Current knowledge on the velvet ants (Hymenoptera: Mutillidae, Myrmosidae) of Vietnam: faunal review and new distributional records Author Okayasu, Juriya Institute for Integrated Innovations, Hokkaido University, North 12 West 7, Sapporo, 060 - 0812 Japan Author Lelej, Arkady S. Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, Vladivostok, 690022, Russia lelej @ biosoil. ru; https: // orcid. org / 0000 - 0001 - 7501 - 0981 lelej@biosoil.ru Author Williams, Kevin A. Plant Pest Diagnostic Center, California Department of Food & Agriculture, 3294 Meadowview Road, Sacramento, CA 95832, USA Kevin. Williams @ cdfa. ca. gov; https: // orcid. org / 0000 - 0002 - 6073 - 1070 Williams@cdfa.ca.gov Author Brabant, Craig M. Department of Entomology, University of Wisconsin-Madison, 1630 Linden Drive, Madison, WI 53706, USA brabant @ entomology. wisc. edu; https: // orcid. org / 0000 - 0002 - 1752 - 5597 brabant@entomology.wisc.edu Author Nguyen, Lien Thi Phuong Institute of Biology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam & Graduate University of Science and Technology, Vietnam Academy of Science & Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam Author Nguyen, Anh Duc Institute of Biology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam & Graduate University of Science and Technology, Vietnam Academy of Science & Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam Author Pham, Thai Hong Systematic Entomology, Research Faculty of Agriculture, Hokkaido University, North 9 West 9, Sapporo, 060 - 8589 Japan mutiphiidae @ gmail. com; https: // orcid. org / 0000 - 0002 - 6307 - 2939 & Mientrung Institute for Scientific Research, Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 321 Huynh Thuc Khang, Hue, Vietnam & phamthai @ vnmn. vast. vn; https: // orcid. org / 0000 - 0002 - 4763 - 3679 mutiphiidae@gmail.com Author Truong, Tuong Thi Cat Systematic Entomology, Research Faculty of Agriculture, Hokkaido University, North 9 West 9, Sapporo, 060 - 8589 Japan mutiphiidae @ gmail. com; https: // orcid. org / 0000 - 0002 - 6307 - 2939 & Mientrung Institute for Scientific Research, Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 321 Huynh Thuc Khang, Hue, Vietnam & Systematic Entomology, Research Faculty of Agriculture, Hokkaido University, North 9 West 9, Sapporo, 060 - 8589 Japan mutiphiidae @ gmail. com; https: // orcid. org / 0000 - 0002 - 6307 - 2939 & tuong. tuong @ gmail. com; https: // orcid. org / 0000 - 0002 - 8593 - 6283 * Corresponding author mutiphiidae@gmail.com text Zootaxa 2025 2025-07-31 5668 1 1 104 https://doi.org/10.11646/zootaxa.5668.1.1 journal article 10.11646/zootaxa.5668.1.1 1175-5334 E1D39158-B8CC-4E80-B610-F70255ECAF60 Orientidia thoawanae Williams, 2019 urn:lsid:zoobank.org:act: 5011918C-07A3-49C4-9D8A-9A13A3C31E4B ( Figs 22C, 22D ) Orientidia thoawanae Williams in Williams et al ., 2019b: 38 , ♀ , holotype ♀ ( Prince of Songkhla University , Hat Yai , Songkhla , Thailand ) [ CSCA ]; Thaochan et al . 2022: 163 , ♀ ; Boutin & Vilhelmsen 2024 : supplementary table S2, S3, ♀ . Orientidia thoawani [sic]: Pagliano et al . 2020: 254 . Incorrect subsequent spelling. Material examined. 7♀ . Vietnam : QUANG NINH : Dong Kho Island , 21°06'N 107°36'E , 20.III.1987 , V. Kuznetzov leg. [ 2♀ IBSS ]; Dong Kho Island , litter, 21.III.1987 , L. Filatova leg. [ 1♀ IBSS ] ; HOA BINH : Ha Son Binh Prov. [= Hoa Binh], Da Bac , Tuly [= Tu Ly], forest, 19.X.1990 , S. Belokobylskij leg. [ 1♀ IBSS ]; Ky Son, Cao Phong, forest, 28.X.1990 , S. Belokobylskij leg. [ 1♀ IBSS ] ; NINH BINH : Cuc Phuong , 27.IV.1996 , Y. Okushima leg. [ 1♀ TUA ]; Nho Quan , Cuc Phuong National Park , 20.354N , 105.588E , 500 m alt., 8.VIII.2016 , T. Mita leg., 314/BTTNVN [ 1♀ VNMN ] . Identification source. Keyed using Williams et al . (2019b) and compared with the original description. Previous record. None. Distribution. Malaysia . Thailand . Vietnam *: Quang Ninh , Hoa Binh , Ninh Binh . Occurrence in Vietnam . March, April, August, October. South China-Vietnam subtropical evergreen forests, Northern Indochina subtropical forests, and Northern Vietnam lowland rain forests ecoregions. 500 m alt. Forest. Sex association. Male unknown. Remarks. Unlike other Orientidia females, this species has a narrow spine-like scutellar scale. Discovery of the male could confirm its placement in Orientidia or necessitate its transfer to a different genus, possibly Promecidia Lelej, 1996 . Genus Pagdenidia Lelej, 1996 urn:lsid:zoobank.org:act: D8D30846-46C8-41F0-84ED-793E361FC57C Pagdenidia Lelej, 1996b: 7 , ♂ ♀ ; Lelej 2005: 69; Lelej & Brothers 2008: 42 ; Pagliano et al . 2020: 254 . Type species: Timulla ( Trogaspidia ) mickeli Pagden, 1949 ( ♂ ♀ , holotype ♂ ), by original designation. Diversity and distribution. This Oriental genus includes seven species (Lelej 2005; Pagliano et al . 2020 ). Remarks. Of the seven Pagdenidia species , six species are known from the male only and one species, Pa . mickeli , is known from both sexes. This genus was reviewed by Pagden (1949) as the Timulla ( Trogaspidia ) erato group. His identification key includes six species known at that time.