Tomarus, Taxa Only, Exact Match: 93 Treatments

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Coleoptera   new species  Grouvelle, A., 1913, H. Sauter’s Formosa-Ausbeute (Rhysodidae, Nitidulidae, Ostomidae, Colydiidae, Passandridae, Cucujidae, Cryptophagidae, Diphyllidae, Lathridiidae, Mycetophagidae, Dermestidae), Archiv für Naturgeschichte, Abteilung A 79, pp. 33-76 : 66 66
Tomarus maimon     Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36 : 7-8 7-8
Tomarus fossor     Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, Insecta Mundi 732 (732), pp. 1-306 : 212 212
Tomarus rostratus     López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision, Zootaxa 5211 (1), pp. 1-119 : 70 70
Tomarus roigjunenti     López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision, Zootaxa 5211 (1), pp. 1-119 : 69-70 69-70
Tomarus     López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán, 2015, The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution, Zootaxa 4048 (4), pp. 451-492 : 476 476
Tomarus maimon     Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36 : 8 8
Tomarus fossor     Taboada-Verona, Carlos, Sermeño-Correa, Carlos, Sierra-Serrano, Oscar & Noriega, Jorge Ari, 2019, Checklist of the superfamily Scarabaeoidea (Insecta, Coleoptera) in an urban area of the Caribbean Colombia, Check List 15 (4), pp. 579-594 : 584-585 584-585
Tomarus     Wijayawardene, Nalin N., Dai, Dong-Qin, Premarathne, Bhagya M., Wimalasena, Madhara K., Jayalal, Udeni, Wickramanayake, Kawmini D., Dangalla, Hasanka, Jayathunga, Hashini, Brahmanage, Rashika S., Karunarathna, Samantha C., Weerakoon, Gothamie, Ariyawansa, Kahandawa G. S. U., Yapa, Neelamanie, Madawala, Sumedha, Nanayakkara, Chandrika M., Fan, Xin-Lei, Kirk, Paul M., Zhang, Gui-Qing, Ediriweera, Aseni, Bhat, Jayarama, Dawoud, Turki M. & Tibpromma, Saowaluck, 2023, Checklist, typification details, and nomenclature status of ascomycetous fungi originally described in Sri Lanka, Phytotaxa 611 (1), pp. 1-105 : 32 32
Tomarus similis     López-García, Margarita M. & Deloya, Cuauhtémoc, 2018, Description of the Female of Tomarus rostratus Dupuis and New Synonyms and Records for the Genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae), The Coleopterists Bulletin 72 (2), pp. 331-338 : 336 336
Tomarus fossor     López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán, 2015, The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution, Zootaxa 4048 (4), pp. 451-492 : 482-484 482-484
Tomarus pilcopataensis     López-García, Margarita M. & Deloya, Cuauhtémoc, 2019, Five New Species of the Dynastine Genus Tomarus Erichson (Coleoptera: Scarabaeidae), with an Illustrated Key to Species, The Coleopterists Bulletin 73 (1), pp. 127-141 : 134 134
Tomarus similis     Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, Insecta Mundi 732 (732), pp. 1-306 : 214 214
Tomarus cicatricosus     López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán, 2015, The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution, Zootaxa 4048 (4), pp. 451-492 : 479-480 479-480
Tomarus bituberculatus     Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, Insecta Mundi 732 (732), pp. 1-306 : 210-211 210-211
Tomarus rostratus     López-García, Margarita M., Giraldo-Mendoza, Alfredo E. & Deloya, Cuauhtémoc, 2019, Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru, Zootaxa 4604 (3), pp. 428-440 : 435-436 435-436
Tomarus pumilus     López-García, Margarita M. & Deloya, Cuauhtémoc, 2018, Description of the Female of Tomarus rostratus Dupuis and New Synonyms and Records for the Genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae), The Coleopterists Bulletin 72 (2), pp. 331-338 : 334 334
Tomarus pullus     López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán, 2015, The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution, Zootaxa 4048 (4), pp. 451-492 : 487-488 487-488
Tomarus villosus     Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36 : 13 13
Tomarus colombianus   sp. nov.  López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán, 2014, A new species of Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini), with a key to the species in Colombia, Zootaxa 3869 (5), pp. 579-584 : 580-583 580-583
Tomarus amazonicus   new status and combination  López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision, Zootaxa 5211 (1), pp. 1-119 : 55-56 55-56
Tomarus adoceteus     Ratcliffe, Brett C. & Cave, Ronald D., 2010, The Dynastinae (Coleoptera: Scarabaeidae) of the Cayman Islands (West Indies), with descriptions of Tomarus adoceteus, new species (Pentodontini) and Caymania nitidissima, new genus and species (Phileurini), Insecta Mundi 2010 (139), pp. 1-15 : 8 8
Tomarus laticaudus     López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision, Zootaxa 5211 (1), pp. 1-119 : 63-64 63-64
Tomarus maimon     López-García, Margarita M. & Deloya, Cuauhtémoc, 2018, Description of the Female of Tomarus rostratus Dupuis and New Synonyms and Records for the Genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae), The Coleopterists Bulletin 72 (2), pp. 331-338 : 335 335
Tomarus similis     López-García, Margarita M., Giraldo-Mendoza, Alfredo E. & Deloya, Cuauhtémoc, 2019, Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru, Zootaxa 4604 (3), pp. 428-440 : 436 436
Tomarus subtropicus     López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision, Zootaxa 5211 (1), pp. 1-119 : 71-72 71-72
Tomarus maternus     López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán, 2015, The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution, Zootaxa 4048 (4), pp. 451-492 : 485-486 485-486
Tomarus gyas     Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36 : 6-7 6-7
Tomarus key    Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, Insecta Mundi 732 (732), pp. 1-306 : 210 210
Tomarus     López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision, Zootaxa 5211 (1), pp. 1-119 : 52-54 52-54
Tomarus bituberculatus     López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision, Zootaxa 5211 (1), pp. 1-119 : 56-58 56-58
Tomarus     Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36 : 14-15 14-15
Tomarus cuniculus     Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, Insecta Mundi 732 (732), pp. 1-306 : 211 211
Tomarus ebenus     Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, Insecta Mundi 732 (732), pp. 1-306 : 212 212
Tomarus maimon     López-García, Margarita M., Giraldo-Mendoza, Alfredo E. & Deloya, Cuauhtémoc, 2019, Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru, Zootaxa 4604 (3), pp. 428-440 : 432-433 432-433
Tomarus ebenus     López-García, Margarita M., Giraldo-Mendoza, Alfredo E. & Deloya, Cuauhtémoc, 2019, Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru, Zootaxa 4604 (3), pp. 428-440 : 430-431 430-431
Tomarus pullus     López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision, Zootaxa 5211 (1), pp. 1-119 : 67-68 67-68
Tomarus cuniculus     Ratcliffe, Brett C. & Cave, Ronald D., 2010, The Dynastinae (Coleoptera: Scarabaeidae) of the Cayman Islands (West Indies), with descriptions of Tomarus adoceteus, new species (Pentodontini) and Caymania nitidissima, new genus and species (Phileurini), Insecta Mundi 2010 (139), pp. 1-15 : 9 9
Tomarus cuniculus     Gillett, Conrad P. D. T. & Gillett, Michael P. T., 2015, The Dynastinae of the island of Saba, Dutch Caribbean (Coleoptera: Scarabaeidae), Insecta Mundi 2015 (433), pp. 1-9 : 4-6 4-6
Tomarus discrepans     López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision, Zootaxa 5211 (1), pp. 1-119 : 59-60 59-60
Tomarus bidentulus     Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36 : 4-5 4-5
Tomarus rubripes     Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36 : 11 11
Tomarus maracaiboensis     López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision, Zootaxa 5211 (1), pp. 1-119 : 65-66 65-66
Tomarus gyas     López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision, Zootaxa 5211 (1), pp. 1-119 : 60-62 60-62
Tomarus     Ratcliffe, Brett C. & Cave, Ronald D., 2010, The Dynastinae (Coleoptera: Scarabaeidae) of the Cayman Islands (West Indies), with descriptions of Tomarus adoceteus, new species (Pentodontini) and Caymania nitidissima, new genus and species (Phileurini), Insecta Mundi 2010 (139), pp. 1-15 : 6-7 6-7
Tomarus cuniculus     López-García, Margarita M. & Deloya, Cuauhtémoc, 2018, Description of the Female of Tomarus rostratus Dupuis and New Synonyms and Records for the Genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae), The Coleopterists Bulletin 72 (2), pp. 331-338 : 335 335
Tomarus rostratus     López-García, Margarita M. & Deloya, Cuauhtémoc, 2018, Description of the Female of Tomarus rostratus Dupuis and New Synonyms and Records for the Genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae), The Coleopterists Bulletin 72 (2), pp. 331-338 : 332-333 332-333
Tomarus gibbosus     Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, Insecta Mundi 2020 (787), pp. 1-83 : 48 48
Tomarus     Meurgey, François & Ramage, Thibault, 2020, Challenging the Wallacean shortfall: A total assessment of insect diversity on Guadeloupe (French West Indies), a checklist and bibliography, Insecta Mundi 2020 (786), pp. 1-183 : 74 74
Tomarus laticaudus     López-García, Margarita M. & Deloya, Cuauhtémoc, 2019, Five New Species of the Dynastine Genus Tomarus Erichson (Coleoptera: Scarabaeidae), with an Illustrated Key to Species, The Coleopterists Bulletin 73 (1), pp. 127-141 : 129-130 129-130
Tomarus pilcopataensis     López-García, Margarita M., Giraldo-Mendoza, Alfredo E. & Deloya, Cuauhtémoc, 2019, Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru, Zootaxa 4604 (3), pp. 428-440 : 435 435
Tomarus villosus     López-García, Margarita M., Giraldo-Mendoza, Alfredo E. & Deloya, Cuauhtémoc, 2019, Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru, Zootaxa 4604 (3), pp. 428-440 : 436-438 436-438
Tomarus bituberculatus     López-García, Margarita M., Giraldo-Mendoza, Alfredo E. & Deloya, Cuauhtémoc, 2019, Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru, Zootaxa 4604 (3), pp. 428-440 : 429-430 429-430
Tomarus rosettae     Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36 : 10 10
Tomarus rosettae     Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36 : 10 10
Tomarus cuniculus     Ratcliffe, Brett C. & Cave, Ronald D., 2008, The Dynastinae (Coleoptera: Scarabaeidae) of the Bahamas with a description of a new species of Cyclocephala from Great Inagua Island, Insecta Mundi 2008 (24), pp. 1-10 : 6 6
Tomarus gibbosus     Ratcliffe, Brett C. & Cave, Ronald D., 2008, The Dynastinae (Coleoptera: Scarabaeidae) of the Bahamas with a description of a new species of Cyclocephala from Great Inagua Island, Insecta Mundi 2008 (24), pp. 1-10 : 7 7
Tomarus pullus     Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, Insecta Mundi 732 (732), pp. 1-306 : 213-214 213-214
Tomarus relictus     Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, Insecta Mundi 2020 (787), pp. 1-83 : 48 48
Tomarus maternus     López-García, Margarita M., Giraldo-Mendoza, Alfredo E. & Deloya, Cuauhtémoc, 2019, Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru, Zootaxa 4604 (3), pp. 428-440 : 433-434 433-434
Tomarus burmeisteri     Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36 : 5-6 5-6
Tomarus maimon     López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision, Zootaxa 5211 (1), pp. 1-119 : 64-65 64-65
Tomarus moroni     López-García, Margarita M. & Deloya, Cuauhtémoc, 2019, Five New Species of the Dynastine Genus Tomarus Erichson (Coleoptera: Scarabaeidae), with an Illustrated Key to Species, The Coleopterists Bulletin 73 (1), pp. 127-141 : 131 131
Tomarus maracaiboensis     López-García, Margarita M. & Deloya, Cuauhtémoc, 2019, Five New Species of the Dynastine Genus Tomarus Erichson (Coleoptera: Scarabaeidae), with an Illustrated Key to Species, The Coleopterists Bulletin 73 (1), pp. 127-141 : 130 130
Tomarus     López-García, Margarita M. & Deloya, Cuauhtémoc, 2019, Five New Species of the Dynastine Genus Tomarus Erichson (Coleoptera: Scarabaeidae), with an Illustrated Key to Species, The Coleopterists Bulletin 73 (1), pp. 127-141 : 135-139 135-139
Tomarus key    López-García, Margarita M., Giraldo-Mendoza, Alfredo E. & Deloya, Cuauhtémoc, 2019, Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru, Zootaxa 4604 (3), pp. 428-440 : 438 438
Tomarus roigjunenti   sp. nov.  Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36 : 8-10 8-10
Tomarus selanderi     López-García, Margarita M. & Deloya, Cuauhtémoc, 2018, Description of the Female of Tomarus rostratus Dupuis and New Synonyms and Records for the Genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae), The Coleopterists Bulletin 72 (2), pp. 331-338 : 335-336 335-336
Tomarus ebenus     López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán, 2015, The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution, Zootaxa 4048 (4), pp. 451-492 : 481-482 481-482
Tomarus rubripes     López-García, Margarita M. & Deloya, Cuauhtémoc, 2018, Description of the Female of Tomarus rostratus Dupuis and New Synonyms and Records for the Genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae), The Coleopterists Bulletin 72 (2), pp. 331-338 : 336 336
Tomarus selanderi     López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision, Zootaxa 5211 (1), pp. 1-119 : 70-71 70-71
Tomarus bidentulus     López-García, Margarita M. & Deloya, Cuauhtémoc, 2019, Five New Species of the Dynastine Genus Tomarus Erichson (Coleoptera: Scarabaeidae), with an Illustrated Key to Species, The Coleopterists Bulletin 73 (1), pp. 127-141 : 128 128
Tomarus peruvianus     López-García, Margarita M., Giraldo-Mendoza, Alfredo E. & Deloya, Cuauhtémoc, 2019, Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru, Zootaxa 4604 (3), pp. 428-440 : 434 434
Tomarus laevicollis     López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision, Zootaxa 5211 (1), pp. 1-119 : 62-63 62-63
Tomarus adoceteus     López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision, Zootaxa 5211 (1), pp. 1-119 : 55 55
Tomarus bituberculatus     López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán, 2015, The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution, Zootaxa 4048 (4), pp. 451-492 : 477-479 477-479
Tomarus gyas     López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán, 2015, The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution, Zootaxa 4048 (4), pp. 451-492 : 484-485 484-485
Tomarus gyas     Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, Insecta Mundi 732 (732), pp. 1-306 : 213 213
Tomarus gyas     López-García, Margarita M., Giraldo-Mendoza, Alfredo E. & Deloya, Cuauhtémoc, 2019, Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru, Zootaxa 4604 (3), pp. 428-440 : 431-432 431-432
Tomarus colombianus     López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán, 2015, The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution, Zootaxa 4048 (4), pp. 451-492 : 480 480
Tomarus maternus     López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision, Zootaxa 5211 (1), pp. 1-119 : 66-67 66-67
Tomarus pilcopataensis     López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision, Zootaxa 5211 (1), pp. 1-119 : 67 67
Tomarus paranaensis     López-García, Margarita M. & Deloya, Cuauhtémoc, 2019, Five New Species of the Dynastine Genus Tomarus Erichson (Coleoptera: Scarabaeidae), with an Illustrated Key to Species, The Coleopterists Bulletin 73 (1), pp. 127-141 : 133 133
Tomarus colombianus     López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision, Zootaxa 5211 (1), pp. 1-119 : 59 59
Tomarus maimon     Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, Insecta Mundi 732 (732), pp. 1-306 : 213 213
Tomarus pumilus     López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision, Zootaxa 5211 (1), pp. 1-119 : 68-69 68-69
Tomarus adoceteus   sp. nov.  Ratcliffe, Brett C. & Cave, Ronald D., 2010, The Dynastinae (Coleoptera: Scarabaeidae) of the Cayman Islands (West Indies), with descriptions of Tomarus adoceteus, new species (Pentodontini) and Caymania nitidissima, new genus and species (Phileurini), Insecta Mundi 2010 (139), pp. 1-15 : 7-8 7-8
Tomarus key    López-García, Margarita M. & Deloya, Cuauhtémoc, 2022, Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision, Zootaxa 5211 (1), pp. 1-119 : 16 16
Tomarus similis     López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán, 2015, The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution, Zootaxa 4048 (4), pp. 451-492 : 488-489 488-489
Tomarus spinipenis   sp. nov.  Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36 : 11-12 11-12
Tomarus spinipenis     Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36 : 12 12
Tomarus villosus     Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36 : 12 12
Tomarus     Gasca-Álvarez, Héctor & Amat-García, Germán, 2010, Synopsis and key to the genera of Dynastinae (Coleoptera, Scarabaeoidea, Scarabaeidae) of Colombia, ZooKeys 34 (34), pp. 153-192 : 185-186 185-186

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