Phylloicus magnus Banks

PRATHER, AYSHA L., 2003, Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae), Zootaxa 275 (1), pp. 1-214 : 68-69

Phylloicus magnus Banks

Figs. 66 , 67

Phylloicus magnus Banks, 1913:236 [Type locality: Colombia, Monte Socorro; MCZ; male]. — Flint 1967:19 [male].

As suggested by its name, P. magnus is probably the largest species of Phylloicus . It is also distinguished by the long coxopodite, the notched apex of tergum X, and the short preanal appendage. The female specimen described here is circumstantially associated. Like the holotype of P. magnus , it is very large. The only Colombian species large enough to be conspecific with this female is P. magnus . The description of the adult is based on this female specimen, which is much less rubbed than the male.

Adult. Forewing length 16.2­18.5 mm, n = 2.

Head chestnut brown, with dorsomesal crest of dark brown setae. Maxillary palps dark brown, covered with dark brown setae. Antenna of male twice forewing length; female antenna 11/2 times forewing length; chestnut brown, with narrow patches of pale sensilla on anteromesal surface of each flagellomere. Dorsal pterothorax chestnut brown; ventrolateral thorax golden brown. Legs golden brown. Metathoracic leg of male with posterior fringe of long setae, setae pale. Tibial spur formula 2,4,4. Forewing flat; chestnut brown; with golden bands and dark patches; with single basal stripe, golden; with irregular gold and dark brown patches posterad of Cu 1; with small golden spots marking nygma and thyridium. Hind wing basal brush absent.

Male. Preterminalic abdominal terga without anteromesal notches. Corematic structures absent, terga III­V unmodified, without membranous lobes or sclerotized processes. Sternum VII without anteromesal process. Sternum VIII similar to anterior sterna, sternum IX not elongate. Tergum IX with paired mesal ridges extending 1/2 length of segment; posterior margin smoothly rounded ( Fig. 66B ); lateral ridge present; dorsal pleural setae approximately 5, ventral pleural setae absent ( Fig. 66A ); sternum IX with paired mesolateral ridges; sternum IX ( Fig. 66C ). Preanal appendage less than 2/3 length of tergum X, narrowly elliptic, setae long, but not filamentous or longer than appendage ( Fig. 66A, B ). Tergum X without basal lobes; basodorsal process absent; basolateral processes absent; apex, in lateral view, truncate, in dorsal view, notched, notch triangular; with paired short basomesal processes, each bearing single spine­like seta ( Fig. 66A, B ). Harpago rounded; peglike setae few, apical ( Fig. 66A, C ). Phallotremal sclerites average size, longest dimension less than diameter of phallobase; dorsal sclerite ovoid, in dorsal view horseshoeshaped ( Fig. 66D, E ).

Female. Preterminal abdominal terga without anteromesal notches. Sternum VII without anteromesal process. Tergum VIII without posterolateral brush; sternum VIII with shallow posteromesal notch; sternum VIII ( Fig. 67C ). Tergum IX without mesal ridge ( Fig. 67B ). Sternum IX anterior and posterior lobes darkly sclerotized and striate, punctate, with shallow pockets anterolateral to vaginal opening ( Fig. 67A ). Tergum X appendage shorter than mesal lobe, base marked by faint suture line, apex rounded; mesal lobe with single mesal seta; digitate lateral processes absent ( Fig. 67B ). Sternum X with patches of short fine setae posterolaterally to anal opening ( Fig. 67A ). Vaginal apparatus anterior and posterior sclerites equal in length; anterior sclerite rounded anteriorly, posterolateral projections rounded; posterior sclerite triangular; posterior end of spermatheca a sclerotized sphere ( Fig. 67A ).

Material examined. COLOMBIA: Monte Socorro, 3600 m, Fassl — holotype male ( MCZ); Caldas: 5 km W Termales de Ruiz, 3200 m, 27­29.ii.1984, C & O Flint — 1 female ( NMNH) .

Distribution. Colombia.

Figures

FIGURE 66. Phylloicus magnus. Male (holotype): A—lateral view; B—dorsal view; C—ventral view; D—phallus, lateral view.

FIGURE 67. Phylloicus magnus. Female (UMSP000068008): A—sterna IX, X and vaginal apparatus, ventral view; B—terga IX and X, dorsal view; C—sternum VIII, ventral view.