Tomarus, Taxa Only, Exact Match: 72 Treatments

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Tomarus pilcopataensis     López-García, Margarita M. & Deloya, Cuauhtémoc, 2019, Five New Species of the Dynastine Genus Tomarus Erichson (Coleoptera: Scarabaeidae), with an Illustrated Key to Species, The Coleopterists Bulletin 73 (1), pp. 127-141: 134 134
Tomarus paranaensis     López-García, Margarita M. & Deloya, Cuauhtémoc, 2019, Five New Species of the Dynastine Genus Tomarus Erichson (Coleoptera: Scarabaeidae), with an Illustrated Key to Species, The Coleopterists Bulletin 73 (1), pp. 127-141: 133 133
Tomarus     López-García, Margarita M. & Deloya, Cuauhtémoc, 2019, Five New Species of the Dynastine Genus Tomarus Erichson (Coleoptera: Scarabaeidae), with an Illustrated Key to Species, The Coleopterists Bulletin 73 (1), pp. 127-141: 135-139 135-139
Tomarus bidentulus     López-García, Margarita M. & Deloya, Cuauhtémoc, 2019, Five New Species of the Dynastine Genus Tomarus Erichson (Coleoptera: Scarabaeidae), with an Illustrated Key to Species, The Coleopterists Bulletin 73 (1), pp. 127-141: 128 128
Tomarus moroni     López-García, Margarita M. & Deloya, Cuauhtémoc, 2019, Five New Species of the Dynastine Genus Tomarus Erichson (Coleoptera: Scarabaeidae), with an Illustrated Key to Species, The Coleopterists Bulletin 73 (1), pp. 127-141: 131 131
Tomarus     Meurgey, François & Ramage, Thibault, 2020, Challenging the Wallacean shortfall: A total assessment of insect diversity on Guadeloupe (French West Indies), a checklist and bibliography, Insecta Mundi 2020 (786), pp. 1-183: 74 74
Tomarus laticaudus     López-García, Margarita M. & Deloya, Cuauhtémoc, 2019, Five New Species of the Dynastine Genus Tomarus Erichson (Coleoptera: Scarabaeidae), with an Illustrated Key to Species, The Coleopterists Bulletin 73 (1), pp. 127-141: 129-130 129-130
Tomarus maracaiboensis     López-García, Margarita M. & Deloya, Cuauhtémoc, 2019, Five New Species of the Dynastine Genus Tomarus Erichson (Coleoptera: Scarabaeidae), with an Illustrated Key to Species, The Coleopterists Bulletin 73 (1), pp. 127-141: 130 130
Tomarus cuniculus     Gillett, Conrad P. D. T. & Gillett, Michael P. T., 2015, The Dynastinae of the island of Saba, Dutch Caribbean (Coleoptera: Scarabaeidae), Insecta Mundi 2015 (433), pp. 1-9: 4-6 4-6
Tomarus fossor     Taboada-Verona, Carlos, Sermeño-Correa, Carlos, Sierra-Serrano, Oscar & Noriega, Jorge Ari, 2019, Checklist of the superfamily Scarabaeoidea (Insecta, Coleoptera) in an urban area of the Caribbean Colombia, Check List 15 (4), pp. 579-594: 584-585 584-585
Tomarus gibbosus     Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, Insecta Mundi 2020 (787), pp. 1-83: 48 48
Tomarus relictus     Freese, Edwin L., Veal, Doug A. & Lago, Paul K., 2020, The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist, Insecta Mundi 2020 (787), pp. 1-83: 48 48
Tomarus rubripes     López-García, Margarita M. & Deloya, Cuauhtémoc, 2018, Description of the Female of Tomarus rostratus Dupuis and New Synonyms and Records for the Genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae), The Coleopterists Bulletin 72 (2), pp. 331-338: 336 336
Tomarus pumilus     López-García, Margarita M. & Deloya, Cuauhtémoc, 2018, Description of the Female of Tomarus rostratus Dupuis and New Synonyms and Records for the Genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae), The Coleopterists Bulletin 72 (2), pp. 331-338: 334 334
Tomarus maimon     López-García, Margarita M. & Deloya, Cuauhtémoc, 2018, Description of the Female of Tomarus rostratus Dupuis and New Synonyms and Records for the Genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae), The Coleopterists Bulletin 72 (2), pp. 331-338: 335 335
Tomarus rostratus     López-García, Margarita M. & Deloya, Cuauhtémoc, 2018, Description of the Female of Tomarus rostratus Dupuis and New Synonyms and Records for the Genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae), The Coleopterists Bulletin 72 (2), pp. 331-338: 332-333 332-333
Tomarus similis     López-García, Margarita M. & Deloya, Cuauhtémoc, 2018, Description of the Female of Tomarus rostratus Dupuis and New Synonyms and Records for the Genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae), The Coleopterists Bulletin 72 (2), pp. 331-338: 336 336
Tomarus cuniculus     López-García, Margarita M. & Deloya, Cuauhtémoc, 2018, Description of the Female of Tomarus rostratus Dupuis and New Synonyms and Records for the Genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae), The Coleopterists Bulletin 72 (2), pp. 331-338: 335 335
Tomarus selanderi     López-García, Margarita M. & Deloya, Cuauhtémoc, 2018, Description of the Female of Tomarus rostratus Dupuis and New Synonyms and Records for the Genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae), The Coleopterists Bulletin 72 (2), pp. 331-338: 335-336 335-336
Tomarus cuniculus     Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, Insecta Mundi 732 (732), pp. 1-306: 211 211
Tomarus maimon     Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, Insecta Mundi 732 (732), pp. 1-306: 213 213
Tomarus bituberculatus     Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, Insecta Mundi 732 (732), pp. 1-306: 210-211 210-211
Tomarus fossor     Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, Insecta Mundi 732 (732), pp. 1-306: 212 212
Tomarus pullus     Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, Insecta Mundi 732 (732), pp. 1-306: 213-214 213-214
Tomarus ebenus     Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, Insecta Mundi 732 (732), pp. 1-306: 212 212
Tomarus similis     Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, Insecta Mundi 732 (732), pp. 1-306: 214 214
Tomarus gyas     Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, Insecta Mundi 732 (732), pp. 1-306: 213 213
Tomarus key    Hielkema, Auke J. & Hielkema, Meindert A., 2019, An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas, Insecta Mundi 732 (732), pp. 1-306: 210 210
Tomarus gibbosus     Ratcliffe, Brett C. & Cave, Ronald D., 2008, The Dynastinae (Coleoptera: Scarabaeidae) of the Bahamas with a description of a new species of Cyclocephala from Great Inagua Island, Insecta Mundi 2008 (24), pp. 1-10: 7 7
Tomarus cuniculus     Ratcliffe, Brett C. & Cave, Ronald D., 2008, The Dynastinae (Coleoptera: Scarabaeidae) of the Bahamas with a description of a new species of Cyclocephala from Great Inagua Island, Insecta Mundi 2008 (24), pp. 1-10: 6 6
Tomarus     Gasca-Álvarez, Héctor & Amat-García, Germán, 2010, Synopsis and key to the genera of Dynastinae (Coleoptera, Scarabaeoidea, Scarabaeidae) of Colombia, ZooKeys 34 (34), pp. 153-192: 185-186 185-186
Tomarus formosianus   new species  Grouvelle, A., 1913, H. Sauter’s Formosa-Ausbeute (Rhysodidae, Nitidulidae, Ostomidae, Colydiidae, Passandridae, Cucujidae, Cryptophagidae, Diphyllidae, Lathridiidae, Mycetophagidae, Dermestidae), Archiv für Naturgeschichte, Abteilung A 79, pp. 33-76: 66 66
Tomarus     Ratcliffe, Brett C. & Cave, Ronald D., 2010, The Dynastinae (Coleoptera: Scarabaeidae) of the Cayman Islands (West Indies), with descriptions of Tomarus adoceteus, new species (Pentodontini) and Caymania nitidissima, new genus and species (Phileurini), Insecta Mundi 2010 (139), pp. 1-15: 6-7 6-7
Tomarus adoceteus     Ratcliffe, Brett C. & Cave, Ronald D., 2010, The Dynastinae (Coleoptera: Scarabaeidae) of the Cayman Islands (West Indies), with descriptions of Tomarus adoceteus, new species (Pentodontini) and Caymania nitidissima, new genus and species (Phileurini), Insecta Mundi 2010 (139), pp. 1-15: 8 8
Tomarus adoceteus   new species  Ratcliffe, Brett C. & Cave, Ronald D., 2010, The Dynastinae (Coleoptera: Scarabaeidae) of the Cayman Islands (West Indies), with descriptions of Tomarus adoceteus, new species (Pentodontini) and Caymania nitidissima, new genus and species (Phileurini), Insecta Mundi 2010 (139), pp. 1-15: 7-8 7-8
Tomarus cuniculus     Ratcliffe, Brett C. & Cave, Ronald D., 2010, The Dynastinae (Coleoptera: Scarabaeidae) of the Cayman Islands (West Indies), with descriptions of Tomarus adoceteus, new species (Pentodontini) and Caymania nitidissima, new genus and species (Phileurini), Insecta Mundi 2010 (139), pp. 1-15: 9 9
Tomarus gyas     López-García, Margarita M., Giraldo-Mendoza, Alfredo E. & Deloya, Cuauhtémoc, 2019, Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru, Zootaxa 4604 (3), pp. 428-440: 431-432 431-432
Tomarus peruvianus     López-García, Margarita M., Giraldo-Mendoza, Alfredo E. & Deloya, Cuauhtémoc, 2019, Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru, Zootaxa 4604 (3), pp. 428-440: 434 434
Tomarus key    López-García, Margarita M., Giraldo-Mendoza, Alfredo E. & Deloya, Cuauhtémoc, 2019, Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru, Zootaxa 4604 (3), pp. 428-440: 438 438
Tomarus maternus     López-García, Margarita M., Giraldo-Mendoza, Alfredo E. & Deloya, Cuauhtémoc, 2019, Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru, Zootaxa 4604 (3), pp. 428-440: 433-434 433-434
Tomarus maimon     López-García, Margarita M., Giraldo-Mendoza, Alfredo E. & Deloya, Cuauhtémoc, 2019, Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru, Zootaxa 4604 (3), pp. 428-440: 432-433 432-433
Tomarus ebenus     López-García, Margarita M., Giraldo-Mendoza, Alfredo E. & Deloya, Cuauhtémoc, 2019, Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru, Zootaxa 4604 (3), pp. 428-440: 430-431 430-431
Tomarus bituberculatus     López-García, Margarita M., Giraldo-Mendoza, Alfredo E. & Deloya, Cuauhtémoc, 2019, Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru, Zootaxa 4604 (3), pp. 428-440: 429-430 429-430
Tomarus pilcopataensis     López-García, Margarita M., Giraldo-Mendoza, Alfredo E. & Deloya, Cuauhtémoc, 2019, Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru, Zootaxa 4604 (3), pp. 428-440: 435 435
Tomarus rostratus     López-García, Margarita M., Giraldo-Mendoza, Alfredo E. & Deloya, Cuauhtémoc, 2019, Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru, Zootaxa 4604 (3), pp. 428-440: 435-436 435-436
Tomarus villosus     López-García, Margarita M., Giraldo-Mendoza, Alfredo E. & Deloya, Cuauhtémoc, 2019, Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru, Zootaxa 4604 (3), pp. 428-440: 436-438 436-438
Tomarus similis     López-García, Margarita M., Giraldo-Mendoza, Alfredo E. & Deloya, Cuauhtémoc, 2019, Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru, Zootaxa 4604 (3), pp. 428-440: 436 436
Tomarus villosus     Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36: 13 13
Tomarus rubripes     Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36: 11 11
Tomarus bidentulus     Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36: 4-5 4-5
Tomarus rosettae     Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36: 10 10
Tomarus spinipenis     Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36: 12 12
Tomarus burmeisteri     Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36: 5-6 5-6
Tomarus maimon     Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36: 8 8
Tomarus     Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36: 14-15 14-15
Tomarus spinipenis   New Species  Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36: 11-12 11-12
Tomarus villosus     Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36: 12 12
Tomarus gyas     Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36: 6-7 6-7
Tomarus rosettae     Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36: 10 10
Tomarus roigjunenti   New Species  Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36: 8-10 8-10
Tomarus maimon     Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa, 2017, The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay, Insecta Mundi 2017 (547), pp. 1-36: 7-8 7-8
Tomarus fossor     López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán, 2015, The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution, Zootaxa 4048 (4), pp. 451-492: 482-484 482-484
Tomarus maternus     López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán, 2015, The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution, Zootaxa 4048 (4), pp. 451-492: 485-486 485-486
Tomarus     López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán, 2015, The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution, Zootaxa 4048 (4), pp. 451-492: 476 476
Tomarus bituberculatus     López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán, 2015, The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution, Zootaxa 4048 (4), pp. 451-492: 477-479 477-479
Tomarus ebenus     López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán, 2015, The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution, Zootaxa 4048 (4), pp. 451-492: 481-482 481-482
Tomarus colombianus     López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán, 2015, The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution, Zootaxa 4048 (4), pp. 451-492: 480 480
Tomarus cicatricosus     López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán, 2015, The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution, Zootaxa 4048 (4), pp. 451-492: 479-480 479-480
Tomarus gyas     López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán, 2015, The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution, Zootaxa 4048 (4), pp. 451-492: 484-485 484-485
Tomarus similis     López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán, 2015, The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution, Zootaxa 4048 (4), pp. 451-492: 488-489 488-489
Tomarus pullus     López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán, 2015, The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution, Zootaxa 4048 (4), pp. 451-492: 487-488 487-488
Tomarus colombianus   new species  López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán, 2014, A new species of Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini), with a key to the species in Colombia, Zootaxa 3869 (5), pp. 579-584: 580-583 580-583

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