Nyctiophylax (Paranyctiophylax) dhauli, Oláh & Johanson, 2010

Oláh, János & Johanson, Kjell Arne, 2010, Generic review of Polycentropodidae with description of 32 new species and 19 new species records from the Oriental, Australian and Afrotropical Biogeographical Regions 2435, Zootaxa 2435 (1), pp. 1-63 : 46

Nyctiophylax (Paranyctiophylax) dhauli , new species

Figures 80–82

Diagnosis: The gonopod shape of this species is intermediate between those of species with short, basally curved gonopods and species with a long ventral lobe of each gonopod. The gonopods in this species are accompanied by slender dorsal and ventral paraproctal processes. There are several species from Sri Lanka to Indonesia with similar genitalia. This species strongly resembles N. (P.) amykos Malicky from Nepal. In N. (P.) dhauli , the sternite IX is vertically tall, not horizontally elongate; the apicoventral mesal lobe of sternite IX is sharply triangular, not blunt and rounded in lateral view; the cerci are elongate, foliaceous, not narrow at basal 1/3rd; the mesoventral branch of each gonopod is longer; the dorsal branch of each gonopod is more slender with a concave dorsum, not stout and convex; the dorsal paraproctal processes are slightly longer than the cerci; and the ventral paraproctal processes are dorsally geniculate.

Description: Male. Body uniformly pale brown. Maxillary palp formula (I,II)-IV-III-V. Forewing length 4.0 mm. Forewing A1, A2, and A3 looped.

Male genitalia. Abdominal segment IX reduced to robust sternite, tall, nearly triangular ( Fig. 80 ); tergum IX membranous, with no narrow sclerotized stripe discernible. Segment X membranous, covering dorsal part of phallic apparatus. Cerci setose, narrow, elongate, foliaceous in lateral view; slightly shorter than dorsal paraproctal processes. Paraproctal complexes well-developed, smooth; each composed of paraproctal base (anterior body formed by meeting point of dorsal and ventral paraproctal processes) heavily sclerotized, narrow; dorsal paraproctal process slender, weakly sclerotized, straight in lateral view; and ventral paraproctal process with dorsum geniculate, apex with several small setae. Gonopods with short, sharply triangular mesoventral branch ( Fig. 80 ); dorsal branch long, slender, with concave dorsum ( Fig. 80 ); dorsal branch with blunt apex. Phallic apparatus ( Fig. 82 ) located dorsally in genitalia, guided by paraproctal processes, membranous segment X, and membranous subphallic bridge. Phallotheca thick, horizontal; endophallus with pair of slightly sclerotized sclerites and ventrad-curving spines; apical part ending in row of elongate spicules.

Holotype male: INDIA: Orissa State, Bhubaneswar, Dhauli , marshy area, 20–28.ii.1987, light [J. Oláh] ( OPC).

Distribution: India.

Etymology: Dhauli, named after the Dhauli Hills at Daya River in Orissa State, near Bhubaneswar, India, where this animal was collected in a marshy area of the famous battlefield of the bloody war between King Ashoka and the King of Kalinga. The water of River Daya turned red and Ashoka, witnessing the mass deaths, thereafter pursued a policy of non-violence. After converting to Buddhism, he was responsible for the proliferation of Buddhist ideals across the whole of East Asia and Southeast Asia.

Figures

FIGURES 80–82. Nyctiophylax (Paranyctiophylax) dhauli, new species, holotype. 80—genitalia, left lateral; 81—left gonopod, ventral; 82—phallic apparatus, left lateral.