Pipunculus calceatus Roser, 1840

Kehlmaier, Christian, 2008, Finnish Pipunculidae (Diptera) studies Part I: Taxonomic notes on Cephalops Fallén, 1810, Pipunculus Latreille, 1802 and Tomosvaryella Aczél, 1939, Zootaxa 1672, pp. 1-42 : 5-6

Pipunculus calceatus Roser, 1840

( Figs 37, 41 & 51–52 )

Pipunculus calceatus Roser, 1840: 55 .

Pipunculus pulchripes Thomson, 1870: 177 . Syn.: Collin (1956: 161).

Material: Germany, Baden-Württemberg: 1Ψ (lectotype P. calceatus ) 1Ψ, Roser ( SMNS); Sweden, Skåne, Bökestad: 1Ψ (lectotype P. pulchripes ), 21/6, ( ZML); Sweden, Skåne, Ivö: 1Ψ (paralectotype P. pulchripes ), ( ZML); Germany, Sachsen, Dresden, Dresdner Heide, 13°46’58’’E 51°05’50’’N, 132m: 1ɗ, 8.VIII.2004, Kehlmaier ( PCCK); Czech Republic, Duchcov, 13.43°E 50.36°N: 1ɗ, 1.VII.1993, Barták ( PCCK); Finland, N, Sibbo, Nevas, 6687:412: 1ɗ, 5.–11.VII.1987; 3ɗ 1Ψ, 13.–20.VII.1985, Albrecht; Spain, Guipúzcoa, San Sebastián, Monte Ulía: 1Ψ, 19.V.2000, Kehlmaier ( PCCK).

Male: Body length: 3.3–4.0mm. Wing length: 3.8–4.7mm. Flagellum dark brown. Frons light brown to grey pollinose. Occiput grey pollinose except posterior to ocellar triangle brown pollinose. Eyes meeting for about length of frons or slightly less (13–15 facets). Postpronotal lobe dark brown. Prescutum and scutum (occ. weakly) brown pollinose. Pleura grey pollinose. Scutellum with apical fringe of about 12 evenly spaced hairs. Subscutellum grey pollinose. LTC about 1.4–2.0 times LFC. Trochanter yellow. Femur with base and apex narrowly yellow (can be more extended on base of hind femur). Hind femur only weakly concave in ventrobasal half or not at all. Tibia yellow, can be darkened midventrally. Tarsi brownish-yellow with distitarsi darkened. Pulvilli of front legs slightly shorter than distitarsi. Tergite 1 dorsolaterally grey, otherwise brown pollinose. Tergites 2–5 laterally with small posterior spots of grey pollinosity, otherwise weakly brown pollinose. Dorsally, tergite 2 in anterior three quarters, tergite 3 in anterior half, tergite 4 in anterior third and tergite 5 only narrowly along anterior margin brown pollinose, otherwise shining. Membranous area key-hole shaped but may appear slot-like in caudal view (the widening of upper half best seen in dorsal view) and normally with distinct keel. Surstyli with blunt apices ( Fig. 37 ).

Female: Body length: 3.1–3.7mm. Wing length: 3.9–4.8mm. Flagellum brown. Frons entirely grey pollinose. Postpronotal lobe light to dark brown. Prescutum and scutum grey pollinose in anterior quarter to third, laterally down to wing base, otherwise brown pollinose. Pleura and subscutellum grey pollinose. LTC about 1.0–1.2 times LFC. Trochanter yellowish. Femur with as much as basal third and apex yellow (especially on hind femur), occasionally base can only be narrowly yellow. Tibia yellow, can be somewhat darkened ventromedially. Tarsi yellow with distitarsi darkened. Pulvilli of front legs at least slightly longer than distitarsi. Tergite 1 anterodorsally, tergite 2 dorsocentrally brown pollinose (but may be considerably reduced and hence shining), otherwise grey pollinose. Tergites 3–6 dorsally shining, laterally with triangular spots of grey pollinosity. Tergites 3–4 can show a very narrow anterior stripe of brown pollinosity. Ovipositor in lateral view with rather short, weakly curved piercer. Transition from base to piercer with distinct ventral step ( Fig. 51 ). Ovipositor viewed dorsally with distinct transition between base and piercer ( Fig. 52 ). Ovipositor’s base can be more round in dorsal and higher in lateral view compared to the figures presented.

Annotations: Roser (1840) described P. calceatus from an unknown number of specimens. Detailed information on the history of Roser’s collection can be found in Kehlmaier (2006). At SMNS, a single female specimen was obtained showing a label stating “ Pipunculus calceatus m.” in Roser’s hand-writing and another label stating “ Type ” [written in red] in Lindner’s hand-writing. The specimen is hereby designated lectotype and was labelled accordingly. One additional male and one female of P. c a l c e a t u s were received labelled as “ Pip. calceatus v. Ros.” by Lindner only. These specimens are not considered having type status here due to the lack of any evidence for this.

Thomson (1870) described P. pulchripes from an unspecified number of females collected in “Funnen i Skåne”. Collin (1956) designated a lectotype and put the name in synonymy with P. calceatus . Study of type material at ZML confirmed the synonymy.

For the eastern Palaearctic region, P. calceatus is known from the Korean Peninsula as mentioned in Kozánek & Belcari (1995) as unpublished data from Kozánek.

Figures

FIGURES 36 – 42. Surstyli in strictly dorsal and outer surstylus in lateral view of male Pipunculus carlestolrai (36 & 40), Pipunculus calceatus (37 & 41) and Pipunculus omissinervis (38 – 39 & 42). Scale bars = 0.1 mm.

FIGURES 43 – 54. Ovipositor in lateral and dorsal view of female Pipunculus violovitshi (43 – 44), Pipunculus elegans (45 – 46), Pipunculus oldenbergi (47 – 48), Pipunculus zugmayeriae (49 – 50), Pipunculus calceatus (51 – 52) and Pipunculus stackelbergi (53 – 54, taken from Kuznetzov (1991 )). Scale bars = 0.2 mm.