Ratcliffe, Brett C., 2002, A checklist of the Scarabaeoidea (Coleoptera) of Panama

Ratcliffe, Brett C., 2002, A checklist of the Scarabaeoidea (Coleoptera) of Panama, Zootaxa 32, pp. 1-48

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Aesalini     5
Lucanini     5
Passalini     5-6
Proculini     6
Ceratocanthinae     7-8
Ochodaeidae     7
Hybosoridae     7
Troginae     8-9
Bolboceratini     9
Athyreini     9
Aphodiini     9-10
Eupariini     10-11
Rhypariini     11
Eurysternini     11
Canthonini     12-13
Ateuchini     13-15
Coprini     15-16
Phanaeini     16-17
Onthophagini     17
Orphninae     18
Sericini     18
Melolonthini     18-19
Macrodactylini     19-20
Hopliini     20
Rutelini     20-24
Anomalini     24-26
Spodochlamyini     26
Anoplognathini     26
Geniatini     26-27
Cyclocephalini     27-31
Pentodontini     31-33
Oryctini     33-35
Phileurini     35-36
Agaocephalini     36
Dynastini     36-37
Gymnetini     38-39
Cetoniini     39
Cremastocheilini     39
Trichiini     39-40