Ischnothyreus pterodactyl, Edward & Harvey, 2014

Edward, Karen L. & Harvey, Mark S., 2014, Australian Goblin Spiders Of The Genus Ischnothyreus (Araneae, Oonopidae), Bulletin of the American Museum of Natural History 2014 (389), pp. 1-144 : 129-133

Ischnothyreus pterodactyl , new species Figures 72–73 , map 14

TYPES: AUSTRALIA: Queensland: Male holotype and female allotype from Mount

Superbus, false summit, 28.23333 ° S, 152.46670 ° E, 1200 m (12 Mar–13 June 1990 T.B. Churchill, R.J. Raven), deposited in QM (³ holotype: QM S16568 , PBI _OON 21959; ♀ allotype: QM S16568 , PBI _ OON 5609 ) .

ETYMOLOGY: The specific epithet is a noun in apposition.

DIAGNOSIS: This species can be easily recognized by the extremely elongate, obtusely bent embolic region of the male palp (fig. 73A, B) and the basal process of the fang consisting of numerous heavily sclerotized lobes (fig. 73C, D). The female possesses a heavily sclerotized circular epigynal atrium, which appears to be fused to the overhanging process and the horizontal sclerotization anterior to the epigynal atrium is less sclerotized than I. rixi , sp. nov. (figs. 72I, 73E).

MALE (PBI_OON 21959, fig. 72 A–C, G, H; paratype (PBI_OON 05631), fig. 73A–D). Total length 1.73. CEPHALOTHORAX: Carapace brown, broadly oval in dorsal view, pars cephalica slightly elevated in lateral view, anteriorly narrowed to between 0.5 and 0.75 times its maximum width, anterolateral corners without extension or projections, surface of elevated portion of pars cephalica strongly reticulate, sides strongly reticulate. Clypeus margin unmodified, curved downward in front view, low, ALE separated from edge of carapace by less than their radius; setae dark. Eyes: ALE largest, ALE circular, PME oval, PLE circular; posterior eye row straight from above; ALE touching, ALE-PLE touching. Sternum as long as wide, pale orange, uniform; setae dark, evenly scattered. Chelicerae, endites, and labium pale orange. Chelicerae slightly divergent, anterior face unmodified; promargin with one larger denticle; fang shape normal, basal process consisting of numerous heavily sclerotized lobes, anterior lobe with distal ridges; setae dark. Labium elongated hexagon, not fused to sternum, anterior margin indented at middle, same as sternum in sclerotization; with six or more setae on anterior margin. Endites distally not excavated, anteromedian tip with one strong, toothlike projection, same as sternum in sclerotization. ABDOMEN: Ovoid; dorsum soft portions pale orange. Book lung covers elliptical. Dorsal scutum brown, covering more than 3/4 of abdomen, no soft tissue visible from above, middle surface finely reticulate, sides finely reticulate. Epigastric scutum small lateral sclerites absent. Postepigastric scutum orange-brown, covering about 2/3 of abdominal length. Dorsum, epigastric area, and postepigastric area setae dark. LEGS: Orange-brown, without color pattern; patella plus tibia I shorter than carapace. Leg spination: femora: I p0-2-0; II p0-1-0; tibiae: I, II p2-2-0; v2-2-0; metatarsi: I, II p1-1-0; v1-1-0. GENITALIA: Palp proximal segments dark red-brown; embolus dark; femur shorter than trochanter; patella about as long as femur; cymbium dark redbrown; bulb dark red-brown, more than two times as long as cymbium, stout; embolic region extremely narrow and elongate, obtusely bent, tip rounded, transparent membranous tissues visible medially (fig. 73A, B).

FEMALE (PBI_OON 5609, fig. 72D–E, I; paratype (PBI_OON 5639), fig. 73E). Total length 1.98. CEPHALOTHORAX: Carapace anteriorly narrowed to 0.49 times its maximum width or less. ABDOMEN: Dorsal scutum covering 1/2 to 3/4 of abdomen, more than 1/2 to most of abdomen width. Postepigastric scutum widely hexagonal, covering about 1/3 of the abdominal length. LEGS: Leg spination: femora: I p0-2-2; II p0-1-0; tibiae: I, II p2-2-0; v2-2-0; metatarsi: I, II p1-1-0; v1-1-0. GENITALIA: Ventral view: strong horizontal scleriotization anterior to rounded epigynal atrium clearly joins posteriorly directed lateral apodemes; heavy sclerotization around epigynal atrium circular, thick; overhanging process slightly triangular, sides appear fused to postepigastric scutum; convoluted duct thicker than apodemes (figs. 72I, 73E).

OTHER MATERIAL EXAMINED: AUS- TRALIA: New South Wales: Beech Picnic Area, Wiangaree , 1000 m, 28.37300 ° S, 153.10070 ° E, Apr. 22, 2002 (G. Monteith, QM S69613 , PBI _OON 22217), 3 ♀ ; Border Ranges National Park, Brindle Creek, rainforest alongside Creek , leaf litter, 28.37277 ° S, 153.06916 ° E, May 9, 2007 (K. Edward and K. Pitz, QM S96010 , PBI _OON 25729), 1 ♀ ; Bruxner Pk, via Coffs Harbour , rainforest, leaf litter, 150 m, 30.23333 ° S, 153.10000 ° E, Mar. 22, 1980 (G. Monteith, QM S16054 ), 1 ♀ ; Bruxner Park , 150 m, 30.23333 ° S, 153.10000 ° E, Mar. 22–Nov. 13, 1980 (G. Monteith, QM S16045 , S16151 ), 7 ³, 9 ♀ ; Bruxner Park , 150 m, 30.23333 ° S, 153.10000 ° E, Nov. 13–Mar. 17, 1980 (G. Monteith, QM S16114 ), 1 ♀ ; Bruxner Pk, via Coffs Harbour , rainforest, leaf litter, 150 m, 30.23333 ° S, 153.10000 ° E, May 14, 1969 (R.W. Taylor, ANIC, PBI _OON 25871), 2 ³ ; Bruxner Pk, via Coffs Harbour , rainforest, leaf mold, rotten wood, 150 m, 30.23333 ° S, 153.10000 ° E, June 25, 1967 (R.W. Taylor, ANIC, PBI _OON 26297), 1 ³, 1 ♀ ; Gibraltar Range National Park , rainforest, 450 m, 29.58333 ° S, 152.21670 ° E, Mar. 30, 1980 (G. Monteith, QM S16112 ), 1 ³ ; Washpool State Forest, Coombadjah Creek , leaf litter, 29.26666 ° S, 152.36666 ° E, Feb. 12, 1982 (C. Horseman, AM KS9354 , PBI _OON 26293), 2 ♀. Queensland: 5.5 km E of One Tree Hill, semievergreen vine thicket, 120 m, 25.33583 ° S, 151.90970 ° E, Dec. 14, 1999, to Mar. 19, 2000 (G. and S. Monteith, QM S57432, PBI _OON 22134), 1 ³ ; 5.5 km E of One Tree Hill , semievergreen vine thicket, 120 m, 25.33583 ° S, 151.90970 ° E, Dec. 14, 1999, to Mar. 19, 2000 (G. and S. Monteith, QM S47072 , PBI _OON 22246), 1 ³, 1 ♀ ; Kroombit Tops, 45 km SSW. of Calliope , rainforest, leaf litter, 890 m, 24.41666 ° S, 151.01666 ° E, Dec. 10, 1983 (G. Monteith, V. Davies, J. Gallon, G. Thomp- son, QM S16096 , PBI _OON 25890), 1 ³ ; Lamington National Park, Binnaburra Lodge , 28.18333 ° S, 153.18333 ° E (July 2, 1986, M.S. Harvey, P.J. Vaughan, WAM T90 /912, PBI _ OON 00004311–2 ), 1 ³, 1 ♀ ; Lamington National Park, O’Reillys Guest House , rainforest, leaf litter, 28.23333 ° S, 153.13333 ° E, Dec. 14, 1981 (G. Monteith, D. Yeates, QM S12990 , S16064 , S16185 , S12985 ), 3 ³, 4 ♀ ; Lamington National Park, O’Reilly’s , rainforest, 28.23333 ° S, 153.13333 ° E, Mar. 2–Apr. 26, 1980 (G. Monteith, D. Yeates, QM S78262 , PBI _ OON 7218 ), 4 ³ ; Lamington National Park, O’Reilly’s , rainforest, 28.23333 ° S, 153.13333 ° E, Mar. 2–Apr. 26, 1980 (G. Monteith, D. Yeates, QM S78249, PBI _ OON 7230 ), 1 ³, 1 ♀ ; Lamington National Park, O’Reilly’s , rainforest, leaf litter, 28.23333 ° S, 153.13333 ° E, Nov. 22–27, 1978 (Lawrence, Weir, ANIC, PBI _OON 26295), 1 ³ ; Lamington National Park, O’Reilly’s , rainforest, leaf litter, 28.23333 ° S, 153.13333 ° E, Nov. 22–27, 1978 (Lawrence, Weir, ANIC, PBI _OON 26296), 1 ³, 1 ♀ ; Lamington National Park , rainforest, leaf litter, 960 m, 28.20000 ° S, 153.16670 ° E, July 9–10, 1977 (R. Raven, QM, S16198 , S86457 , S86440, S86437 , S86465– S86466 , S86448, S86459), 8 ³, 3 ♀ ; Lamington National Park, Binna Burra , rainforest, leaf litter, 28.18333 ° S, 153.18333 ° E, Mar. 21–23, 2006 (M. Ramírez, QM S72917, PBI _OON 21974), 1 ♀ ; Lamington National Park, IBISCA Plot # IQ–700–CKA, rainforest, leaf litter, 720 m, 28.23694 ° S, 153.15194 ° E, Jan. 22, 2008 (C. Burwell, QM, S86441 , S86443 , S86455, S16018 – S16021 ), 11 ³, 4 ♀ ; Mount Superbus, Site 7, 850 m, 28.23333 ° S, 152.48330 ° E, Mar. 12–June 13, 1990 (T.B. Churchill, R.J. Raven, QM S15910 , PBI _OON 21994), 1 ♀ ; Mount Superbus , summit, 1350 m, 28.22183 ° S, 152.45280 ° E, Oct. 24, 1998 (G. Monteith, QM S54213 , PBI _OON 21973), 1 ♀ ; Mount Superbus, Trap 1, 1350 m, 28.23333 ° S, 152.48330 ° E, Mar. 12–June 13, 1990 (T.B. Churchill, R.J. Raven, QM S15911 , PBI _OON 22004), 1 ♀ ; Tamborine National Park, Curtis Falls Track , rainforest, leaf litter, 27.93194 ° S, 153.19388 ° E, May 11, 2007 (K. Edward and K. Pitz, QM S96011 , PBI _OON 25728), 1 ♀ ; Upper Tallebudgera Creek , 550 m, 28.25000 ° S, 153.26670 ° E, Jan. 8, 1984 (G. Monteith, G. Thompson, QM S69632, PBI _OON 22041), 1 ³ ; Upper Tallebudgera valley, below Springbrook , rainforest, leaf litter, 550 m, 28.25000 ° S, 153.26666 ° E, Jan. 8–Mar. 17, 1985 (G. Monteith, D. Cook, G. Thompson, QM S16473 , PBI _ OON 5910 ), 2 ♀ ; Lamington National Park, Binna Burra, along Border Track , 900 m, 28.19861 ° S, 153.1875 ° E, Apr 29–30, 2009 (H. Wood, CASENT 9035029, PBI _ OON 00005631 ), 1 ³ ; Lamington National Park, Binna Burra, along Border Track , 900 m, 28.19861 ° S, 153.1875 ° E, Apr 29–30, 2009 (H. Wood, CASENT 9035032, PBI _ OON 00005639 , 1 ♀.

DISTRIBUTION: This species is relatively widespread from Coffs Harbour, New South- Wales to Bundaberg, northeastern Queensland (map 14).

Figures

Fig. 72. Ischnothyreus pterodactyl, sp. nov. Holotype male (PBI_OON 00021959): A. habitus, dorsal view; B. habitus, ventral view; C. habitus, lateral view; G. carapace, anterior view; H. sternum, ventral view. Allotype female (PBI_OON 00005609): D. habitus, dorsal view; E. habitus, ventral view; F. habitus, lateral view; I. Epigynum, ventral view.

Fig. 73. Ischnothyreus pterodactyl, sp. nov. Paratype male (PBI_OON 00005631): A. left palp, prolateral view; B. left palp, retrolateral view; C. left chelicerae, anterior view; D. left chelicerae, posterior view. Paratype female (PBI_OON 00005639): E. epigynum, ventral view. Scale lines 5 0.1 mm.