Orthobula quadrinotata, Deeleman-Reinhold, Christa, 2001

Deeleman-Reinhold, Christa, 2001, Forest Spiders of South East Asia With a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanteriidae)., Forest Spiders of South East Asia With a revision of the sac and ground spiders- Family Liocranidae, Leiden, Netherlands: Brill Leiden; Boston; Köln, pp. 400-505 : 435-436

Orthobula quadrinotata sp. n. ( figs 700-704 , map 36 )

Type locality. — Indonesia, SE Sulawesi.

Material examined. — Holotype ♂ from the type locality, 40 km W of Kendari, Sampara, secondary forest litter , 12.viii. 1980, P.R. and C.L. Deeleman; paratypes: 2 ♂ 2 ♀, same data .

Other material. — C Kalimantan, Kaharian , 2°02'S, 113°40'E, primary peat swamp forest, 1 ♀, leaf litter, 2-16.ix. 1985, S. Djojosudharmo.

Diagnosis. — A close relative of impressa , trinotata and bilobata . O. quadrinotata differs from trinotata by the presence of three anterior dark marks on a creamy white background on the abdomen in addition to a large broad posterior one, further by the shorter more compact embolus with proximal hook as in bilobata ; from crucifera by the proximal hook on the embolus and in the epigyne by the conspicuous komma-shaped ducts in front of the spermathecae, reminiscent of that in bilobata . A clearly smaller and lighter species than bilobata , distinct from that species by the tegulum being less expanded and from bilobata and yaginumai by the presence of a long spine adjacent to the tibial apophysis; from yag ­ inumai also by the dorsal abdominal pattern and the spine at the tibial apophysis.

Description. — MALE. Total length 1.90 mm. Carapace length 0.95 mm, width 0.75 mm, height at coxae II 0.30 mm, head width 0.45 mm, width eye region 0.30 mm; abdomen 0.80 mm long, 0.70 mm wide. Leg lengths: leg I 2.25 mm (0.60-0.85-0.45-0.35), leg II 1.65 mm (0.50-0.55-0.35-0.25), leg III 1.60 mm (0.50-0.55-0.30-0.25), leg IV 2.25 mm (0.60-0.75-0.65-0.25), palp 0.25-0.15- 0.12-0.45 mm. Carapace and mouthparts deep orange, femora orange, rest of legs uniform pale yellow, abdomen predominantly creamy white, with dark grey rear part and additionally three dark grey spots on the anterior median and lateral sides as in fig. 700 ; in dark specimens the grey area is extended over the flanks of the abdomen. Carapace with smooth median band, rows of large round depressions towards the sides, sternum with some round impressions. ALE a little larger AME, AME slightly separated, adjacent to ALE; PME separated by a little less than their d, almost adjacent to PLE. Leg spination: tibiae I with 6pv and 5rv, tibia II with 5pv and 4rv or 5rv; metatarsus I and II with 2-2-2-2v, tarsus I and II with 2- 2v. Abdomen shiny, dorsally entirely covered by scutum, ventrally an epigastric and a partial postgenital scutum. Palp figs 701-702 , tibial apophysis accompanied by a long spine; tegulum slightly bilobed posteriorly, maximal width about 2 x that of cymbium.

FEMALE. Total length 2.15 mm. Carapace length 1.00 mm, width 0.80 mm, head width 0.50 mm, eye group width 0.30 mm; abdomen 1.05 mm long, 0.80 mm wide, epigynal plate 0.25 mm wide, 2.5 mm long. Leg lengths: leg I 2.40 mm (0.65-0.90-0.55-0.30), leg II 2.15 mm (0.65-0.75-0.50-0.25), leg III 1.80 mm (0.55-0.60-0.45-0.20), leg IV 2.55 mm (0.75-0.85-0.60-0.35), palp 0.30-0.12-0.14- 0.22 mm. Carapace, eyes, mouthparts and legs as in male. Abdomen colouration as male. Epigyne ( fig. 703 ) characterized by conspicuous komma-shaped openings annex ducts; surface showing a round depression in that area. Vulva ( fig. 704 ) with large membranous bursae, issued from a chitinous coiled base situated between openings and spermathecae.

Distribution. — Indonesia. Found in SE Sulawesi and C Kalimantan.

Etymology. — In analogy with Orthobula trinotata , this species is called quadrinotata , composed of Latin quadri- = four, notata = markings.

Figures

Map 36: (t) Orthobula trinotata (c) O. crucifera (y) O. yaginum ai ● O. bilobata (i) O. impressa ■ O. quadrinotata ▲ O. pura Ө Unidentified specimen 1 - 4.

Figs 697 - 699. Orthobula impressa. 697. Fem ale Tissam aharam a, pattern of abdom en. 698. M ale Ratnapura, left palpal bulb, ventral. 699. Epigyne. 700 - 704. Orthobula quadrinotata sp. n. 700. M ale, abdom en, dorsal. 701. Left palpal cym bium and bulb, ventral. 702. Palp, retrolateral. 703. Fem ale Tissam aharam a, epigyne. 704. Vulva, in clove oil, dorsal.