New species of Ochodaeus Dejean from Madagascar II (Coleoptera: Scarabaeoidea: Ochodaeidae) Author Paulsen, M. J. text Insecta Mundi 2019 2019-05-31 706 706 1 10 journal article 10.5281/zenodo.3673403 f01c9bd1-7635-40c7-b5ee-5fc57320b880 1942-1354 3673403 08CB0DA0-C83E-438F-AA97-8CD637F044A5 Updated key couplets to Ochodaeus spp. in Madagascar The following couplet changes to the species key in Paulsen (2019) allow for determination of all Madagascan species having a granulate pronotum, including the three newly described species. 2(1). Elytral intervals with setose tubercles of 2 sizes, with setae of larger tubercles flattened, scale-like........................................................ O. pygmaeus Paulian — Elytral intervals with tubercles either absent or of uniform size and without flattened setae. 3 3(2). Clypeus with median tubercle or small horn that is distinct from anterior margin ( Fig. 4–6 )........................................................................... 4 — Clypeus lacking free tubercle or horn, anterior margin usually thickened medially ( Fig. 7–8 ). 6 4(3). Mentum with a knobby protuberance on each side ( Fig. 1d ).... O. meridialis Paulsen , n. sp. — Mentum lacking protuberances laterally............................................ 5 Figures 4–8. Key characters of Madagascar Ochodaeus species. 4) Head of O. meridialis Paulsen , n. sp. , showing central tubercle of clypeus. 5) Head of O. umbonulus Paulsen , n. sp. , showing tubercle anterior to straight frontoclypeal suture. 6) Head of O. iniquipes Paulsen , n. sp. , showing minute horn on posteriorly curved frontoclypeal suture. 7) Head of O. isoanalensis Paulian showing broadly thickened clypeal margin. 8) Head of O. miliaris (Klug) , showing narrowly thickened clypeal margin. 5(4). Clypeus ( Fig. 5 ) with median tubercle anterior to frontoclypeal suture; anterior margin of clypeus indistinct and eroded medially........................ O. umbonulus Paulsen , n. sp. — Clypeus ( Fig. 6 ) with small horn located on frontoclypeal suture; anterior margin of clypeus distinct and entire throughout.......................... O. iniquipes Paulsen , n. sp. 6(3). Median thickening of clypeal bead variable, lateral thickening always evident ( Fig. 7 ); body length usually> 9.0 mm........................................ O. isoanalensis Paulian — Median thickening of clypeal bead simply convex, bead slender laterally ( Fig. 8 ); body length usually 4.5–8.0 mm............................................ O. miliaris Klug