authorityName="Hacker" authorityYear="2012" box="[188,515,1747,1773]" class="Insecta" family="Nolidae" genus="Meganola" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="154" phylum="Arthropoda" rank="species" species="diagrapha">
<emphasis id="B957EA9AFFB08C61FF67F948534EFEEF" box="[188,350,1747,1773]" italics="true" pageId="19" pageNumber="154">M. diagrapha</emphasis>
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<taxonomicName id="4C234D0BFFB08C61FF67F9695362FF0C" ID-CoL="KZM8X" authorityName="László" authorityYear="2025" box="[188,370,1781,1806]" class="Insecta" family="Nolidae" genus="Meganola" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="154" phylum="Arthropoda" rank="species" species="saotomeana" status="sp. n.">
<emphasis id="B957EA9AFFB08C61FF67F9695362FF0C" box="[188,370,1781,1806]" italics="true" pageId="19" pageNumber="154">M. saotomeana</emphasis>
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<taxonomicNameLabel id="A26457E1FFB08C61FEA2F96853ABFF0C" box="[377,443,1780,1806]" pageId="19" pageNumber="154" rank="species">sp. n.</taxonomicNameLabel>
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<taxonomicName id="4C234D0BFFB08C61FF67F88B5034FF32" ID-CoL="DHL2W" authority="Hacker, 2012" authorityName="Hacker" authorityYear="2012" box="[188,548,1814,1840]" class="Insecta" family="Nolidae" genus="Meganola" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="154" phylum="Arthropoda" rank="species" species="pachygrapha">
<emphasis id="B957EA9AFFB08C61FF67F88B536FFF32" box="[188,383,1814,1840]" italics="true" pageId="19" pageNumber="154">M. pachygrapha</emphasis>
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<taxonomicName id="4C234D0BFFB08C61FF67F8A553FEFF50" ID-CoL="DHKV9" authority="Hacker, 2012" authorityName="Hacker" authorityYear="2012" box="[188,494,1848,1874]" class="Insecta" family="Nolidae" genus="Meganola" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="154" phylum="Arthropoda" rank="species" species="dananae">
<emphasis id="B957EA9AFFB08C61FF67F8A55359FF50" box="[188,329,1848,1874]" italics="true" pageId="19" pageNumber="154">M. dananae</emphasis>
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</taxonomicName>
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<taxonomicName id="4C234D0BFFB08C61FF67F8C6533CFF71" ID-CoL="KZMGF" authorityName="László" authorityYear="2025" box="[188,300,1881,1907]" class="Insecta" family="Nolidae" genus="Meganola" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="154" phylum="Arthropoda" rank="species" species="baaka" status="sp. n.">
<emphasis id="B957EA9AFFB08C61FF67F8C6533CFF71" box="[188,300,1881,1907]" italics="true" pageId="19" pageNumber="154">M. baaka</emphasis>
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<emphasis id="B957EA9AFFB08C61FEE8F8C65365FF76" bold="true" box="[307,373,1882,1908]" pageId="19" pageNumber="154">
<taxonomicNameLabel id="A26457E1FFB08C61FEE8F8C65365FF76" box="[307,373,1882,1908]" pageId="19" pageNumber="154" rank="species">sp. n.</taxonomicNameLabel>
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<paragraph id="8B9C3688FFB08C61FF67F8E05010FF97" blockId="19.[188,775,1645,1941]" box="[188,512,1915,1941]" pageId="19" pageNumber="154">
<taxonomicName id="4C234D0BFFB08C61FF67F8E05010FF97" ID-CoL="DHL57" authority="Hacker, 2012" authorityName="Hacker" authorityYear="2012" box="[188,512,1915,1941]" class="Insecta" family="Nolidae" genus="Meganola" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="154" phylum="Arthropoda" rank="species" species="subderrae">
<emphasis id="B957EA9AFFB08C61FF67F8E0534BFF97" box="[188,347,1915,1941]" italics="true" pageId="19" pageNumber="154">M. subderrae</emphasis>
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