Goja garridoi, Carrejo & Obando & Casasola-González & García Aldrete, 2021

Carrejo, Nancy, Obando, Ranulfo González, Casasola-González, José Arturo & García Aldrete, Alfonso N., 2021, New Colombian Goja Navás (Psocodea: ‘ Psocoptera’: Epipsocidae) with peculiar genitalia, and the first Goja with brachypterous male, from Oaxaca, Mexico, Zootaxa 5040 (4), pp. 451-481 : 463

Goja garridoi n. sp. Male

( Figs 7–12 )

Diagnosis. With G. meremberg n. sp., G. tenerife n. sp., and G. vavilovi n. sp., it constitutes an assemblage of related species, characterized by having the hypandrium with a median posterior process, by having a large process on each side of the posterior end of the aedeagal arch, and by having endophallic sclerites unlike those in the known species of Goja . It differs from the other species in the assemblage, by having the hypandrium with a wide, short, rectangular process, by having the posterior processes of the phallosome club-shaped, the right one with a curved, rectangular flap, and by having the right (lateral) endophallic sclerites wider than in G. meremberg and more elongated than in G. tenerife . ( Fig. 12 ).

Color (in 80% ethanol). Body cream. Head pattern ( Fig. 9 ), vertex with broad dark brown to ochre area between the compound eyes, covering triangular area, slightly concave posteriorly; forehead, clypeus and labrum brown, postclypeus with convergent oblique dark brown striae; labral sclerites and upper part of labrum dark brown; genae and postgenae yellowish, genae with small dark brown spots in lower area. Compound eyes dark brown; ocellar triangle black. Antennae pale brown. Maxillary palps pale brown, Mx 4 brown. Thorax brown, tergal lobes of meso- and metathorax dark brown, metapleura brown. Legs pale brown. Forewings ( Fig. 7 ) almost hyaline; pterostigma with median and distal pale brown bands; veins brown. Hindwings ( Fig. 8 ), hyaline, veins brown. Abdomen cream, with subcuticular spots brown. Clunium pale brown, posterior margin brown. Hypandrium brown. Phallosome with aedeagal arch and endophallic sclerites dark brown. Epiproct and paraprocts pale brown.

Morphology. Head: H/MxW: 1.55; compound eyes large, H/D: 3.04; IO/MxW: 0.66. Outer cusp of lacinial tips broad, with nine denticles. Mx4/Mx2: 1.06. Forewings ( Fig. 7 ): FW/W: 2.82; Rs 4-6-branched; M 5-7-branched, main branches may be forked; pterostigma elongate: lp/wp: 6.19; areola postica low, elongate: al/ah: 3.41. Hindwings ( Fig. 8 ): HW/w: 3.25; Rs 2-3-branched; M 2-4-branched; Rs and M joined by a crossvein. Legs: fore- tarsi without ctenidiobothria; mid- and hind- tarsi with ctenidiobothria. Hypandrium ( Fig. 11 ) broad, setose, pigmented area deeply concave. Phallosome ( Fig. 12 ): concave anteriorly side struts straight, widening distally; apex of aedeagal arch with two large, rounded processes. Two pairs of endophallic sclerites, asymmetric, lateral sclerites broad, wider in the middle, posteriorly blunt; mesal sclerites curved, distally acuminate. Paraprocts ( Fig. 10 ) large, oval, with setae and macrosetae in distal border; sensory fields with 32 trichobothria in basal rosettes. Epiproct ( Fig. 10 ) broad, almost straight anteriorly, rounded posteriorly, one macroseta mesally, near anterior border, distal field with setae and macrosetae as illustrated.

Measurements. FW: 6550, HW: 4715, F: 1485, T: 2715, t1: 1225, t2: 345, ctt1: 42, f1: 1450, f 2: 1237, f3: 912, f4: 575, f5: 320, f6; 312, f7: 257, f8: 240, f9: 227, f10: 207, f11: 262, Mx4: 370, IO: 460, D: 355, d: 340, IO/d: 1.35, PO: 0.96.

Specimen studied. Holotype male. COLOMBIA. Cauca. Puracé, Cueva de los Guácharos , 2°18’26.3”N: 76°15’4.6”W. 2789 m. 7.VI.2019. D. H. Garrido-Torres. Beating vegetation.

Etymology. This species is dedicated to its collector Daniel Hernando Garrido-Torres.

Figures

FIGURES 7‒12. Goja garridoi n. sp. Male. 7. Forewing. 8. Hindwing. 9. Front view of head. 10. Left paraproct and epiproct. 11. Hypandrium. 12. Phallosome. Scales in mm.